World Spider Catalog

Linyphiidae Blackwall, 1859

Family | ACCEPTED

N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; Tian et al., 2022 show five subfamilies: Linyphiinae (incl. Mynogleninae Lehtinen, 1967: 250 and Dubiaraneinae Millidge, 1993c: 145), Erigoninae Emerton, 1882: 7, Micronetinae Hull, 1920a: 8, Ipainae Saaristo, 2007b: 34 and Stemonyphantinae Wunderlich, 1986: 120; da Silva-Moreira, Kulkarni & Hormiga, 2025: 60-62 list resp. newly circumscribe six subfamilies: Erigoninae, Lepthyphantinae Simon, 1929: 562, Linyphiinae (syn. nov.: Dubiaraneinae), Mynogleninae, Pocobletinae Silva-Moreira & Hormiga, 2025: 61 (new subfamily) and Stemonyphantinae, state Ipainae and Micronetinae to be polyphyletic, leave several linyphiid clades unranked and also several genera as incertae sedis.

In synonymy:

Scleroschema Hull, 1911 = Araeoncus Simon, 1884 (Denis, 1942a: 84)

Gen. Araeoncus Simon, 1884 urn:lsid:nmbe.ch:spidergen:00840

type Walckenaeria humilis Blackwall, 1841; gender masculine

N.B.: type species fixed by Simon, 1894a: 652; considered a senior synonym of Scleroschema Hull, 1911b: 51 [urn:lsid:nmbe.ch:spidergen:04156] (type Walckenaeria humilis Blackwall, 1841) by Denis, 1942a: 84; a generic re-organisation of the Savignia group (incl. Araeoncus, Dicymbium, Diplocephalus, Erigonella, Glyphesis) is pending (Frick, Nentwig & Kropf, 2010); the proposed synonymy with Savignia Blackwall, 1833 by Breitling, 2021: 22 is not followed here as it is based mainly on COI (see FAQ 7).

Transferred to other genera:

Araeoncus aequus O. Pickard-Cambridge, 1911 -- see Acartauchenius

Araeoncus americanus Chamberlin & Ivie, 1944 -- see Coloncus

Araeoncus brunneus Bösenberg, 1902 -- see Diplocephalus

Araeoncus duriusculus Caporiacco, 1935 -- see Styloctetor

Araeoncus erratus O. Pickard-Cambridge, 1908 -- see Acartauchenius

Araeoncus hiemalis (Blackwall, 1841) -- see Erigonella

Araeoncus krugeri Simon, 1894 -- see Tybaertiella

Araeoncus longicephalus Saito, 1988 -- see Saitonia

Araeoncus muscus Saito, 1989 -- see Saitonia

Araeoncus ojiroensis Saito, 1990 -- see Saitonia

Araeoncus orientalis Oi, 1960 -- see Saitonia

Araeoncus patellatus Emerton, 1917 -- see Scotinotylus

Araeoncus prospiciens (Thorell, 1875) -- see Archaraeoncus

Araeoncus pusillus Tullgren, 1955 -- see Saloca

Araeoncus ruderalis Sørensen, 1898 -- see Arctilaira

Araeoncus stigmosus Xia, Zhang, Gao, Fei & Kim, 2001 -- see Tibioploides

Araeoncus strandi Ermolajev, 1937 -- see Tanasevitchia

Araeoncus tianschanicus Hu & Wu, 1989 -- see Archaraeoncus

In synonymy:

Araeoncus affinitatus (O. Pickard-Cambridge, 1863, T from Walckenaeria) = Araeoncus crassiceps (Westring, 1861) (Simon, 1884a: 638)

Araeoncus excelsus (L. Koch, 1879) = Araeoncus crassiceps (Westring, 1861) (Holm, 1973: 78)

Araeoncus globiceps (Menge, 1868, T from Lophocarenum) = Araeoncus humilis (Blackwall, 1841) (Simon, 1884a: 636)

