World Spider Catalog

Gnaphosidae Banks, 1892

Family | ACCEPTED

N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.

In synonymy:

Phoeodrassus Lohmander, 1944 = Haplodrassus Chamberlin, 1922 (Lohmander, 1944: 13, as subgenus)

Tuvadrassus Marusik & Logunov, 1995 = Haplodrassus Chamberlin, 1922 (Murphy, 2007: 9)

Gen. Haplodrassus Chamberlin, 1922 urn:lsid:nmbe.ch:spidergen:02429

N.B.: considered a senior synonym of Tuvadrassus Marusik & Logunov, 1995b: 194 [urn:lsid:nmbe.ch:spidergen:02493] (type Drassodes tegulatus Schenkel, 1963) by Murphy, 2007: x, 9; Phoeodrassus Lohmander, 1944: 13 [urn:lsid:nmbe.ch:spidergen:06023] (type Drassus cognatus Westring, 1861) was described as subgenus of Haplodrassus; see note under Aracus Thorell, 1887.

Transferred to other genera:

Haplodrassus grazianoi Caporiacco, 1948 -- see Berinda

Haplodrassus magister Chamberlin, 1933 -- see Urozelotes

Haplodrassus severus (C. L. Koch, 1839) -- see Drassodes

Haplodrassus viveki Gajbe, 1992 -- see Drassodes

In synonymy:

Haplodrassus acrotirius (Roewer, 1928) = Haplodrassus omissus O. Pickard-Cambridge, 1872 (Bosmans et al., 2018: 25)

Haplodrassus admes Chamberlin, 1922 = Haplodrassus bicornis Emerton, 1909 (Platnick & Shadab, 1975b: 17)

Haplodrassus altanus Chamberlin, 1933 = Haplodrassus stuxbergi L. Koch, 1879 (Ubick & Roth, 1973a: 4, sub Haplodrassus hiemalis)

Haplodrassus buddebergi (Simon, 1887) = Haplodrassus cognatus Westring, 1861 (Simon, 1914: 208, sub Drassodes capnodes)

Haplodrassus capnodes (Thorell, 1875) = Haplodrassus cognatus Westring, 1861 (Grimm, 1985: 134)

Haplodrassus carinatus (Strand, 1906, T from Drassodes) = Haplodrassus dalmatensis L. Koch, 1866 (Chatzopoulou & Chatzaki, 2009: 358)

Haplodrassus clavator (O. Pickard-Cambridge, 1860) = Haplodrassus signifer C. L. Koch, 1839 (O. Pickard-Cambridge, 1872: 232, sub Drassus troglodytes)

Haplodrassus cognatus ermolajewi Lohmander, 1942 = Haplodrassus cognatus Westring, 1861 (Lohmander, 1942: 86, only provisional name)

Haplodrassus corticalis (Lucas, 1846, T from Drassodes) = Haplodrassus rufipes Lucas, 1846 (Bosmans et al., 2018: 33)

Haplodrassus criminalis (O. Pickard-Cambridge, 1875) = Haplodrassus silvestris Blackwall, 1833 (Smith, 1900a: 330, sub Drassus)

Haplodrassus dalmatensis pictus (Thorell, 1875) = Haplodrassus dalmatensis L. Koch, 1866 (Simon, 1909b: 15, sub Drassodes)

Haplodrassus deceptus (Banks, 1900, T from Zelotes) = Haplodrassus signifer C. L. Koch, 1839 (Chamberlin, 1922: 163)

Haplodrassus delinquens (O. Pickard-Cambridge, 1875) = Haplodrassus dalmatensis L. Koch, 1866 (F. O. Pickard-Cambridge, 1895b: 38, sub Drassus)

Haplodrassus denotatus (O. Pickard-Cambridge, 1874, T from Drassodes) = Haplodrassus dalmatensis L. Koch, 1866 (Levy, 2004: 24)

Haplodrassus desertorum (Caporiacco, 1928, T from Pseudodrassus) = Haplodrassus dalmatensis L. Koch, 1866 (Levy, 2004: 24)

Haplodrassus dubius (Wider, 1834, T from Filistata) = Haplodrassus silvestris Blackwall, 1833 (Zonstein & Marusik, 2019b: 83)

Haplodrassus dystactus Chamberlin & Gertsch, 1940 = Haplodrassus signifer C. L. Koch, 1839 (Platnick & Shadab, 1975b: 11)

Haplodrassus ferrumequinum (F. O. Pickard-Cambridge, 1899, T from Drassodes) = Haplodrassus signifer C. L. Koch, 1839 (Platnick & Shadab, 1975b: 11)

Haplodrassus gridellii Caporiacco, 1949 = Haplodrassus rufipes Lucas, 1846 (Bosmans et al., 2018: 33)

Haplodrassus hiemalis (Emerton, 1909) = Haplodrassus stuxbergi L. Koch, 1879 (Mikhailov & Marusik, 2023: 226)

Haplodrassus infuscatus (Westring, 1861, T from Drassus) = Haplodrassus silvestris Blackwall, 1833 (Simon, 1893a: 361, sub Drassodes)

Haplodrassus insularis Denis, 1962 = Haplodrassus dalmatensis L. Koch, 1866 (Schmidt, 1977: 58)

Haplodrassus isaevi Ponomarev & Tsvetkov, 2006 = Haplodrassus orientalis L. Koch, 1866 (Bosmans et al., 2018: 61)

Haplodrassus lithobius (Roewer, 1928, T from Drassodes) = Haplodrassus dalmatensis L. Koch, 1866 (Chatzaki, Thaler & Mylonas, 2002a: 587)

Haplodrassus macellinus hebes (O. Pickard-Cambridge, 1874) = Haplodrassus macellinus Thorell, 1871 (Bosmans et al., 2018: 54)

Haplodrassus magnipalpus Paik & Sohn, 1984 = Haplodrassus kulczynskii Lohmander, 1942 (Paik & Kang, 1988: 63)

Haplodrassus maroccanus Denis, 1956 = Haplodrassus omissus O. Pickard-Cambridge, 1872 (Bosmans & Van Keer, 2012a: 11, sub H. parvicorpus, contra Levy, 2004: 24, who considered it a S of H. dalmatensis)

Haplodrassus minusculus (L. Koch, 1866, T from Drassus) = Haplodrassus dalmatensis L. Koch, 1866 (Simon, 1909b: 15, sub Drassodes)

Haplodrassus mysticus (O. Pickard-Cambridge, 1895) = Haplodrassus signifer C. L. Koch, 1839 (Bristowe, 1939: 17, sub Drassodes)

Haplodrassus nigriceps (Banks, 1895, T from Syrisca) = Haplodrassus signifer C. L. Koch, 1839 (Chamberlin, 1922: 163)

Haplodrassus nigroscriptus deminutus (Simon, 1909, T from Drassodes) = Haplodrassus nigroscriptus Simon, 1909 (Bosmans et al., 2018: 22)

Haplodrassus nyctelius (Thorell, 1871) = Haplodrassus macellinus Thorell, 1871 (Simon, 1878a: 178, sub Drassus)

Haplodrassus occidentalis (Thorell, 1871) = Haplodrassus macellinus Thorell, 1871 (Simon, 1914: 209, sub Drassodes)

Haplodrassus palaestinensis (Strand, 1915, T from Drassodes) = Haplodrassus dalmatensis L. Koch, 1866 (Levy, 2004: 24)

Haplodrassus parvicorpus (Roewer, 1951) = Haplodrassus omissus O. Pickard-Cambridge, 1872 (Bosmans et al., 2018: 25)

Haplodrassus placidus (Banks, 1896, T from Drassus) = Haplodrassus signifer C. L. Koch, 1839 (Chamberlin, 1922: 163)

Haplodrassus pseudopugnans (Schenkel, 1963) = Haplodrassus pugnans Simon, 1880 (Song & Hubert, 1983: 17, sub Drassodes)

Haplodrassus robustus (Emerton, 1890, T from Drassodes) = Haplodrassus signifer C. L. Koch, 1839 (Chamberlin, 1922: 163)

Haplodrassus seditiosus (Caporiacco, 1928) = Haplodrassus omissus O. Pickard-Cambridge, 1872 (Bosmans et al., 2018: 25)

Haplodrassus similis (L. Koch, 1866, removed from S of Haplodrassus = Drassodes severus) = Haplodrassus rufipes Lucas, 1846 (Bosmans et al., 2018: 33)

Haplodrassus simplex (Kulczyński, 1926, T from Drassodes) = Haplodrassus moderatus Kulczyński, 1897 (Mikhailov & Marusik, 2023: 226)

Haplodrassus troglodytes (C. L. Koch, 1839, T from Drassus) = Haplodrassus signifer C. L. Koch, 1839 (Simon, 1914: 209, sub Drassodes)

