World Spider Catalog

Linyphiidae Blackwall, 1859

Family | ACCEPTED

N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; Tian et al., 2022 show five subfamilies: Linyphiinae (incl. Mynogleninae Lehtinen, 1967: 250 and Dubiaraneinae Millidge, 1993c: 145), Erigoninae Emerton, 1882: 7, Micronetinae Hull, 1920a: 8, Ipainae Saaristo, 2007b: 34 and Stemonyphantinae Wunderlich, 1986: 120; da Silva-Moreira, Kulkarni & Hormiga, 2025: 60-62 list resp. newly circumscribe six subfamilies: Erigoninae, Lepthyphantinae Simon, 1929: 562, Linyphiinae (syn. nov.: Dubiaraneinae), Mynogleninae, Pocobletinae Silva-Moreira & Hormiga, 2025: 61 (new subfamily) and Stemonyphantinae, state Ipainae and Micronetinae to be polyphyletic, leave several linyphiid clades unranked and also several genera as incertae sedis.

In synonymy:

Anacotyle Simon, 1926 = Styloctetor Simon, 1884 (Marusik & Tanasevitch, 1998: 154)

Gen. Styloctetor Simon, 1884 urn:lsid:nmbe.ch:spidergen:01285

type Styloctetor inuncans Simon, 1884; gender masculine; syn. of Styloctetor romanus (O. Pickard-Cambridge, 1873)

N.B.: removed from the synonymy of Ceratinopsis Emerton, 1882 (rejecting Wunderlich, 1970: 406) and considered a senior synonym of Anacotyle Simon, 1926: 490 [urn:lsid:nmbe.ch:spidergen:03838] (type Erigone stativa Simon, 1881) by Marusik & Tanasevitch, 1998: 154.

Transferred to other genera:

Styloctetor austriacus Kulczyński, 1898 -- see Trichoncoides

Styloctetor cristatopalpus (Ohlert, 1867) -- see Moebelia

Styloctetor machadoi Miller, 1970 -- see Ceratinopsis

Styloctetor penicillatus (Westring, 1851) -- see Moebelia

Styloctetor simplex Kulczyński, 1908 -- see Semljicola

Styloctetor uncinus O. Pickard-Cambridge, 1906 -- see Mecynargus

In synonymy:

Styloctetor asiaticus (Andreeva & Tystshenko, 1970, T from Thyreosthenius) = Styloctetor romanus (O. Pickard-Cambridge, 1873) (Tanasevitch, 1983b: 1786, sub Ceratinopsis)

Styloctetor distinctus (Caporiacco, 1935, T from Alioranus) = Styloctetor romanus (O. Pickard-Cambridge, 1873) (Tanasevitch, 2025b: 118)

Styloctetor duriusculus (Caporiacco, 1935, T from Araeoncus) = Styloctetor romanus (O. Pickard-Cambridge, 1873) (Tanasevitch, 2025b: 118)

Styloctetor incautus (O. Pickard-Cambridge, 1872, T from Lophomma) = Styloctetor romanus (O. Pickard-Cambridge, 1873) (Bosmans, 1994a: 234, sub Ceratinopsis)

Styloctetor inuncans Simon, 1884 = Styloctetor romanus (O. Pickard-Cambridge, 1873) (Simon, 1926: 488)

Styloctetor lehtineni Marusik & Tanasevitch, 1998 = Styloctetor vacerosus (Keyserling, 1886) (Slowik & Marusik, 2025: 300)

Styloctetor lucidus (O. Pickard-Cambridge, 1873, T from Cornicularia) = Styloctetor compar (Westring, 1861) (Bristowe, 1939: 67, sub Anacotyle stativa)

Styloctetor obscurus (Emerton, 1919, T from Ceratinopsis) = Styloctetor compar (Westring, 1861) (Wunderlich, 1970: 406, sub Ceratinopsis stativus)