Araeoncus meridionalis Denis, 1954 = Araeoncus humilis (Blackwall, 1841) (Bosmans, 1996: 125)

Araeoncus praeceps Holm, 1962 = Araeoncus victorianyanzae Berland, 1936 (Scharff, 1990a: 9)

Araeoncus altissimus Simon, 1884 | | France, Italy, Azerbaijan urn:lsid:nmbe.ch:spidersp:009419

Araeoncus altissimus Simon, 1884a: 640, f. 471-473 (Dm) Original description

Araeoncus altissimus Simon, 1894a: 611, f. 642-643 (m)

Araeoncus altissimus Simon, 1926: 366, 492, f. 649 (m, Df)

Araeoncus altissimus Denis, 1948a: 19, f. 3-5 (mf)

Araeoncus altissimus Tanasevitch, 1987a: 331, f. 54-58 (m)

Araeoncus altissimus Tanasevitch, 1990: 111, f. 22.2, 28.23-24 (m)

Araeoncus altissimus Pesarini, 1996: 414, f. 1-2, 8 (mf)

Araeoncus altissimus Bosmans, 1996: 146, f. 74-81 (mf)

Araeoncus altissimus IJland, van Helsdingen & Miller, 2012: 10, f. 12-13 (f)

Araeoncus anguineus (L. Koch, 1869) | | France to Bulgaria urn:lsid:nmbe.ch:spidersp:009420

Erigone anguinea L. Koch, 1869: 180 (Dmf) Original description

Araeoncus anguineus Simon, 1884a: 642, f. 474-476 (mf)

Araeoncus anguineus Simon, 1894a: 652, f. 629 (m)

Araeoncus anguineus Schenkel, 1927b: 243, f. 7 (f)

Araeoncus anguineus Wiehle, 1960a: 243, f. 440-445 (mf)

Diplocephalus helleri Wiehle, 1967a: 7, f. 22-24 (f; misidentified per Wunderlich, 1973d: 395)

Araeoncus anguineus Millidge, 1977: 34, f. 125 (m)

Araeoncus anguineus Deltshev, 1985a: 359, f. 2-5 (mf)

Araeoncus anguineus Heimer & Nentwig, 1991: 114, f. 328 (mf)

Diplocephalus helleri Heimer & Nentwig, 1991: 142, f. 400.3-4 (f; misidentified sensu Wunderlich, 1973d: 395)

Araeoncus anguineus Pesarini, 1996: 414, f. 3-4, 7 (mf)

Araeoncus anguineus Breitling, 2020: 344, f. 17A (f)

Araeoncus banias Tanasevitch, 2013 | | Israel urn:lsid:nmbe.ch:spidersp:045834

Araeoncus banias Tanasevitch, 2013a: 103, f. 1-7 (Dmf) Original description

Araeoncus caucasicus Tanasevitch, 1987 | | Ukraine, Caucasus, Iran, Central Asia urn:lsid:nmbe.ch:spidersp:009421

Araeoncus caucasicus Tanasevitch, 1987a: 333, f. 59-64 (Dmf) Original description

Araeoncus caucasicus Tanasevitch, 1990: 111, f. 22.3, 28.21-22 (m)

Araeoncus caucasicus Marusik, Guseinov & Aliev, 2005: 140, f. 10 (f)

Araeoncus caucasicus Tanasevitch, 2013a: 103, f. 8 (f)

Araeoncus caucasicus Tanasevitch, 2013c: 270, f. 17-18, 23-24 (m)

Araeoncus clavatus Tanasevitch, 1987 | | Turkey, Armenia urn:lsid:nmbe.ch:spidersp:009422

Araeoncus clavatus Tanasevitch, 1987a: 334, f. 65-69 (Dm) Original description

Araeoncus clavatus Tanasevitch, 1990: 111, f. 22.4, 28.17-28 (m)

Araeoncus clavatus Karabulut & Türkeş, 2012: 93, f. 3A-G (m, Df)