Haplodrassus uncifer Chamberlin, 1936 = Haplodrassus bicornis Emerton, 1909 (Platnick & Shadab, 1975b: 17)

Haplodrassus utus Chamberlin & Ivie, 1946 = Haplodrassus eunis Chamberlin, 1922 (Platnick & Shadab, 1975b: 20)

Haplodrassus vicarius (Simon, 1878) = Haplodrassus macellinus Thorell, 1871 (Simon, 1914: 209, sub Drassodes macellinus hebes)

Haplodrassus vignai Di Franco, 1996 = Haplodrassus macellinus Thorell, 1871 (Bosmans et al., 2018: 54, contra Levy, 2004: 31, who considered it a S of H. invalidus))

Haplodrassus walteri (Roewer, 1928, T from Scotophaeus) = Haplodrassus creticus Roewer, 1928 (Levy, 2004: 28)

Nomina dubia:

Haplodrassus barberi (Banks, 1902a: 212, pl. 7, f. 7, 11, f, USA, originally in Prosthesima) -- Platnick & Shadab, 1975b: 5

Haplodrassus pacificus (Banks, 1904b: 336, pl. 39, f. 15, m, USA, originally in Zelotes) -- Platnick & Shadab, 1975b: 5

Haplodrassus spinicrus (Caporiacco, 1928: 84, j, Libya, originally in Drassodes) -- Bosmans et al., 2018: 44

Haplodrassus aenus Thaler, 1984 | | Switzerland, Austria urn:lsid:nmbe.ch:spidersp:027318

Haplodrassus aenus Thaler, 1984a: 189, f. 1-8, 9a-c, g-h (Dmf) Original description

Haplodrassus aeneus Grimm, 1985: 132, f. 149, 172-173 (mf, lapsus)

Haplodrassus aeneus Heimer & Nentwig, 1991: 426, f. 1126 (mf, lapsus)

Haplodrassus alexeevi Ponomarev & Shmatko, 2017 | | Russia (Europe) urn:lsid:nmbe.ch:spidersp:049538

Haplodrassus alexeevi Ponomarev & Shmatko, 2017: 11, f. 1, 3-4, 8, 11-12 (Dmf) Original description

Haplodrassus ambalaensis Gajbe, 1992 | | India urn:lsid:nmbe.ch:spidersp:027319

Haplodrassus ambalaensis Gajbe, 1992a: 305, f. 6-10 (Df) Original description

Haplodrassus atarot Levy, 2004 | | Israel urn:lsid:nmbe.ch:spidersp:040608

Haplodrassus atarot Levy, 2004: 33, f. 74-75 (Dm) Original description

Haplodrassus atarot Levy, 2009: 5, f. 6-7 (Df)

Haplodrassus belgeri Ovtsharenko & Marusik, 1988 | | Russia (South to north-east Siberia, Far East) urn:lsid:nmbe.ch:spidersp:027320

Haplodrassus belgeri Ovtsharenko & Marusik, 1988: 213, f. 45, 47, 50, 53 (Dmf) Original description

Haplodrassus belgeri Ovtsharenko & Marusik, 1996: 121, pl. 2, f. 11-12 (mf)

Haplodrassus bengalensis Gajbe, 1992 | | India urn:lsid:nmbe.ch:spidersp:037337

Haplodrassus bengalensis Gajbe, 1992a: 306, f. 11-15 (Df) Original description

Haplodrassus bicornis (Emerton, 1909) | | Canada, USA urn:lsid:nmbe.ch:spidersp:027321

Drassus bicornis Emerton, 1909: 218, pl. 9, f. 2 (Dmf) Original description

Haplodrassus bicornis Chamberlin, 1922: 161

Haplodrassus admes Chamberlin, 1922: 162 (Df) [urn:lsid:nmbe.ch:spidersp:058449]

Haplodrassus uncifer Chamberlin, 1936a: 18, f. 17-19 (Dmf) [urn:lsid:nmbe.ch:spidersp:058456]

Haplodrassus bicornis Levi & Field, 1954: 458, f. 67, 75-76 (mf)

Haplodrassus bicornis Platnick & Shadab, 1975b: 16, f. 23-30 (mf, S of Haplodrassus admes and H. uncifer)

Haplodrassus bicornis Platnick & Dondale, 1992: 220, f. 330-333 (mf)

Haplodrassus bicornis Paquin & Dupérré, 2003: 78, f. 673-675 (mf)

Haplodrassus bohemicus Miller & Buchar, 1977 | | Czechia, Slovakia, Austria, Hungary, Serbia, North Macedonia, Bulgaria, Greece, Ukraine, Russia (Europe, Caucasus) urn:lsid:nmbe.ch:spidersp:027322

Haplodrassus bohemicus Miller & Buchar, 1977: 163, pl. II, f. 1-6 (Dmf) Original description

Haplodrassus bohemicus Stefanovska et al., 2008: 37, f. 10-15 (mf)

Haplodrassus bohemicus Kovblyuk, Kastrygina & Omelko, 2012: 62, f. 1-6, 10-12 (mf; probably belongs to another species per Bosmans et al., 2018: 50)

Haplodrassus bohemicus Bosmans et al., 2018: 50, f. 198-204 (mf)

Haplodrassus bohemicus Grbić, Hänggi & Krnjajić, 2021: 26, f. 6A-D (mf)

Haplodrassus bohemicus Ponomarev et al., 2021: 359, f. 17-20 (mf)

Haplodrassus bohemicus Šestáková et al., 2022: 17, f. 6a-e, 7a-d, 8a-e, 9a-c, 10a-d (mf)

Haplodrassus canariensis Schmidt, 1977 | | Canary Is. urn:lsid:nmbe.ch:spidersp:027323

Haplodrassus canariensis Schmidt, 1977: 58, f. 5-6 (Dmf) Original description

Haplodrassus caspius Ponomarev & Belosludtsev, 2008 | | Greece, Russia (Europe, Caucasus), Azerbaijan, Iran, Kazakhstan urn:lsid:nmbe.ch:spidersp:042161

Haplodrassus caspius Ponomarev & Belosludtsev in Ponomarev, Belosludtsev & Dvadnenko, 2008: 164, f. 7-8 (Df) Original description

Haplodrassus caspius Kovblyuk et al., 2013b: 67, f. 1-11 (f, Dm)

Haplodrassus caspius Zamani & Marusik, 2018c: 70, f. 2A, 3F (f)

Haplodrassus caspius Chatzaki, 2021: 385, f. 10A-C (m)

Haplodrassus caucasius Ponomarev & Dvadnenko, 2013 | | Caucasus (Russia, Georgia) urn:lsid:nmbe.ch:spidersp:046078

Haplodrassus caucasius Ponomarev & Dvadnenko, 2013: 51, f. 1a-b, 2a-b, 3a-c, 4a-b (Dmf) Original description

Haplodrassus chamberlini Platnick & Shadab, 1975 | | North America urn:lsid:nmbe.ch:spidersp:027324

Haplodrassus chamberlini Platnick & Shadab, 1975b: 27, f. 63-70 (Dmf) Original description

Haplodrassus chamberlini Platnick & Dondale, 1992: 213, f. 317-321 (mf)

Haplodrassus chotanagpurensis Gajbe, 1987 | | India urn:lsid:nmbe.ch:spidersp:027325

Haplodrassus chotanagpurensis Gajbe, 1987a: 277, f. 1-5 (Df) Original description

Haplodrassus cognatus (Westring, 1861) | | Europe, Russia (Europe to Far East), Caucasus, Kazakhstan, Japan urn:lsid:nmbe.ch:spidersp:027326

Drassus fuscus C. L. Koch, 1843: 123, f. 837 (f, misidentified)

Drassus cognatus Westring, 1861: 343 (Df) Original description

Drassus fuscus L. Koch, 1866: 86, pl. 4, f. 58 (f)

Drassus capnodes Thorell, 1875a: 96 (Dm) [urn:lsid:nmbe.ch:spidersp:058451]

Drassus capnodes Thorell, 1875c: 92 (Dm)

Drassus cognatus Dahl, 1883: 56 (Dm)

Drassus buddebergi Simon, 1887c: 142 (Df) [urn:lsid:nmbe.ch:spidersp:058461]

Drassus cognatus Chyzer & Kulczyński, 1897: 215, pl. 8, f. 53 (mf)

Drassus capnodes Bösenberg, 1902: 296, pl. 28, f. 435 (f)

Drassus cognatus Bösenberg, 1902: 297, pl. 28, f. 436 (mf)

Drassodes cognatus Simon, 1914: 136, 208, f. 231-233 (mf)

Drassodes capnodes Simon, 1914: 208 (S of Drassus buddebergi)