Styloctetor stativus (Simon, 1882, T from Gongylidiellum) = Styloctetor compar (Westring, 1861) (Breitling et al., 2015: 71)

Styloctetor tauricus (Thorell, 1875, T from Diplocephalus) = Styloctetor romanus (O. Pickard-Cambridge, 1873) (Millidge, 1977: 55, sub Ceratinopsis)

Styloctetor unicolor (Crosby, 1905) = Styloctetor purpurescens (Keyserling, 1886) (Crosby, 1905a: 25, sub Ceratinopsis)

Homonym replaced:

Styloctetor orientalis (Eskov, 1989) -- see Styloctetor okhotensis Eskov, 1993 Marusik et al., 1993a

Styloctetor austerus (L. Koch, 1884) | | Alps (Switzerland, Austria, Italy) urn:lsid:nmbe.ch:spidersp:012775

Erigone austera L. Koch, 1876b: 239 (nomen nudum)

Lophomma austerum L. Koch in Simon, 1884a: 541 (Dm) Original description

Anacotyle austera Simon, 1926: 360

Anacotyle austera Schenkel, 1934: 95, f. 3 (Df)

Styloctetor austerus Thaler, 1970: 266, f. 29-32, 36 (mf, T from Anacotyle)

Ceratinopsis austera Heimer & Nentwig, 1991: 136, f. 383 (mf)

Styloctetor austerus Marusik & Tanasevitch, 1998: 158 (T from Ceratinopsis)

Styloctetor compar (Westring, 1861) | | Europe, Russia (Europe to Far East), Kazakhstan, Alaska, Canada urn:lsid:nmbe.ch:spidersp:010683

Erigone compar Westring, 1861: 286 (Dm) Original description

Neriene lucida O. Pickard-Cambridge, 1873b: 452, pl. 35, f. 27a-e (Dm; N.B.: the f, which Roewer, 1942a: 692 listed under Diplocephalus cristatus, was never described) [urn:lsid:nmbe.ch:spidersp:060661]

Erigone stativa Simon, 1882d: 256 (Dm) [urn:lsid:nmbe.ch:spidersp:012781]

Micryphantes stativus Förster & Bertkau, 1883: 271

Tmeticus compar Simon, 1884a: 420

Gongylidium lucidum Simon, 1884a: 499

Lophomma stativum Simon, 1884a: 540, f. 343 (m)

Micryphantes stativus Bösenberg, 1902: 153, pl. 13, f. 205 (m, Df)

Lophomma stativa O. Pickard-Cambridge, 1906a: 64, pl. A, f. 19-21 (mf)

Centromerus compar Reimoser, 1919: 80

Ceratinopsis obscura Emerton, 1919a: 105, pl. 7, f. 2a-b (Dm) [urn:lsid:nmbe.ch:spidersp:057241]

Anacotyle stativa Simon, 1926: 360, 490, f. 640 (m)

Ceratinopsis obscura Bishop & Crosby, 1930: 25, pl. 5, f. 29-30 (m)

Gongylidiellum compar Hull, 1933: 105, 108

Styloctetor stativus Hull, 1933: 107

Anacotyle stativa Bristowe, 1939: 67 (S of Cornicularia lucida)

Anacotyle stativa Vogelsanger, 1944: 164, f. 4 (f)

Anacotyle stativum Miller, 1947: 26, pl. VI, f. 15-16 (mf)

Anacotyle stativa Locket & Millidge, 1953: 260, f. 158A-C (mf)

Anacotyle stativa Tullgren, 1955: 308, f. 13a-c (m)

Anacotyle stativa Wiehle, 1960a: 250, f. 452-460 (mf)

Styloctetor stativus Merrett, 1963b: 435, f. 116A-C (m, T)

Styloctetor stativus Thaler, 1970: 266, f. 33-35 (m)

Ceratinopsis stativa Wunderlich, 1970: 406 (S of Ceratinopsis obscurus)