Araeoncus clivifrons Deltshev, 1987 | | Bulgaria urn:lsid:nmbe.ch:spidersp:009423

Araeoncus clivifrons Deltshev, 1987b: 97, f. 1-5 (Dmf) Original description

Araeoncus convexus Tullgren, 1955 | | Sweden, Estonia urn:lsid:nmbe.ch:spidersp:009424

Araeoncus convexus Tullgren, 1955: 322, f. 25a-b (Df) Original description

Araeoncus crassiceps (Westring, 1861) | | Europe, Russia (Europe to South Siberia) urn:lsid:nmbe.ch:spidersp:009425

Erigone crassiceps Westring, 1861: 231 (Dmf) Original description

Walckenaera affinitata O. Pickard-Cambridge, 1863: 8591 (Dmf) [urn:lsid:nmbe.ch:spidersp:060277]

Walckenaera affinitata O. Pickard-Cambridge, 1873b: 454, pl. 35, f. 30 (m)

Erigone excelsa L. Koch, 1879c: 76, pl. 2, f. 29 (Dm) [urn:lsid:nmbe.ch:spidersp:060099]

Walckenaera crassiceps O. Pickard-Cambridge, 1879d: 151

Araeoncus crassiceps Simon, 1884a: 638, f. 463-466 (m, S of Walckenaeria affinitata)

Araeoncus excelsus Simon, 1884a: 643

Araeoncus crassiceps Smith, 1901a: 334, f. 1t, 2q (m)

Araeoncus crassiceps Bösenberg, 1902: 194, pl. 17, f. 268 (mf)

Araeoncus crassiceps Falconer, 1911: 284, f. 2 (f)

Savignia crassiceps Dahl, 1912b: 609

Araeoncus crassiceps Locket & Millidge, 1953: 298, f. 180C, E, G, 181D (mf)

Araeoncus crassiceps Tullgren, 1955: 320, f. 22a-d (mf)

Araeoncus crassiceps Wiehle, 1960a: 239, f. 432-439 (mf)

Araeoncus crassiceps Miller, 1971: 278, pl. LIX, f. 11-13 (mf)

Araeoncus crassiceps Holm, 1973: 78 (S of Araeoncus excelsus)

Araeoncus crassiceps Palmgren, 1976: 42, f. 15.5-10 (mf)

Araeoncus crassiceps Roberts, 1987: 90, f. 40b, 41c (mf)

Araeoncus crassipes Heimer & Nentwig, 1991: 116, f. 330 (mf)

Araeoncus crassiceps Hormiga, 2000: 28, f. 1A-G (mf)

Araeoncus crassiceps Tanasevitch, 2013c: 270, f. 11-16, 19-22, 25-27 (mf)

Araeoncus curvatus Tullgren, 1955 | | Sweden, Estonia urn:lsid:nmbe.ch:spidersp:009426

Araeoncus curvatus Tullgren, 1955: 321, f. 23a-b (Df) Original description

Araeoncus cypriacus Tanasevitch, 2011 | | Cyprus urn:lsid:nmbe.ch:spidersp:043958

Araeoncus cypriacus Tanasevitch, 2011b: 51, f. 1-5 (Dm) Original description

Araeoncus discedens (Simon, 1882) | | Spain, France, Italy urn:lsid:nmbe.ch:spidersp:009427

Erigone discedens Simon, 1882d: 235 (Dm) Original description

Araeoncus discedens Simon, 1884a: 639, f. 467-470 (m)

Araeoncus discedens Simon, 1926: 368, 492, f. 651 (m, Df)

Araeoncus discedens Bosmans, 1996: 148, f. 82-88 (mf)

Erigone discedens Frétey, 2023: 124 (year of description corrected)

Araeoncus dispar Tullgren, 1955 | | Sweden urn:lsid:nmbe.ch:spidersp:009428

Araeoncus dispar Tullgren, 1955: 323, f. 26a-b (Df) Original description

Araeoncus etinde Bosmans & Jocqué, 1983 | | Cameroon urn:lsid:nmbe.ch:spidersp:009430