Drassodes cognatus Ermolajev, 1928b: 98, f. 1 (f)

Haplodrassus cognatus Reimoser, 1937a: 16, f. 34-35 (mf)

Haplodrassus capnodes Reimoser, 1937a: 17, f. 38 (m)

Haplodrassus cognatus ermolajewi Lohmander, 1942: 86 (onyl provisional name for f of Ermolajev, 1928b: 98) [urn:lsid:nmbe.ch:spidersp:027327]

Haplodrassus cognatus Palmgren, 1943: 95, f. 99 (mf)

Haplodrassus (Phoeodrassus) cognatus Lohmander, 1944: 13 (new subgenus)

Haplodrassus cognatus Tullgren, 1946: 106, f. 31C, pl. 17, f. 221-224 (mf)

Haplodrassus cognatus Brændegaard, 1966: 98, f. 85 (f)

Haplodrassus cognatus Miller, 1971: 82, pl. VII, f. 17, pl. XXVII, f. 19 (mf)

Haplodrassus cognatus Miller & Buchar, 1977: 168, pl. III, f. 8-10 (mf)

Haplodrassus cognatus Grimm, 1985: 134, f. 155, 168-169 (mf, S of Haplodrassus capnodes)

Haplodrassus cognatus Izmailova, 1989: 105, f. 88 (f)

Haplodrassus cognatus Heimer & Nentwig, 1991: 428, f. 1130 (mf)

Haplodrassus cognatus Kamura, 1995b: 126, f. 7-8 (f)

Haplodrassus cognatus Roberts, 1998: 109, f. (mf)

Haplodrassus cognatus Almquist, 2006: 407, f. 351a-f (mf)

Haplodrassus cognatus Kamura, 2007: 97, f. 7-8 (m)

Haplodrassus cognatus Kamura, 2009: 485, f. 32-34 (mf)

Haplodrassus cognatus Kovblyuk, Kastrygina & Omelko, 2012: 66, f. 15-16 (f)

Haplodrassus concertor (Simon, 1878) | | Spain, France, Italy urn:lsid:nmbe.ch:spidersp:027328

Drassus concertor Simon, 1878a: 129, pl. 16, f. 24 (Dmf) Original description

Drassodes concertor Simon, 1914: 140, 210, f. 251 (f; not f. 249, m = H. signifer per Bosmans et al., 2018: 50)

Haplodrassus concertor Denis, 1958c: 248, f. 1-2 (f)

Haplodrassus concertor Bosmans et al., 2018: 50, f. 205-213 (mf)

Haplodrassus crassipes (Lucas, 1846) | | Morocco, Algeria, Tunisia urn:lsid:nmbe.ch:spidersp:026875

Drassus crassipes Lucas, 1846: 217, pl. 13, f. 4 (Dm) Original description

Drassodes macellinus Simon, 1914: 139, 209, f. 248 (mf, misidentified per Bosmans et al., 2018: 8)

Haplodrassus crassipes Bosmans et al., 2018: 8, f. 1-9, 18-23 (mf, T from Drassodes)

Haplodrassus creticus (Roewer, 1928) | | Greece (Crete) urn:lsid:nmbe.ch:spidersp:026876

Drassodes creticus Roewer, 1928b: 104, pl. 1, f. 9 (Dm) Original description

Scotophaeus walteri Roewer, 1928b: 107, pl. 1, f. 18 (Df) [urn:lsid:nmbe.ch:spidersp:027908]

Haplodrassus creticus Chatzaki, Thaler & Mylonas, 2002a: 585, f. 55-60 (m, Df, T from Drassodes)

Haplodrassus creticus Levy, 2004: 28 (S of Scotophaeus walteri)

Haplodrassus creticus Bosmans et al., 2018: 53, f. 214-222 (mf)

Haplodrassus dalmatensis (L. Koch, 1866) | | Europe, North Africa, Turkey, Caucasus, Middle East, Russia (Europe) to Central Asia urn:lsid:nmbe.ch:spidersp:027329

Drassus dalmatensis L. Koch, 1866: 89, pl. 4, f. 59 (Df) Original description

Drassus minusculus L. Koch, 1866: 110, pl. 5, f. 70 (Df) [urn:lsid:nmbe.ch:spidersp:058465]

Drassus denotatus O. Pickard-Cambridge, 1874a: 398, pl. 52, f. 24 (Df) [urn:lsid:nmbe.ch:spidersp:026880]

Drassus pictus Thorell, 1875a: 97 (Dmf) [urn:lsid:nmbe.ch:spidersp:027330]

Drassus pictus Thorell, 1875c: 90 (Dmf)

Drassus delinquens O. Pickard-Cambridge, 1875e: 245, pl. 8, f. 4 (Df)

Drassus minusculus L. Koch, 1876b: 302 (Dm)

Drassus minusculus F. O. Pickard-Cambridge, 1895b: 38 (S of Drassus delinquens)

Drassus minusculus Chyzer & Kulczyński, 1897: 217, pl. 8, f. 58, 63 (mf)

Drassodes minusculus pictus Kulczyński, 1899: 349, pl. 6, f. 29 (mf)

Drassodes carinatus Strand, 1906b: 609 (Dmf) [urn:lsid:nmbe.ch:spidersp:026869]

Drassodes dalmatensis Simon, 1909b: 15 (S of Drassus minusculus and D. pictus)

Drassodes dalmatensis Simon, 1914: 137, 209, f. 237-239 (mf)

Drassodes palaestinensis Strand, 1915c: 137 (Dm) [urn:lsid:nmbe.ch:spidersp:026964]

Drassodes lithobius Roewer, 1928b: 101, pl. 1, f. 7 (Df) [urn:lsid:nmbe.ch:spidersp:026932]

Scotophaeus desertorum Caporiacco, 1928b: 86, f. 3 (Df) [urn:lsid:nmbe.ch:spidersp:027770]

Pseudodrassus desertorum Caporiacco, 1935a: 286

Drassodes dalmatensis Miller, 1936b: 262, f. 1-2 (mf)

Haplodrassus dalmatensis Tullgren, 1946: 100, pl. 16, f. 201-203 (mf)

Haplodrassus dalmatensis Miller, 1947: 55, pl. II, f. 3 (f)

Drassodes dalmatensis Locket & Millidge, 1951: 103, f. 53C, F, H, 54B (mf)

Haplodrassus insularis Denis, 1962a: 35, f. 22 (Df) [urn:lsid:nmbe.ch:spidersp:058454]

Haplodrassus dalmatensis Brændegaard, 1966: 99, f. 86 (f)

Pseudodrassus desertorum Denis, 1966e: 106, f. 5 (f)

Haplodrassus dalmatensis Tystshenko, 1971: 97, f. 176 (m)

Haplodrassus dalmatensis Miller, 1971: 83, pl. VII, f. 5-7 (mf)

Haplodrassus dalmatensis Merrett, 1972: 180, f. 1d (m)

Haplodrassus dalmatensis Locket, Millidge & Merrett, 1974: 6, f. 3D (m)

Haplodrassus dalmatensis Schmidt, 1977: 58 (S of Haplodrassus insularis)

Haplodrassus dalmatensis Miller & Buchar, 1977: 170, pl. IV, f. 1-3 (mf)

Haplodrassus dalmatensis Grimm, 1985: 138, f. 156, 164-165 (mf)

Haplodrassus dalmatensis Roberts, 1985: 66, f. 24a (mf)

Haplodrassus dalmatensis Heimer & Nentwig, 1991: 428, f. 1129 (mf)

Haplodrassus dalmatensis Noordam, 1992: 6 (mf)

Haplodrassus dalmatensis Roberts, 1995: 106, f. (mf)

Haplodrassus dalmatensis Roberts, 1998: 110, f. (mf)

Haplodrassus dalmatensis Chatzaki, Thaler & Mylonas, 2002a: 587, f. 62 (m, S of Drassodes lithobius)

Haplodrassus dalmatensis Levy, 2004: 23, f. 60 (f, S of Drassodes palaestinensis, Drassus denotatus and Pseudodrassus desertorum; mf 57-59, 61 misidentifed, see H. omissus)

Haplodrassus dalmatensis Almquist, 2006: 408, f. 352a-e (mf)

Haplodrassus dalmatensis Chatzopoulou & Chatzaki, 2009: 358 (S of Drassodes carinatus)

Haplodrassus dalmatensis Kovblyuk, Kastrygina & Omelko, 2012: 66, f. 22-24, 28-29 (mf)

Haplodrassus dalmatensis Lecigne, 2016a: 111, f. 7C-H (mf)

Haplodrassus dalmatensis Bosmans et al., 2018: 12, f. 10-17, 24-29 (mf)

Haplodrassus dalmatensis Zamani, Al-Yacoub & Najim, 2022: 144, f. 4A (f)