Anacotyle stativa Tystshenko, 1971: 262, f. 882 (f)

Anacotyle stativa Miller, 1971: 271, pl. LVI, f. 22-23 (mf)

Anacotyle stativa Palmgren, 1976: 40, f. 18.20-24 (mf)

Ceratinopsis stativa Millidge, 1977: 19, f. 102 (m)

Ceratinopsis stativa Roberts, 1987: 70, f. 29b (mf)

Ceratinopsis stativa Heimer & Nentwig, 1991: 136, f. 385 (mf)

Styloctetor stativus Marusik & Tanasevitch, 1998: 156, f. 5-7, 14 (mf, T from Ceratinopsis)

Styloctetor stativus Paquin & Dupérré, 2003: 121, f. 1265-1267 (mf)

Styloctetor compar Breitling et al., 2015: 71 (T from Gongylidiellum, S of Styloctetor stativus)

Styloctetor compar Lecigne, 2016d: 23, f. 12A-B (m)

Styloctetor logunovi (Eskov & Marusik, 1994) | | Kazakhstan, Iran, Russia (South Siberia), Mongolia urn:lsid:nmbe.ch:spidersp:012777

Ceratinopsis logunovi Eskov & Marusik, 1994: 42, f. 6-9 (Dmf) Original description

Styloctetor logunovi Marusik & Tanasevitch, 1998: 154, f. 1-3, 12 (mf, T from Ceratinopsis)

Styloctetor logunovi Zamani et al., 2020: 583, f. 8A-E (mf)

Styloctetor okhotensis (Eskov, 1993) | | Russia (north-eastern Siberia, Far North East) urn:lsid:nmbe.ch:spidersp:012778

Ceratinopsis orientalis Eskov, 1989a: 106, f. 59-62 (Dmf; preoccupied by Locket, 1982, see Ceratinopsis orientalis) [urn:lsid:nmbe.ch:spidersp:055609]

Ceratinopsis okhotensis Marusik et al., 1993b: 74 (replacement name) Original description

Styloctetor okhotensis Marusik & Tanasevitch, 1998: 158, f. 13, 19 (mf, T from Ceratinopsis)

Styloctetor purpurescens (Keyserling, 1886) | | Canada, USA urn:lsid:nmbe.ch:spidersp:012779

Erigone purpurescens Keyserling, 1886b: 187, pl. 17, f. 248 (Dm; f f. 248a see Tutaibo anglicanus) Original description

Ceratinopsis unicolor Crosby, 1905a: 341 (Dm) [urn:lsid:nmbe.ch:spidersp:062902]

Ceratinopsis purpurescens Bishop & Crosby, 1930: 25, pl. 5, f. 31-35 (m, Df, S of Ceratinopsis unicolor)

Ceratinopsis purpurescens Muma, 1943: 31, pl. XI, f. 12-13 (mf)

Styloctetor purpurescens Marusik & Tanasevitch, 1998: 156 (T from Ceratinopsis)

Styloctetor romanus (O. Pickard-Cambridge, 1873) | | Europe, North Africa, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, Iran, Central Asia, Pakistan, China urn:lsid:nmbe.ch:spidersp:012780 TYPE SPECIES

Erigone incauta O. Pickard-Cambridge, 1872: 289 (Dm; suppressed for lack of usage) [urn:lsid:nmbe.ch:spidersp:057240]

Erigone romana O. Pickard-Cambridge, 1873a: 752, pl. 65, f. 6 (Dmf) Original description

Erigone taurica Thorell, 1875b: 63 (Dm) [urn:lsid:nmbe.ch:spidersp:057242]

Erigone taurica Thorell, 1875c: 39 (m)

Plaesiocraerus tauricus Simon, 1884a: 777

Lophomma incautum Simon, 1884a: 538

Styloctetor inuncans Simon, 1884a: 735, f. 621-623 (Dmf) [urn:lsid:nmbe.ch:spidersp:062927]