Araeoncus etinde Bosmans & Jocqué, 1983: 582, f. 1a-e (Dmf) Original description

Araeoncus femineus (Roewer, 1942) | | Equatorial Guinea (Bioko) urn:lsid:nmbe.ch:spidersp:009431

Gonatium femineum Roewer, 1942b: 247, f. 3 (Df) Original description

Diplocephalus femineus Millidge, 1981d: (T from Gonatium)

Araeoncus femineum Bosmans & Jocqué, 1983: 584 (T from Diplocephalus)

Araeoncus galeriformis (Tanasevitch, 1987) | | Russia (Caucasus), Azerbaijan urn:lsid:nmbe.ch:spidersp:009432

Savignia galeriformis Tanasevitch, 1987a: 361, f. 123-127 (Dmf) Original description

Araeoncus galeriformis Tanasevitch, 1990: 44, f. 22.5, 28.19-20 (mf, T from Savignia)

Araeoncus gertschi Caporiacco, 1949 | | Kenya urn:lsid:nmbe.ch:spidersp:009433

Araeoncus gertschi Caporiacco, 1949a: 371, f. 32 (Df) Original description

Araeoncus gertschi Scharff, 1990b: 144 (misplaced in this genus, incertae sedis, species inquirenda)

Araeoncus hanno Simon, 1884 | | Algeria urn:lsid:nmbe.ch:spidersp:009434

Araeoncus hanno Simon, 1884a: 638 (Dm) Original description

Araeoncus hanno Denis, 1954a: 314, f. 6-7 (Df)

Araeoncus hanno Bosmans, 1996: 127, f. 10-17 (mf)

Araeoncus humilis (Blackwall, 1841) | | Europe, North Africa, Russia (Europe to South Siberia), Iran, Japan. Introduced to New Zealand urn:lsid:nmbe.ch:spidersp:009435 TYPE SPECIES

Walckenaera humilis Blackwall, 1841a: 636 (Dmf) Original description

Argus humilis Walckenaer, 1847a: 506

Walckenaera humilis Blackwall, 1864a: 307, pl. 21, f. 223 (mf)

Lophocarenum globiceps Menge, 1868: 207, pl. 40, f. 204 (Dm) [urn:lsid:nmbe.ch:spidersp:060278]

Erigone humilis Dahl, 1883: 47

Araeoncus humilis Simon, 1884a: 636, f. 460-462 (mf, S of Lophocarenum globiceps)

Diplocephalus humilis Chyzer & Kulczyński, 1894: 111, pl. 4, f. 23 (mf)

Araeoncus humilis Becker, 1896: (3): 126, pl. 11, f. 12 (mf)

Araeoncus humilis Smith, 1901a: 304, p. 334: f. 1s, 2o (m)

Araeoncus humilis Bösenberg, 1902: 195, pl. 17, f. 269 (mf)

Araeoncus humilis Simon, 1926: 367, 492, f. 650 (mf)

Araeoncus humilis Peelle & Saito, 1932: 94, f. 7 (m)

Araeoncus humilis Miller, 1947: 37, pl. VI, f. 6 (f)

Araeoncus humilis Denis, 1948a: 19, f. (m)

Araeoncus humilis Locket & Millidge, 1953: 298, f. 180A-B, D, F, 181A-C (mf)

Araeoncus meridionalis Denis, 1954a: 313, f. 4-5 (Df) [urn:lsid:nmbe.ch:spidersp:060100]

Araeoncus humilis Tullgren, 1955: 319, f. 21a-d (mf)

Araeoncus humilis Saito, 1959: 79, f. 83a-f (m)

Araeoncus humilis Wiehle, 1960a: 235, f. 423-431 (mf)

Araeoncus humilis Merrett, 1963b: 400, f. 65A-C (m)