Haplodrassus dalmatensis Rehfeldt & Cassar, 2024: 23, f. 9a-c (f)

Haplodrassus dentatus Xu & Song, 1987 | | China urn:lsid:nmbe.ch:spidersp:027331

Haplodrassus dentatus Xu & Song, 1987: 83, f. 1-4 (Dmf) Original description

Haplodrassus dentatus Xu, 1991b: 37, f. 15 (f)

Haplodrassus dentatus Song, Zhu & Chen, 1999: 451, f. 263B, J (mf)

Haplodrassus dentatus Song, Zhu & Zhang, 2004: 132, f. 77A-D (mf)

Haplodrassus dentatus Omelko & Marusik, 2012: 344, f. 13-15, 19, 25 (mf)

Haplodrassus dentifer Bosmans & Abrous, 2018 | | Morocco, Algeria, Tunisia, Spain urn:lsid:nmbe.ch:spidersp:050459

Haplodrassus dentifer Bosmans & Abrous in Bosmans et al., 2018: 15, f. 30-40, 49-53 (Dmf) Original description

Haplodrassus deserticola Schmidt & Krause, 1996 | | Canary Is. urn:lsid:nmbe.ch:spidersp:027332

Haplodrassus deserticola Schmidt & Krause, 1996: 266, f. 4-5 (Df) Original description

Haplodrassus deserticola Wunderlich, 2011: 44, f. 84-88 (f, Dm)

Haplodrassus deserticola Kovblyuk, Kastrygina & Omelko, 2012: 74, f. 40-42, 50-51, 66-67 (mf)

Haplodrassus dixiensis Chamberlin & Woodbury, 1929 | | USA urn:lsid:nmbe.ch:spidersp:027333

Haplodrassus dixiensis Chamberlin & Woodbury, 1929: 134, pl. 2, f. 6 (Df) [text of pl. 1 belongs to pl. 2] Original description

Haplodrassus dixiensis Platnick & Shadab, 1975b: 23, f. 39-46 (f, Dm)

Haplodrassus dumdumensis Tikader, 1982 | | India urn:lsid:nmbe.ch:spidersp:027334

Haplodrassus dumdumensis Tikader, 1982b: 468, f. 391-393 (Df; probably misplaced) Original description

Haplodrassus eunis Chamberlin, 1922 | | Canada, USA urn:lsid:nmbe.ch:spidersp:027335

Haplodrassus eunis Chamberlin, 1922: 162 (Df) Original description

Haplodrassus utus Chamberlin & Ivie, 1946: 8, f. 9-11 (Dmf) [urn:lsid:nmbe.ch:spidersp:058457]

Haplodrassus eunis Platnick & Shadab, 1975b: 20, f. 31-38 (mf, S of Haplodrassus utus)

Haplodrassus eunis Platnick & Dondale, 1992: 222, f. 334-337 (mf)

Haplodrassus eunis Paquin & Dupérré, 2003: 78, f. 679-681 (mf)

Haplodrassus guiyangensis Yan & Yu, 2021 | | China urn:lsid:nmbe.ch:spidersp:053645

Haplodrassus guiyangensis Yan & Yu in Yan, Qin & Yu, 2021: 124, f. 1A-D, 2A-F (Dm) Original description

Haplodrassus hatsushibai Kamura, 2007 | | Korea, Japan urn:lsid:nmbe.ch:spidersp:040609

Haplodrassus montanus Chikuni, 1989b: 121, f. 17 (mf, misidentified)

Haplodrassus hatsushibai Kamura, 2007: 100, f. 23-31 (Dmf) Original description

Haplodrassus hatsushibai Kamura, 2009: 487, f. 43-45 (mf)

Haplodrassus hatsushibai Omelko & Marusik, 2012: 344, f. 5-6, 10-12, 18, 23-24 (mf)

Haplodrassus hatsushibai Seo, 2017d: 118, f. 4A-D,I-J (f)

Haplodrassus hatsushibai Suzuki, 2019e: 108, f. 6-9 (mf)

Haplodrassus huarong Yin & Bao, 2012 | | China urn:lsid:nmbe.ch:spidersp:046214

Haplodrassus huarong Yin & Bao in Yin et al., 2012: 1174, f. 625a-h (Dmf) Original description

Haplodrassus hunanensis Yin & Bao, 2012 | | China urn:lsid:nmbe.ch:spidersp:046215

Haplodrassus hunanensis Yin & Bao in Yin et al., 2012: 1175, f. 626a-e (Df) Original description

Haplodrassus ibericus Melic, Silva & Barrientos, 2016 | | Portugal, Spain urn:lsid:nmbe.ch:spidersp:048590

Haplodrassus ibericus Melic, Silva & Barrientos in Melic et al., 2016: 108, f. 1a-g (Dmf) Original description

Haplodrassus invalidus (O. Pickard-Cambridge, 1872) | | Greece, Cyprus, Turkey, Israel, Egypt, Azerbaijan urn:lsid:nmbe.ch:spidersp:027339

Drassus invalidus O. Pickard-Cambridge, 1872: 237, pl. 15, f. 14 (Dm) Original description

Drassus invalidus Simon, 1878a: 135 (Df)

Haplodrassus invalidus Roewer, 1955c: 399

Haplodrassus invalidus Levy, 2004: 31, f. 70-73 (mf, S of H. vignai, rejected by Bosmans et al., 2018: 54, see H. macellinus)

Haplodrassus invalidus Kovblyuk, Kastrygina & Omelko, 2012: 68, f. 32-34 (m)

Haplodrassus invalidus Bosmans et al., 2018: 54, f. 223-231 (mf)

Haplodrassus invalidus Chatzaki & Van Keer, 2019: 443, f. 21-22 (f)

Haplodrassus ivlievi Ponomarev, 2015 | | Russia (Europe) urn:lsid:nmbe.ch:spidersp:048470

Haplodrassus ivlievi Ponomarev, 2015: 260, f. 5-6 (Df) Original description

Haplodrassus jacobi Gajbe, 1992 | | India urn:lsid:nmbe.ch:spidersp:027340

Haplodrassus jacobi Gajbe, 1992b: 315, f. 9-13 (Df) Original description

Haplodrassus kanenoi Kamura, 1995 | | Japan urn:lsid:nmbe.ch:spidersp:027341

Haplodrassus kanenoi Kamura, 1995b: 123, f. 1-6 (Dmf) Original description

Haplodrassus kanenoi Kamura, 2007: 97, f. 19-20 (m)

Haplodrassus kanenoi Kamura, 2009: 485, f. 39-42 (mf)

Haplodrassus kulczynskii Lohmander, 1942 | | Europe, Turkey, Russia (Europe to Far East), China, Korea urn:lsid:nmbe.ch:spidersp:027342

Drassus microps Chyzer & Kulczyński, 1897: 217, pl. 8, f. 57 (mf; misidentified per Lohmander, 1942: 101)

Drassus microps Bösenberg, 1902: 300, pl. 28, f. 64, 442 (mf; misidentified per Lohmander, 1942: 101)

Drassodes microps Lessert, 1910b: 54 (misidentified per Lohmander, 1942: 101)

Drassodes microps Simon, 1914: 138, 209, f. 240-241 (mf; misidentified per Lohmander, 1942: 101)

Drassodes microps Reimoser, 1937a: 18, f. 41-42 (mf; misidentified per Lohmander, 1942: 101)

Haplodrassus kulczynskii Lohmander, 1942: 101 (Dmf) Original description

Haplodrassus microps Denis, 1951a: 104, f. 1-2 (m, misidentified)

Haplodrassus microps Buchar & Zdárek, 1960: 89, f. 1A-B (m, misidentified)

Haplodrassus microps Roșca, 1968: 86, f. 19a-b (mf, misidentified)

Haplodrassus microps Tystshenko, 1971: 97, f. 177 (m, misidentified)

Haplodrassus kulczynskii Miller, 1971: 82, pl. VII, f. 14-16 (mf)

Haplodrassus kulczynskii Miller & Buchar, 1977: 170, pl. IV, f. 7-10 (mf)

Haplodrassus magnipalpus Paik & Sohn, 1984: 106, f. 1-13 (Dmf) [urn:lsid:nmbe.ch:spidersp:058455]

Haplodrassus kulczynskii Grimm, 1985: 141, f. 152, 162-163 (mf)

Haplodrassus kulczynskii Paik & Kang, 1988: 63 (S of Haplodrassus magnipalpus)

Haplodrassus kulczynskii Heimer & Nentwig, 1991: 426, f. 1124 (mf)

Haplodrassus kulczynskii Esyunin & Efimik, 1995: 83, f. 22 (f)

Haplodrassus kulczynskii Roberts, 1998: 111, f. (mf)