Styloctetor romanus Chyzer & Kulczyński, 1894: 96, pl. 4, f. 7 (mf)

Micryphantes inuncans Bösenberg, 1902: 153, pl. 13, f. 204 (Df)

Styloctetor romanus Simon, 1926: 357, 488, f. 629-631 (mf, S of Styloctetor inuncans)

Alioranus distinctus Caporiacco, 1935b: 177, pl. 2, f. 8 (Dm) [urn:lsid:nmbe.ch:spidersp:009375]

Araeoncus duriusculus Caporiacco, 1935b: 180, pl. 1, f. 19 (Df) [urn:lsid:nmbe.ch:spidersp:009429]

Styloctetor romanus Miller, 1947: 37, pl. V, f. 19-23 (mf)

Styloctetor romanus Locket & Millidge, 1953: 258, f. 157A-C (mf)

Styloctetor romanus Wiehle, 1960a: 254, f. 461-467 (mf)

Styloctetor romanus Merrett, 1963b: 435, f. 115A-B (m)

Thyreostenius asiaticus Andreeva & Tystshenko, 1970: 40, f. 1 (Df) [urn:lsid:nmbe.ch:spidersp:057239]

Styloctetor romanus Tystshenko, 1971: 264, f. 883, 886 (mf)

Styloctetor romanus Miller, 1971: 276, pl. LVIII, f. 8-9 (mf)

Thyreosthenius asiaticus Andreeva, 1976: 63, f. 69-71 (f)

Ceratinopsis romana Millidge, 1977: 55 (S of Diplocephalus tauricus)

Ceratinopsis romana Tanasevitch, 1983b: 1786 (S of Thyreosthenius asiaticus)

Ceratinopsis romana Roberts, 1987: 70, f. 29a (mf)

Sphecozone romana Wunderlich, 1987a: 172 (T from Ceratinopsis; not accepted by later authors)

Ceratinopsis romana Tanasevitch, 1990: 104, f. 22.49, 23.35 (mf)

Ceratinopsis romana Heimer & Nentwig, 1991: 136, f. 384 (mf)

Ceratinopsis romana Bosmans, 1994a: 234 (S of Lophomma incautum)

Styloctetor romanus Marusik & Tanasevitch, 1998: 154, f. 4, 11 (mf, T from Ceratinopsis)

Ceratinopsis romana Xia et al., 2002: 80, f. 1-6 (mf)

Styloctetor romanus Bosmans, 2007: 139, f. 148-152 (mf)

Styloctetor romanus Türkeş et al., 2015: 563, f. 6A-B (f)

Styloctetor romanus Wunderlich, 2023: 39, f. 57-60 (mf)

Styloctetor romanus Tanasevitch, 2025b: 118 (S of Alioranus distinctus and Araeoncus duriusculus)

Styloctetor tuvinensis Marusik & Tanasevitch, 1998 | | Russia (South Siberia) urn:lsid:nmbe.ch:spidersp:012782

Styloctetor tuvinensis Marusik & Tanasevitch, 1998: 158, f. 15-18 (Dm) Original description

Styloctetor vacerosus (Keyserling, 1886) | | Russia (Urals to Far East), Kazakhstan, Alaska urn:lsid:nmbe.ch:spidersp:057665

Erigone vacerosa Keyserling, 1886b: 200, pl. 18, f. 260 (Df) Original description

Erigone vacerosa Roewer, 1955c: 1503 (nicht zu deuten = nomen dubium)

Styloctetor lehtineni Marusik & Tanasevitch, 1998: 158, f. 8-10, 20 (Dmf) [urn:lsid:nmbe.ch:spidersp:012776]

Styloctetor lehtineni Nekhaeva, Kim & Yeszhanov, 2024: 400, f. 7B-C (m)

Styloctetor vacerosus Slowik & Marusik, 2025: 300, f. 1E (f, T from Erigone, removed from nomen dubium, S of Styloctetor lehtineni)