Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Leptodrassex Murphy, 2007 urn:lsid:nmbe.ch:spidergen:03793
N.B.: see Murphy, 2007: f. 514-517 for figures of two species from Kenya.

Transferred to other genera:

Leptodrassex memorialis (Spassky, 1940) -- see Leptopilos

Leptodrassex algericus (Dalmas, 1919) | | Algeria, Libya urn:lsid:nmbe.ch:spidersp:027454
Leptodrassus algericus Dalmas, 1919b: 245, f. 7 (Dmf) Original description
Leptodrassex algericus Murphy, 2007: 63, f. 519-520 (m chelicera and eyes, T from Leptodrassus)
Micaria sp. Haddad & Bosmans, 2013: 398 (f) [misidentified per Chatzaki & Russell-Smith, 2017: 243]
Leptodrassex algericus Chatzaki & Russell-Smith, 2017: 243, f. 12C-D (f, N.B.: 12C is paralectotype of Castanilla marchesii)
Leptodrassex capensis Haddad & Booysen, 2022 | | South Africa urn:lsid:nmbe.ch:spidersp:055865
Leptodrassex capensis Haddad & Booysen, 2022: 16, f. 45, 80-81 (Df) Original description
Leptodrassex hylaestomachi (Berland, 1934) | | Spain (Canary Is., Balearic Is.), Portugal (Savage Is.) urn:lsid:nmbe.ch:spidersp:027460
Leptodrassus hylaestomachi Berland, 1934c: 423, f. 1-4 (Df) Original description
Leptodrassus hylaestomachi Denis, 1941a: 109, f. 1 (f)
Leptodrassus hylaestomachi Schmidt, 1977: 59, f. 8 (Dm)
Leptodrassex hylaestomachi Murphy, 2007: 63, f. 519-520 (m chelicera and eyes, T from Leptodrassus)
Leptodrassex hylaestomachi Crespo et al., 2009b: 344, f. 2 (f)
Leptodrassex hylaestomachi Wunderlich, 2011: 38, f. 154-157 (mf)
Leptodrassex hylaestomachi Febrer & Barrientos, 2023: 248, f. 7A-D (f)
Leptodrassex murphyi Haddad & Booysen, 2022 | | Mozambique, South Africa urn:lsid:nmbe.ch:spidersp:055864
Leptodrassex murphyi Haddad & Booysen, 2022: 17, f. 46-79, 82-86 (Dmf) Original description
Leptodrassex simoni (Dalmas, 1919) | | Portugal, Spain, France, Lebanon urn:lsid:nmbe.ch:spidersp:027466
Leptodrassus tenerrimus Simon, 1914: 141, 210, f. 252-253 (Dmf, preoccupied)
Leptodrassus simoni Dalmas, 1919b: 245, f. 5-6 (replacement name) Original description
Leptodrassus masculinus Strand, 1942: 397 (superfluous replacement name)
Leptodrassus simoni Levy, 1999d: 446, f. 35-37 (mf)
Leptodrassex simoni Murphy, 2007: 63, f. 512-513 (Tmf from Leptodrassus)
Leptodrassex simoni Wunderlich, 2011: 38, f. 74-76 (mf)

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