Haplodrassus kulczynskii Namkung, 2002: 471, f. 37.12a-c (mf)

Haplodrassus kulczynskii Namkung, 2003: 474, f. 37.12a-c (mf)

Haplodrassus kulczynskii Song, Zhu & Zhang, 2004: 133, f. 78A-F (f)

Haplodrassus kulczynskii Jung et al., 2005: 181, f. 64, 89, 127 (mf)

Haplodrassus kulczynskii Marusik et al., 2007: 43, f. 5-10 (mf)

Haplodrassus kulczynskii Kovblyuk, Kastrygina & Omelko, 2012: 73, f. 35-39 (mf)

Haplodrassus kulczynskii Omelko & Marusik, 2012: 340, f. 28 (m)

Haplodrassus kulczynskii Kim & Lee, 2013: 111, f. 76A-D (mf)

Haplodrassus lilliputanus Levy, 2004 | | Israel urn:lsid:nmbe.ch:spidersp:040610

Haplodrassus lilliputanus Levy, 2004: 34, f. 76-77 (Dm) Original description

Haplodrassus longivulva Bosmans & Hervé, 2018 | | Morocco, Algeria urn:lsid:nmbe.ch:spidersp:050460

Haplodrassus longivulva Bosmans & Hervé in Bosmans et al., 2018: 17, f. 41-48, 54-57 (Dmf) Original description

Haplodrassus lyndae Abrous & Bosmans, 2018 | | Morocco, Algeria, Spain urn:lsid:nmbe.ch:spidersp:050461

Haplodrassus lyndae Abrous & Bosmans in Bosmans et al., 2018: 17, f. 58-66, 77-82 (Dmf) Original description

Haplodrassus lyndae Breitling, 2020: 343, f. 11D-E (m)

Haplodrassus macellinus (Thorell, 1871) | | France, Italy, Portugal?, Spain? urn:lsid:nmbe.ch:spidersp:027343

Drassus macellinus Thorell, 1871a: 185 (Dmf) Original description

Drassus nyctelius Thorell, 1871a: 185 (Df) [urn:lsid:nmbe.ch:spidersp:058458]

Drassus occidentalis Thorell, 1871a: 184 (Df) [urn:lsid:nmbe.ch:spidersp:058459]

Drassus hebes O. Pickard-Cambridge, 1874a: 390, pl. 51, f. 15 (Dm) [urn:lsid:nmbe.ch:spidersp:027344]

Drassus vicarius Simon, 1878a: 132 (Df) [urn:lsid:nmbe.ch:spidersp:058460]

Drassus macellinus Simon, 1878a: 138 (S of Drassus nyctelius and D. hebes, the latter elevated to subspecies by Simon, 1914: 140, 209)

Drassodes macellinus hebes Simon, 1914: 140, 209, f. 248 (mf, S of Drassus vicarius)

Drassodes macellinus Simon, 1914: 209 (S of Drassus occidentalis and D. typhon, the latter revalidated by Bosmans et al., 2018: 49)

Haplodrassus vignai Di Franco, 1996: 173, f. 1-4 (Dmf) [urn:lsid:nmbe.ch:spidersp:027369]

Haplodrassus macellinus Bosmans et al., 2018: 54, f. 232-240 (mf, S of H. macellinus hebes and H. vignai)

Haplodrassus macellinus Lecigne, 2018b: 67, f. 7B-E (m)

Haplodrassus maculatus (Banks, 1904) | | USA, Mexico urn:lsid:nmbe.ch:spidersp:027345

Zelotes maculata Banks, 1904b: 336, pl. 40, f. 38 (Dm) Original description

Haplodrassus maculatus Chamberlin, 1922: 161

Haplodrassus maculatus Platnick & Shadab, 1975b: 26, f. 55-62 (m, Df)

Haplodrassus mayumiae Kamura, 2007 | | Korea, Japan urn:lsid:nmbe.ch:spidersp:040611

Haplodrassus mayumiae Kamura, 2007: 98, f. 9-12, 17-18 (Dmf) Original description

Haplodrassus mayumiae Kamura, 2009: 485, f. 35-38 (mf)

Haplodrassus mayumiae Lee et al., 2015b: 52, f. 1A-F (m)

Haplodrassus medes Zamani, Chatzaki, Esyunin & Marusik, 2021 | | Iran urn:lsid:nmbe.ch:spidersp:053230

Haplodrassus medes Zamani et al., 2021a: 86, f. 11A-J (Dm) Original description

Haplodrassus mediterraneus Levy, 2004 | | Turkey, Syria, Lebanon, Israel, Jordan urn:lsid:nmbe.ch:spidersp:040612

Drassodes invalidus Simon, 1914: 138, 210, f. 244 (f, misidentified)

Haplodrassus mediterraneus Levy, 2004: 28, f. 66-69 (Dmf) Original description

Haplodrassus mediterraneus Seyyar, 2011: 1246, f. 1-6 (mf)

Haplodrassus mediterraneus Bosmans et al., 2018: 59, f. 241-251 (mf)

Haplodrassus mimus Chamberlin, 1922 | | USA urn:lsid:nmbe.ch:spidersp:027347

Haplodrassus mimus Chamberlin, 1922: 161 (Df) Original description

Haplodrassus mimus Platnick & Shadab, 1975b: 24, f. 47-54 (f, Dm)

Haplodrassus minor (O. Pickard-Cambridge, 1879) | | Europe, Turkey, Russia (Europe to West Siberia) urn:lsid:nmbe.ch:spidersp:027348

Drassus minor O. Pickard-Cambridge, 1879e: 192, pl. 12, f. 1 (Df) Original description

Drassus minor Chyzer & Kulczyński, 1897: 219, pl. 8, f. 55, 65 (f, Dm)

Drassus minor Bösenberg, 1902: 300, pl. 28, f. 443 (mf)

Drassodes minor Simon, 1914: 138, f. 242 (f)

Haplodrassus minor Reimoser, 1937a: 18, f. 39-40 (mf)

Drassodes minor Locket & Millidge, 1951: 103, f. 54C (mf)

Drassodes minor Locket & Millidge, 1957: 481, f. 1A-B (m)

Drassodes minor Cooke, 1963: 156, f. 4-5 (m)

Haplodrassus minor Miller, 1971: 82, pl. VI, f. 24 (m)

Haplodrassus minor Locket, Millidge & Merrett, 1974: 7, f. 2C-D (m)

Haplodrassus minor Miller & Buchar, 1977: 170, pl. IV, f. 4-6 (m)

Haplodrassus minor Grimm, 1985: 144, f. 176-178 (mf)

Haplodrassus minor Roberts, 1985: 66, f. 24c (mf)

Haplodrassus minor Heimer & Nentwig, 1991: 426, f. 1123 (mf)

Haplodrassus minor Roberts, 1995: 107, f. (mf)

Haplodrassus minor Roberts, 1998: 111, f. (mf)

Haplodrassus minor Chatzaki, Thaler & Mylonas, 2002a: 589, f. 63 (f)

Haplodrassus minor Tuneva & Esyunin, 2003a: 232, f. 27-33 (mf)

Haplodrassus minor Deltshev, 2004: 72, f. 6-8 (m)

Haplodrassus minor Kovblyuk, Kastrygina & Omelko, 2012: 73, f. 43-49, 52-65 (mf)

Haplodrassus minor Omelko & Marusik, 2012: 340, f. 27 (m)

Haplodrassus minor Coşar, Danışman & Kartaler, 2018: 36, f. 3A-B, 4A-B, 5 (f)

Haplodrassus minor Bosmans et al., 2018: 61, f. 252-256 (m)

Haplodrassus miryangensis Seo, 2017 | | Korea urn:lsid:nmbe.ch:spidersp:052085

Haplodrassus miryangensis Seo, 2017d: 119, f. 5A-F (Df) Original description

Haplodrassus moderatus (Kulczyński, 1897) | | Europe, Russia (Europe to Far East), China urn:lsid:nmbe.ch:spidersp:027349

Drassus moderatus Kulczyński in Chyzer & Kulczyński, 1897: 216, pl. 8, f. 60 (Df) Original description

Drassodes simplex Kulczyński, 1926: 39, pl. 2, f. 4 (Df) [urn:lsid:nmbe.ch:spidersp:026996]

Haplodrassus moderatus Lohmander, 1942: 82, 85 (Dm)

Haplodrassus moderatus Palmgren, 1943: 96, f

Haplodrassus moderatus Tullgren, 1946: 104, f. 31A, pl. 17, f. 214-216 (mf)

Haplodrassus modestus Roewer, 1955c: 400 (lapsus)

Haplodrassus moderatus Miller, 1971: 82, pl. VIII, f. 2 (m)

Haplodrassus moderatus Wunderlich, 1975b: 42, f. 1-5 (mf)

