Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Leptopilos Levy, 2009 urn:lsid:nmbe.ch:spidergen:03887

In synonymy:

Leptopilos leclerci (Denis, 1966, T from Leptodrassus) = Leptopilos tenerrimus O. Pickard-Cambridge, 1872 (Levy, 2009: 9)

Leptopilos butleri Haddad & Booysen, 2022 | | Botswana, Zimbabwe, South Africa urn:lsid:nmbe.ch:spidersp:055866
Leptopilos butleri Haddad & Booysen, 2022: 23, f. 87-88, 121-126 (Dmf) Original description
Leptopilos digitus Haddad & Booysen, 2022 | | South Africa urn:lsid:nmbe.ch:spidersp:055867
Leptopilos digitus Haddad & Booysen, 2022: 26, f. 89-90, 93-120, 126-130 (Dmf) Original description
Leptopilos hadjissaranti (Chatzaki, 2002) | | Greece (Crete) urn:lsid:nmbe.ch:spidersp:038064
Leptodrassus hadjissaranti Chatzaki in Chatzaki, Thaler & Mylonas, 2002a: 590, f. 68-69, 76-77 (Dmf) Original description
Leptopilos hadjissaranti Levy, 2009: 9 (Tmf from Leptodrassus)
Leptopilos lakhish Levy, 2009 | | Israel urn:lsid:nmbe.ch:spidersp:042292
Leptopilos lakhish Levy, 2009: 13, f. 27-28 (Dm) Original description
Leptopilos levantinus Levy, 2009 | | Greece, Israel, Iran urn:lsid:nmbe.ch:spidersp:042291
Leptodrassus pupa Chatzaki, Thaler & Mylonas, 2002a: 592, f. 74-75 (f, misidentified)
Leptodrassus pupa Chatzaki, Thaler & Mylonas, 2002b: 618, f. 44-45 (m, misidentified)
Leptopilos levantinus Levy, 2009: 10, f. 19-22 (Dmf) Original description
Leptopilos manolisi (Chatzaki, 2002) | | Greece (Crete), Turkey, Israel urn:lsid:nmbe.ch:spidersp:038065
Leptodrassus manolisi Chatzaki in Chatzaki, Thaler & Mylonas, 2002a: 592, f. 70-71 (Dm) Original description
Leptopilos manolisi Levy, 2009: 11, f. 23-26 (Tm from Leptodrassus, Df)
Leptopilos manolisi Wunderlich, 2011: 38, f. 158-159 (mf)
Leptopilos manolisi Akar & Türkeş, 2022: 27, f. 2a-c (m)
Leptopilos memorialis (Spassky, 1940) | | Bulgaria, Greece, Ukraine, Russia (Europe to Central Asia), Kazakhstan, Pakistan, Mongolia urn:lsid:nmbe.ch:spidersp:027463
Leptodrassus memorialis Spassky, 1940: 355, pl. 7, f. 2-5 (Dmf) Original description
Leptodrassus nemoralis Eskov & Marusik, 1995: 77, f. 45-47 (mf)
Leptodrassus memorialis Tuneva & Esyunin, 2002: 218, f. 7-10 (mf)
Leptodrassex memorialis Kovblyuk & Nadolny, 2010: 192, f. 20-39 (Tmf from Leptodrassus)
Leptodrassex memorialis Wunderlich, 2011: 38, f. 157a (m)
Leptodrassex memorialis Marusik & Kovblyuk, 2011: 156, f. 17.18, 22 (mf)
Leptodrassex memorialis Marusik et al., 2014b: 418, f. 2, 12-14 (f)
Leptodrassex memorialis van Helsdingen, IJland & Komnenov, 2018: 14, f. 4 (f)
Leptopilos memorialis Chatzaki & Van Keer, 2019: 444 (T from Leptodrassex)
Leptopilos memorialis Naumova, Blagoev & Deltshev, 2021: 235, f. 7A-E (mf)
Leptopilos pupa (Dalmas, 1919) | | Egypt urn:lsid:nmbe.ch:spidersp:027465
Leptodrassus pupa Dalmas, 1919b: 248, f. 9 (Dmf) Original description
Leptodrassus pupa Levy, 1999d: 446, f. 38-40 (mf)
Leptopilos pupa Levy, 2009: 9 (Tmf from Leptodrassus)
Leptopilos tenerrimus (O. Pickard-Cambridge, 1872) | | Libya, Israel urn:lsid:nmbe.ch:spidersp:027468
Drassus tenerrimus O. Pickard-Cambridge, 1872: 233, pl. 15, f. 10 (Dm) Original description
Leptodrassus tenerrimus Dalmas, 1919b: 247
Leptodrassus leclerci Denis, 1966e: 105, f. 3 (Df) [urn:lsid:nmbe.ch:spidersp:027461]
Leptodrassus tenerrimus Levy, 1999d: 445, f. 32-34 (m)
Leptopilos tenerrimus Levy, 2009: 9, f. 17-18 (mf, T from Leptodrassus, S of Leptodrassus leclerci)
Leptopilos vasivulva Haddad & Booysen, 2022 | | Botswana, Zimbabwe, South Africa urn:lsid:nmbe.ch:spidersp:055868
Leptopilos vasivulva Haddad & Booysen, 2022: 23, f. 91-92, 131-135 (Dmf) Original description

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