Araeoncus humilis Tystshenko, 1971: 262, f. 772 (f)

Araeoncus humilis Miller, 1971: 278, pl. LIX, f. 14-15 (mf)

Araeoncus humilis Palmgren, 1976: 42, f. 15.11-13 (mf)

Araeoncus humilis Roberts, 1987: 90, f. 40a, 41b (mf)

Araeoncus humilis Deltshev, 1987b: 101, f. 6-13 (mf)

Araeoncus humilis Millidge, 1988a: 61, f. 293-295, 297-298, 300, 306 (mf)

Araeoncus humilis Heimer & Nentwig, 1991: 114, f. 329 (mf)

Araeoncus humilis Melic, 1994a: 114, f. 6-8 (m)

Araeoncus humilis Bosmans, 1996: 125, f. 1-9 (mf, S of Araeoncus meridionalis)

Araeoncus humilis Hormiga, 2000: 29, f. 2A-C, pl. 13A-F, 14A-D (m)

Araeoncus humilis Ono, Matsuda & Saito, 2009: 277, f. 227-229 (mf)

Araeoncus humilis Frick, Nentwig & Kropf, 2010: 309, Suppl. pl. 3A-O (mf)

Araeoncus hyalinus Song & Li, 2010 | | China urn:lsid:nmbe.ch:spidersp:043214

Araeoncus hyalinus Song & Li, 2010a: 120, f. 1A-E, 2A-G, 3A-E (Dmf) Original description

Araeoncus impolitus Holm, 1962 | | Kenya urn:lsid:nmbe.ch:spidersp:009436

Araeoncus impolitus Holm, 1962: 90, f. 26c, 27c (Df) Original description

Araeoncus longispineus Song & Li, 2010 | | China urn:lsid:nmbe.ch:spidersp:043215

Araeoncus longispineus Song & Li, 2010a: 125, f. 4A-D, 5A-H (Dmf) Original description

Araeoncus longiusculus (O. Pickard-Cambridge, 1875) | | France (Corsica), Italy (incl. Sardinia) urn:lsid:nmbe.ch:spidersp:009437

Erigone longiuscula O. Pickard-Cambridge, 1875b: 192, pl. 27, f. 3 (Dm) Original description

Araeoncus longiusculus Simon, 1884a: 634, f. 458-459 (m, Df)

Araeoncus longiusculus Simon, 1926: 365, 491, f. 647 (mf)

Araeoncus longiusculus Pantini & Sassu, 2009: 625, f. 1-4 (mf)

Araeoncus macrophthalmus Miller, 1970 | | Angola urn:lsid:nmbe.ch:spidersp:009438

Araeoncus macrophthalmus Miller, 1970: 87, pl. IV, f. 1-6 (Dm) Original description

Araeoncus malawiensis Jocqué, 1981 | | Malawi urn:lsid:nmbe.ch:spidersp:009439

Araeoncus malawiensis Jocqué, 1981a: 481, f. 21-23 (Df) Original description

Araeoncus martinae Bosmans, 1996 | | Morocco, Algeria urn:lsid:nmbe.ch:spidersp:009440

Araeoncus martinae Bosmans, 1996: 129, f. 18-26 (Dmf) Original description

Araeoncus mitriformis Tanasevitch, 2008 | | Turkey, Iran urn:lsid:nmbe.ch:spidersp:041997

Araeoncus mitriformis Tanasevitch, 2008b: 473, f. 1-4 (Dm) Original description

Araeoncus mitriformis Tanasevitch, 2009a: 384, f. 7-11 (Df)

Araeoncus mitriformis Danişman & Cosar, 2013: 567, f. 1A-E (mf)

Araeoncus obtusus Bosmans & Jocqué, 1983 | | Cameroon urn:lsid:nmbe.ch:spidersp:009441