Haplodrassus moderatus Izmailova, 1977: 71, f. 3 (f)

Haplodrassus moderatus Izmailova, 1978a: 10, f. b (f)

Haplodrassus moderatus Grimm, 1985: 145, f. 154, 174-175 (mf)

Haplodrassus moderatus Izmailova, 1989: 106, f. 89 (f)

Haplodrassus moderatus Heimer & Nentwig, 1991: 428, f. 1132 (mf)

Haplodrassus moderatus Marusik & Logunov, 1995b: 192, f. 76-80 (mf)

Haplodrassus moderatus Roberts, 1998: 109, f. (mf)

Haplodrassus moderatus Song, Zhu & Zhang, 2004: 134, f. 79A-C (f)

Haplodrassus moderatus Almquist, 2006: 409, f. 353a-g (mf)

Haplodrassus moderatus Marusik & Kovblyuk, 2011: 156, f. 19 (f)

Haplodrassus simplex Mikhailov, 2013b: 172 (no T stated)

Haplodrassus moderatus Grbić, Hänggi & Krnjajić, 2021: 26, f. 7A-E (mf)

Haplodrassus moderatus Mikhailov & Marusik, 2023: 226 (S of Drassodes simplex)

Haplodrassus montanus Paik & Sohn, 1984 | | Russia (Far East), Korea urn:lsid:nmbe.ch:spidersp:027350

Haplodrassus montanus Paik & Sohn, 1984: 107, f. 14-26 (Dmf) Original description

Haplodrassus montanus Namkung, 2002: 470, f. 37.11a-b (mf)

Haplodrassus montanus Namkung, 2003: 473, f. 37.11a-b (mf)

Haplodrassus montanus Jung et al., 2005: 182, f. 65, 128 (mf)

Haplodrassus montanus Omelko & Marusik, 2012: 345, f. 1-4, 7-9, 16-17, 20-22 (mf)

Haplodrassus montanus Kim & Lee, 2013: 112, f. 77A-D (mf)

Haplodrassus montanus Seo, 2017d: 119, f. 4E-H (f)

Haplodrassus morosus (O. Pickard-Cambridge, 1872) | | Greece, Turkey, Israel urn:lsid:nmbe.ch:spidersp:026954

Drassus morosus O. Pickard-Cambridge, 1872: 232, pl. 15, f. 9 (Dm) Original description

Haplodrassus morosus Levy, 2004: 26, f. 62-65 (m, Df, T from Drassodes, S of Haplodrassus omissus, rejected by Bosmans et al., 2018: 25)

Haplodrassus nabozhenkoi Ponomarev, 2023 | | Russia (Caucasus) urn:lsid:nmbe.ch:spidersp:059516

Haplodrassus nabozhenkoi Ponomarev in Ponomarev & Shmatko, 2023b: 214, f. 1-7, 12-14 (Dmf) Original description

Haplodrassus nigroscriptus (Simon, 1909) | | Morocco urn:lsid:nmbe.ch:spidersp:026957

Drassodes nigroscriptus Simon, 1909b: 16 (Dmf) Original description

Drassodes nigroscriptus deminutus Simon, 1909b: 17 (Dmf) [urn:lsid:nmbe.ch:spidersp:026958]

Haplodrassus nigroscriptus Bosmans et al., 2018: 22, f. 67-76, 83-86 (mf, T from Drassodes, S of Drassodes nigroscriptus deminutus)

Haplodrassus nojimai Kamura, 2007 | | Korea, Japan urn:lsid:nmbe.ch:spidersp:040613

Haplodrassus nojimai Kamura, 2007: 102, f. 32-33 (Df) Original description

Haplodrassus nojimai Kamura, 2009: 487, f. 46 (f)

Haplodrassus nojimai Kim & Yoo, 2021: 17, f. 1A-F (f)

Haplodrassus omissus (O. Pickard-Cambridge, 1872) | | Canary Is., Morocco, Mediterranean urn:lsid:nmbe.ch:spidersp:050462

Drassus omissus O. Pickard-Cambridge, 1872: 239, pl. 15 (Df)

Drassus parvulus L. Koch, 1882: 632, pl. 1, f. 10-11 (Dm; preoccupied by Lucas, 1846, see Setaphis parvula)

Drassodes acrotirius Roewer, 1928b: 102, pl. 1, f. 8 (Df) [urn:lsid:nmbe.ch:spidersp:026843]

Drassodes seditiosus Caporiacco, 1928b: 83, f. 1 (Df) [urn:lsid:nmbe.ch:spidersp:027358]

Haplodrassus omissus Caporiacco, 1934a: 131

Haplodrassus seditiosus Caporiacco, 1934b: 127, f. 3 (Dm) [urn:lsid:nmbe.ch:spidersp:027358]

Drassodes parvicorpus Roewer, 1951: 443 (replacement name for Drassus parvulus) [urn:lsid:nmbe.ch:spidersp:026966] Original description

Haplodrassus maroccanus Denis, 1956a: 196, f. 4-6 (Dmf) [urn:lsid:nmbe.ch:spidersp:027346]

Drassodes acrotirius Chatzaki, Thaler & Mylonas, 2002a: 587, f. 62A-B (f)

Haplodrassus dalmatensis Levy, 2004: 23, f. 57-59, 61 (mf; misidentified per Bosmans et al., 2018: 25)

Haplodrassus acrotirius Bosmans & Chatzaki, 2005: 82 (T from Drassodes, confirmed by Bosmans et al., 2013: 10)

Haplodrassus parvicorpus Bosmans & Van Keer, 2012a: 11, f. 27-28 (mf, T from Drassodes, S of H. maroccanus)

Haplodrassus omissus Bosmans et al., 2018: 25, f. 87-97, 108-112 (mf, removed from S of H. morosus, contra Levy, 2004: 26, S of Haplodrassus acrotirius, H. maroccanus, H. parvicorpus [= Drassus parvulus] and H. seditiosus)

Haplodrassus orientalis (L. Koch, 1866) | | Bulgaria, Greece, Turkey, Ukraine, Russia (Europe), Georgia, Kazakhstan urn:lsid:nmbe.ch:spidersp:026963

Drassus orientalis L. Koch, 1866: 106, pl. 5, f. 68 (Df) Original description

Haplodrassus isaevi Ponomarev & Tsvetkov, 2006: 9, f. 12-14 (Dmf) [urn:lsid:nmbe.ch:spidersp:041362]

Haplodrassus isaevi Piterkina & Ovtsharenko, 2007: 1426, f. 1.1-6 (mf)

Haplodrassus isaevi Kovblyuk, Kastrygina & Omelko, 2012: 70, f. 25-27, 30-31 (mf)

Haplodrassus orientalis Bosmans et al., 2018: 61, f. 262-270 (f, T from Drassodes, S of Haplodrassus isaevi)

Haplodrassus orientalis Esyunin & Tuneva, 2020: 115, f. 50 (m)

Haplodrassus orientalis Naumova, Blagoev & Deltshev, 2021: 235, f. 5A-H (mf)

Haplodrassus orientalis Seyyar, Türkeş & Demir, 2022a: 332, f. 2, 3A-E (m)

Haplodrassus orientalis Seropian et al., 2023: 241, f. 22-23 (f)

Haplodrassus ovatus Bosmans & Hervé, 2018 | | Algeria, Tunisia urn:lsid:nmbe.ch:spidersp:050463

Haplodrassus ovatus Bosmans & Hervé in Bosmans et al., 2018: 29, f. 98-107, 113-117 (Dmf) Original description

Haplodrassus ovtchinnikovi Ponomarev, 2008 | | Turkey, Russia (Europe), Iran, Kazakhstan urn:lsid:nmbe.ch:spidersp:041586

Haplodrassus ovtchinnikovi Ponomarev, 2008a: 57, f. 26-28 (Dmf) Original description

Haplodrassus ovtchinnikovi Kovblyuk et al., 2009: 177, f. 42-50 (mf)

Haplodrassus paramecus Zhang, Song & Zhu, 2001 | | China urn:lsid:nmbe.ch:spidersp:038063

Haplodrassus paramecus Zhang, Song & Zhu, 2001: 53, f. 2A-C (Df) Original description

Haplodrassus paramecus Song, Zhu & Zhang, 2004: 136, f. 80A-C (f)

Haplodrassus pargongsanensis Paik, 1992 | | Korea urn:lsid:nmbe.ch:spidersp:027352

Haplodrassus pargongsanensis Paik, 1992j: 88, f. 9-14 (Df; N.B.: erroneously spelled H. pargongensis in legend) Original description

Haplodrassus pargongsanensis Jung et al., 2005: 183, f. 23, 66, 90 (f)

Haplodrassus pargongsanensis Kim & Lee, 2013: 114, f. 78A-B (f)

