Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Leptodrassus Simon, 1878 urn:lsid:nmbe.ch:spidergen:02443

Transferred to other genera:

Leptodrassus algericus Dalmas, 1919 -- see Leptodrassex

Leptodrassus hadjissaranti Chatzaki, 2002 -- see Leptopilos

Leptodrassus hylaestomachi Berland, 1934 -- see Leptodrassex

Leptodrassus kalbicus Savelyeva, 1972 -- see Agroeca

Leptodrassus leclerci Denis, 1966 -- see Leptopilos

Leptodrassus manolisi Chatzaki, 2002 -- see Leptopilos

Leptodrassus memorialis Spassky, 1940 -- see Leptodrassex

Leptodrassus pupa Dalmas, 1919 -- see Leptopilos

Leptodrassus simoni Dalmas, 1919 -- see Leptodrassex

Leptodrassus tenerrimus (O. Pickard-Cambridge, 1872) -- see Leptopilos

Nomen nudum:

Leptodrassus japonicus Kishida -- Brignoli, 1983c: 564

Leptodrassus albidus Simon, 1914 | | Azores, Canary Is., Spain to Greece, Turkey, Israel urn:lsid:nmbe.ch:spidersp:027453
Leptodrassus albidus Simon, 1914: 142, 210, f. 255-256 (Dmf) Original description
Leptodrassus albidus Dalmas, 1919b: 244, f. 3-4 (mf)
Leptodrassus albidus Di Franco & Pantini, 2000: 479, f. 1-2 (m)
Leptodrassus albidus Chatzaki, Thaler & Mylonas, 2002a: 589, f. 64-65, 72 (mf)
Leptodrassus albidus Borges & Wunderlich, 2008: 264, f. 18a-d (f)
Leptodrassus albidus Levy, 2009: 8, f. 15-16 (f)
Leptodrassus albidus Seyyar & Demir, 2010: 15, f. 1A-D (m)
Leptodrassus albidus Komnenov, 2014: 166, f. 32-33 (m)
Leptodrassus albidus Suárez, 2018: 61, f. 2a-b (f)
Leptodrassus albidus Naumova, Blagoev & Deltshev, 2021: 235, f. 6A-C (m)
Leptodrassus bergensis Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:027455
Leptodrassus bergensis Tucker, 1923: 317, pl. 9, f. 43 (Df) Original description
Leptodrassus bergensis Dippenaar-Schoeman et al., 2021C: 22, 1 f. (f)
Leptodrassus croaticus Dalmas, 1919 | | Croatia urn:lsid:nmbe.ch:spidersp:027456
Leptodrassus femineus Chyzer & Kulczyński, 1897: 221, pl. 8, f. 35 (f, misidentified)
Leptodrassus croaticus Dalmas, 1919b: 244 (Df) Original description
Leptodrassus diomedeus Caporiacco, 1951 | | Italy urn:lsid:nmbe.ch:spidersp:027457
Leptodrassus diomedeus Caporiacco, 1951a: 84, f. 7a-b (Dm) Original description
Leptodrassus femineus (Simon, 1873) | | Portugal to Bulgaria and Greece, Cyprus, Israel urn:lsid:nmbe.ch:spidersp:027458
Drassus femineus Simon, 1873a: 165 (Dmf) Original description
Leptodrassus femineus Simon, 1878a: 209, pl. 15, f. 3 (mf)
Leptodrassus femineus Simon, 1893a: 355, f. 323
Leptodrassus femineus Simon, 1914: 142, 210, f. 254, 257 (mf)
Leptodrassus femineus Dalmas, 1919b: 243, f. 1-2 (mf)
Leptodrassus femineus Di Franco, 1987: 144, f. 6-7 (m)
Leptodrassus femineus Di Franco, 2001: 201, f. 1-2 (f)
Leptodrassus femineus Chatzaki, Thaler & Mylonas, 2002a: 590, f. 66-67, 73 (f)
Leptodrassus femineus Murphy, 2007: 61, f. 504-505 (mf)
Leptodrassus femineus Levy, 2009: 7, f. 8-14 (mf)
Leptodrassus fragilis Dalmas, 1919 | | Algeria, Libya urn:lsid:nmbe.ch:spidersp:027459
Leptodrassus fragilis Dalmas, 1919b: 246 (Dm) Original description
Leptodrassus incertus Banks, 1898 | | Mexico urn:lsid:nmbe.ch:spidersp:041299
Leptodrassus incertus Banks, 1898b: 216, pl. 13, f. 10 (Df) Original description
Leptodrassus incertus Ubick & Roth, 1973b: 3 (belongs to a not specified genus of Clubionidae)
Leptodrassus licentiosus Dalmas, 1919 | | South Africa urn:lsid:nmbe.ch:spidersp:027462
Leptodrassus licentiosus Dalmas, 1919b: 248, f. 10 (Df) Original description
Leptodrassus licentiosus Dippenaar-Schoeman et al., 2021C: 23, 1 f. (f)
Leptodrassus punicus Dalmas, 1919 | | Tunisia urn:lsid:nmbe.ch:spidersp:027464
Leptodrassus punicus Dalmas, 1919b: 247, f. 8 (Df) Original description
Leptodrassus strandi Caporiacco, 1947 | | Ethiopia urn:lsid:nmbe.ch:spidersp:027467
Leptodrassus strandi Caporiacco, 1947c: 180, pl. 2, f. 36 (Df) Original description
Leptodrassus tropicus Dalmas, 1919 | j | Sierra Leone urn:lsid:nmbe.ch:spidersp:027469
Leptodrassus tropicus Dalmas, 1919b: 247 (Dj) Original description

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