Araeoncus obtusus Bosmans & Jocqué, 1983: 586, f. 2a-b (Dm) Original description

Araeoncus picturatus Holm, 1962 | | Tanzania urn:lsid:nmbe.ch:spidersp:009443

Araeoncus picturatus Holm, 1962: 88, f. 26b, 27b (Df) Original description

Araeoncus rhodes Tanasevitch, 2011 | | Greece (Rhodes) urn:lsid:nmbe.ch:spidersp:043959

Araeoncus rhodes Tanasevitch, 2011b: 53, f. 6-12, 20 (Dmf) Original description

Araeoncus sicanus Brignoli, 1979 | | Italy (Sicily) urn:lsid:nmbe.ch:spidersp:009444

Araeoncus sicanus Brignoli, 1979e: 280, f. 9 (Df) Original description

Araeoncus subniger Holm, 1962 | | Kenya urn:lsid:nmbe.ch:spidersp:009445

Araeoncus subniger Holm, 1962: 90, f. 26d, 27d (Df) Original description

Araeoncus subniger Jocqué, 1985a: 277, f. 1-3 (Dm)

Araeoncus tauricus Gnelitsa, 2004 | | Bulgaria, Greece (Crete), Turkey, Ukraine urn:lsid:nmbe.ch:spidersp:039874

Araeoncus tauricus Deltshev, Lazarov & Blagoev, 2004: 193, f. 175 (mf, nomen nudum)

Araeoncus tauricus Gnelitsa, 2004: 81, f. 1-10 (Dmf) Original description

Araeoncus tauricus Danişman & Coşar, 2014: 72, f. 1-2 (m)

Araeoncus tauricus Türkeş et al., 2015: 562, f. 3A-B (f)

Araeoncus toubkal Bosmans, 1996 | | Portugal, Morocco urn:lsid:nmbe.ch:spidersp:009446

Araeoncus altissimus Denis, 1968d: 156, f. 26-31 (mf; misidentified per Bosmans, 1996: 132)

Araeoncus toubkal Bosmans, 1996: 132, f. 27-34 (Dmf) Original description

Araeoncus tuberculatus Tullgren, 1955 | | Sweden urn:lsid:nmbe.ch:spidersp:009447

Araeoncus tuberculatus Tullgren, 1955: 324, f. 27a-b (Df) Original description

Araeoncus vaporariorum (O. Pickard-Cambridge, 1875) | | France, Italy urn:lsid:nmbe.ch:spidersp:009448

Erigone vaporariorum O. Pickard-Cambridge, 1875b: 198, pl. 27, f. 8 (Dm) Original description

Araeoncus vaporariorum Simon, 1884a: 633, f. 455-457 (m, Df)

Araeoncus vaporariorum Smith, 1901a: 334, f. 1u, 2p (m)

Araeoncus vaporariorum Simon, 1926: 366, 491, f. 648 (mf)

Araeoncus vaporariorum Pesarini, 1996: 414, f. 5-6 (m)

Araeoncus victorianyanzae Berland, 1936 | | Kenya, Tanzania urn:lsid:nmbe.ch:spidersp:009449

Araeoncus victorianyanzae Berland in Fage & Simon, 1936: 326, f. 17 (Dmf) Original description

Araeoncus praeceps Holm, 1962: 85, f. 25a-d, 26a, 27a, pl. 3, f. 1 (Dmf) [urn:lsid:nmbe.ch:spidersp:060101]

Araeoncus victorianyanzae Scharff, 1990a: 9, f. 2-6 (m, S of Araeoncus praeceps)

Araeoncus viphyensis Jocqué, 1981 | | Malawi urn:lsid:nmbe.ch:spidersp:009450

Araeoncus viphyensis Jocqué, 1981a: 482, f. 24-26 (Df) Original description

Araeoncus vorkutensis Tanasevitch, 1984 | | Russia (Europe to South Siberia), Kazakhstan urn:lsid:nmbe.ch:spidersp:009451

Araeoncus vorkutensis Tanasevitch, 1984a: 387, pl. 4, f. 1-8 (Dmf) Original description