Haplodrassus ponomarevi Kovblyuk & Seyyar, 2009 | | Greece, Turkey urn:lsid:nmbe.ch:spidersp:042878

Haplodrassus ponomarevi Kovblyuk & Seyyar in Kovblyuk et al., 2009: 177, f. 51-62 (Dmf) Original description

Haplodrassus ponomarevi Bosmans et al., 2018: 61, f. 257-261 (m)

Haplodrassus pseudosignifer Marusik, Hippa & Koponen, 1996 | | Ukraine, Russia (Europe to Central Asia), Iran urn:lsid:nmbe.ch:spidersp:027353

Haplodrassus pseudosignifer Marusik, Hippa & Koponen, 1996: 26, f. 63-65, 69 (Dmf) Original description

Haplodrassus pseudosignifer Kovblyuk, Kastrygina & Omelko, 2012: 78, f. 68-70, 75-76 (mf)

Haplodrassus pseudosignifer Šestáková et al., 2022: 17, f. 9g-i (m)

Haplodrassus pugnans (Simon, 1880) | | Egypt, Israel, Russia (Europe to Far East), Iran, China, Korea, Japan urn:lsid:nmbe.ch:spidersp:027354

Drassus pugnans Simon, 1880b: 118, pl. 3, f. 20-21 (Dmf) Original description

Drassodes pugnans Schenkel, 1963: 43, f. 20a-c (mf)

Drassodes pseudopugnans Schenkel, 1963: 45, f. 21a-c (Dm) [urn:lsid:nmbe.ch:spidersp:027351]

Drassodes pugnans Song & Hubert, 1983: 17, f. 50-54 (mf, S of Drassodes pseudopugnans)

Haplodrassus signifer Hayashi, 1984: 13, f. 12-26 (mf, misidentified)

Haplodrassus pugnans Yaginuma, 1986a: 193, f. 108.1 (mf, T from Drassodes)

Haplodrassus pugnans Song, 1987: 340, f. 295 (mf)

Drassodes pugnans Zhang, 1987: 175, f. 147.1-5 (mf)

Haplodrassus pugnans Ovtsharenko & Marusik, 1988: 213, f. 56-58 (mf)

Haplodrassus pugnans Chikuni, 1989b: 121, f. 16 (mf)

Haplodrassus pugnans Ovtsharenko & Marusik, 1996: 120, pl. 2, f. 5-6 (mf)

Haplodrassus pugnans Song, Zhu & Chen, 1999: 451, f. 263C, K (mf)

Haplodrassus pugnans Hu, 2001: 255, f. 144.1-5 (mf)

Haplodrassus pugnans Song, Zhu & Chen, 2001: 345, f. 223A-D (mf)

Haplodrassus pugnans Levy, 2004: 20, f. 53-56 (mf, S of H. gridellii, rejected by Bosmans et al., 2018: 33, see H. rufipes)

Haplodrassus pugnans Song, Zhu & Zhang, 2004: 137, f. 81A-J (mf)

Haplodrassus pugnans Kamura, 2007: 95, f. 1-6 (mf)

Haplodrassus pugnans Kamura, 2009: 485, f. 29-31 (mf)

Haplodrassus pugnans Kovblyuk, Kastrygina & Omelko, 2012: 64, f. 7-9, 13-14 (mf)

Haplodrassus pugnans Kim, Lee & Yoo, 2018: 384, f. 1A-H (m)

Haplodrassus pugnans Zhang, Peng & Zhang, 2022: 237, f. 174A-G (mf)

Haplodrassus qashqai Zamani, Chatzaki, Esyunin & Marusik, 2021 | | Iran urn:lsid:nmbe.ch:spidersp:053231

Haplodrassus qashqai Zamani et al., 2021a: 87, f. 12A-H, 13A-D, 14A-H (Dmf) Original description

Haplodrassus reginae Schmidt & Krause, 1998 | | Cape Verde Is. urn:lsid:nmbe.ch:spidersp:027355

Haplodrassus reginae Schmidt & Krause, 1998: 421, f. 4 (Df) Original description

Haplodrassus rhodanicus (Simon, 1914) | | Portugal, Spain, France, Italy (Sardinia), Algeria, Tunisia, Iran urn:lsid:nmbe.ch:spidersp:026980

Drassodes rhodanicus Simon, 1914: 122, 134, 208 (Dm) Original description

Haplodrassus rhodanicus Bosmans et al., 2018: 29, f. 118-125, 136-139 (m, Df, T from Drassodes)

Haplodrassus rufipes (Lucas, 1846) | | Morocco, Algeria, Tunisia, Libyia, Portugal, Spain, France, Italy urn:lsid:nmbe.ch:spidersp:026985

Drassus severus C. L. Koch, 1843: 126, f. 838 (m, misidentified, only material from Italy, material from Greece is H. severus, per Bosmans et al., 2018: 33)

Drassus corticalis Lucas, 1846: 216, pl. 13, f. 3 (Df) [urn:lsid:nmbe.ch:spidersp:026874]

Drassus rufipes Lucas, 1846: 215, pl. 13, f. 2 (Df) Original description

Drassus similis L. Koch, 1866: 103, pl. 4, f. 67 (Dmf, removed from S of Haplodrassus severus per Bosmans et al., 2018: 33) [urn:lsid:nmbe.ch:spidersp:058462]

Drassodes severus Simon, 1893a: 361, f. 322 (m, misidentified per Bosmans et al., 2018: 36)

Drassodes severus Simon, 1914: 136, 210, f. 234-236 (mf, misidentified per Bosmans et al., 2018: 36)

Haplodrassus gridellii Caporiacco, 1949b: 117, f. 3 (Df) [urn:lsid:nmbe.ch:spidersp:027337]

Haplodrassus rufipes Bosmans et al., 2018: 33, f. 126-135, 140-144 (mf, T from Drassodes, S of Drassodes corticalis, D. similis (removed from S of H. severus) and of Haplodrassus gridellii (removed from S of H. pugnans, contra Levy, 2004 : 20))

Haplodrassus rufipes Breitling, 2020: 343, f. 11F-G (m)

Haplodrassus rufus (Savelyeva, 1972) | | Kazakhstan urn:lsid:nmbe.ch:spidersp:027356

Aphantaulax rufa Savelyeva, 1972a: 1238, f. 5-8 (Dmf) Original description

Aphantaulax rufus Brignoli, 1983c: 565

Haplodrassus rufus Ovtsharenko & Savelyeva, 1992: 132 (Tmf from Aphantaulax)

Haplodrassus rugosus Tuneva, 2004 | | Kazakhstan urn:lsid:nmbe.ch:spidersp:040614

Haplodrassus rugosus Tuneva, 2004: 322, f. 6-9 (Dm) Original description

Haplodrassus rugosus Ponomarev, Bragina & Shmatko, 2017: 7, f. 4 (m)

Haplodrassus sataraensis Tikader & Gajbe, 1977 | | India urn:lsid:nmbe.ch:spidersp:027357

Haplodrassus sataraensis Tikader & Gajbe, 1977c: 70, f. 4A-E (Dmf) Original description

Haplodrassus sataraensis Tikader, 1982b: 466, f. 385-390 (mf)

Haplodrassus sataraensis Gajbe, 2007: 479, f. 167-172 (mf)

Haplodrassus securifer Bosmans & Abrous, 2018 | | Morocco, Algeria, Tunisia, Portugal, Spain, France, Italy, Belgium urn:lsid:nmbe.ch:spidersp:050464

Drassodes invalidus Simon, 1914: 138, 210, f. 243 (m, misidentified per Bosmans et al., 2018: 39; not f, = H. mediterraneus)

Haplodrassus securifer Bosmans & Abrous in Bosmans et al., 2018: 39, f. 145-153, 162-166 (Dmf) Original description

Haplodrassus signifer (C. L. Koch, 1839) | | North America, Europe, North Africa, Turkey, Caucasus, Russia (Europe to Far East), Israel, Iran, Central Asia, China, Korea urn:lsid:nmbe.ch:spidersp:027360

Drassus signifer C. L. Koch, 1839a: vol. 7, p. 31, f. 452 (Df) Original description

Drassus troglodytes C. L. Koch, 1839a: vol. 7, p. 35, f. 455-456 (Dmf) [urn:lsid:nmbe.ch:spidersp:058467]

Clubiona troglodytes Walckenaer, 1841: 480

Drassus clavator O. Pickard-Cambridge, 1860: 171 (Dmf) [urn:lsid:nmbe.ch:spidersp:058468]

Drassus clavator Blackwall, 1861a: 109, pl. 6, f. 66 (m)

Drassus troglodytes L. Koch, 1866: 116, pl. 5, f. 73-74 (mf)

Drassus troglodytes O. Pickard-Cambridge, 1872: 232 (S of Drassus clavator)

Drassus troglodytes Menge, 1875: 378, pl. 65, f. 216 (mf)

Drassus robustus Emerton, 1890: 179, pl. 4, f. 8 (Dmf) [urn:lsid:nmbe.ch:spidersp:058475]

Drassus mysticus O. Pickard-Cambridge, 1895b: 104 (Dm) [urn:lsid:nmbe.ch:spidersp:058469]

Teminius nigriceps Banks, 1895b: 421 (Df) [urn:lsid:nmbe.ch:spidersp:058476]

Drassus troglodytes Becker, 1896: 257, pl. 16, f. 13 (mf)

Drassus placidus Banks, 1896e: 63 (Df) [urn:lsid:nmbe.ch:spidersp:058477]

Drassus troglodytes Chyzer & Kulczyński, 1897: 217, pl. 8, f. 56, 61 (mf)

Drassodes ferrum-equinum F. O. Pickard-Cambridge, 1899a: 60, pl. 4, f. 12 (Df) [urn:lsid:nmbe.ch:spidersp:058453]

Prosthesima decepta Banks, 1900c: 531 (Df) [urn:lsid:nmbe.ch:spidersp:058478]

Drassus troglodytes Bösenberg, 1902: 298, pl. 28, f. 439 (mf)

Drassodes robustus Bryant, 1908: 7

Drassodes troglodytes Lessert, 1910b: 52

Zelotes deceptus Petrunkevitch, 1911: 149

Syrisca nigriceps Petrunkevitch, 1911: 514

Drassodes signifer Comstock, 1912: 313, f. 303-304

Drassodes signifer Simon, 1914: 122, 140, 209, f. 249-250 (mf, S of Drassus troglodytes)

Haplodrassus concertor Simon, 1914: 140, f. 249 (m; misidentified per Bosmans et al., 2018: 50)

Drassodes beaufortensis Strand, 1916b: 95 (provisional name)

Haplodrassus signifer Chamberlin, 1922: 163 (S of Drassodes robustus, Drassus placidus, Syrisca nigriceps and Zelotes deceptus)

Drassus troglodytes Spassky, 1925: 35, f. 4 (f)

Haplodrassus altanus Chamberlin, 1933a: 5, f. 10 (f; misidentified per Platnick & Shadab, 1975b: 12)

Haplodrassus signifer Reimoser, 1937a: 17, f. 36-37 (mf)

Drassodes signifer Bristowe, 1939: 17 (S of Drassus mysticus)

Haplodrassus dystactus Chamberlin & Gertsch, 1940: 8, f. 6, 9-10 (Dmf) [urn:lsid:nmbe.ch:spidersp:058452]

Haplodrassus signifer Comstock, 1940: 326, f. 303 (f)

Haplodrassus signifer Palmgren, 1943: 95, f

Haplodrassus signifer Tullgren, 1946: 98, f. 30A, pl. 16, f. 197-200 (mf)

Drassodes signifer Brændegaard, 1946: 55, g. 35-36 (mf)

Haplodrassus signifer Kaston, 1948: 350, f. 1170-1172, 1186 (mf)

Drassodes signifer Locket & Millidge, 1951: 101, f. 53A, D, G, 54A (mf)

Haplodrassus signifer Brændegaard, 1966: 94, f. 81-82 (mf)

Haplodrassus signifer Tystshenko, 1971: 97, f. 175 (m)

Haplodrassus signifer Miller, 1971: 82, pl. VII, f. 18, pl. XXVII, f. 20-21 (mf)

Haplodrassus signifer Platnick & Shadab, 1975b: 11, f. 11-22 (mf, S of Drassodes ferrumequinum and Haplodrassus dystactus)

Haplodrassus signifer Miller & Buchar, 1977: 168, pl. II, f. 7-10 (mf)

Haplodrassus signifer Paik, 1978e: 417, f. 188.1-5 (f)

Drassodes signifer Jia & Zhu, 1983: 167, f. a-e (mf)

Haplodrassus signifer Thaler, 1984a: 189, f. 9d-f, i (m)

Haplodrassus signifer Grimm, 1985: 146, f. 146-148, 170-171 (mf)

Haplodrassus signifer Roberts, 1985: 66, f. 23g (mf)

Haplodrassus signifer Heiss & Allen, 1986: 57, f. 95-96 (mf)

Haplodrassus aeneus Hu & Li, 1987b: 301, f. 32.1-4 (mf, misidentified)

Haplodrassus signifer Izmailova, 1989: 107, f. 90 (m)

Haplodrassus signifer Heimer & Nentwig, 1991: 426, f. 1125 (mf)

Haplodrassus signifer Platnick & Dondale, 1992: 215, f. 322-325 (mf)

Haplodrassus signifer Noordam, 1992: 6 (mf)

Haplodrassus signifer Paik, 1992j: 89, text-f. 1A-B (f)

Haplodrassus signifer Roberts, 1995: 106, f. (mf)

Haplodrassus signifer Marusik, Hippa & Koponen, 1996: 26, f. 66-68, 70 (mf)

Haplodrassus signifer Ovtsharenko & Marusik, 1996: 121, pl. 2, f. 7-8 (mf)

Haplodrassus signifer Agnarsson, 1996: 31, f. 11A-C (mf)

Haplodrassus signifer Mcheidze, 1997: 114, f. 157-160 (mf)

Haplodrassus signifer Bellmann, 1997: 170, f. (f)

Haplodrassus signifer Roberts, 1998: 109, f. (mf)

Drassodes signifer Hu, 2001: 240, f. 132.1-2 (mf)

Haplodrassus aeneus Hu, 2001: 255, f. 143.1-4 (mf; misidentified per Song, Zhu & Zhang, 2004: 140)

Haplodrassus signifer Chatzaki, Thaler & Mylonas, 2002a: 587, f. 61 (m)

Haplodrassus signifer Paquin & Dupérré, 2003: 78, f. 682-684 (mf)

Haplodrassus signifer Levy, 2004: 19, f. 46-52 (mf)

Haplodrassus signifer Song, Zhu & Zhang, 2004: 139, f. 82A-I (mf)

Haplodrassus signifer Almquist, 2006: 411, f. 354a-i (mf)

Haplodrassus signifer Kovblyuk, Kastrygina & Omelko, 2012: 81, f. 71-74, 77-78 (mf)

Haplodrassus signifer Omelko & Marusik, 2012: 340, f. 26 (m)

Haplodrassus signifer Kim & Lee, 2013: 114, f. 79A-D (mf)

Haplodrassus signifer Marusik, 2015: 675, f. 2J, 9A, C, 10 (mf)

Haplodrassus signifer Bosmans et al., 2018: 42, f. 154-161, 167-171 (mf)

Haplodrassus signifer Breitling, 2020: 343, f. 11H-I (m)

Haplodrassus signifer Febrer & Barrientos, 2022: 138, f. 3A-B (f)

Haplodrassus signifer Šestáková et al., 2022: 17, f. 9d-f (m)

Haplodrassus signifer Ponomarev & Shmatko, 2023b: 214, f. 8-11, 15-17 (mf)

Haplodrassus silvestris (Blackwall, 1833) | | Europe, Turkey, Caucasus urn:lsid:nmbe.ch:spidersp:027361

Drassus sylvestris Blackwall, 1833: 440 (Df; emendation now amply protected by usage) Original description

Filistata dubia Wider, 1834: 202, pl. 14, f. 6a-b (Df) [urn:lsid:nmbe.ch:spidersp:051355]

Drassus infuscatus Westring, 1861: 347 (Df) [urn:lsid:nmbe.ch:spidersp:058463]

Drassus sylvestris Blackwall, 1861a: 113, pl. 6, f. 68 (f, Dm)

Drassus infuscatus L. Koch, 1866: 99, pl. 4, f. 64 (f, Dm)

Drassus infuscatus Menge, 1875: 381, pl. 66, f. 218 (mf)

Drassus criminalis O. Pickard-Cambridge, 1875e: 244, pl. 8, f. 3 (Df) [urn:lsid:nmbe.ch:spidersp:058464]

Drassodes silvestris Simon, 1893a: 361 (S of Drassus infuscatus)

Drassus infuscatus Becker, 1896: 256, pl. 16, f. 12 (mf)

Drassus silvestris Chyzer & Kulczyński, 1897: 216, pl. 8, f. 45, 54 (mf)

Drassus silvestris Smith, 1900a: 330 (S of Drassus criminalis)

Drassus infuscatus Bösenberg, 1902: 299, pl. 28, f. 440 (mf)

Drassodes silvestris Simon, 1914: 135, 208, f. 228-230 (mf)

Haplodrassus silvestris Reimoser, 1937a: 15, f. 32-33 (mf)

Haplodrassus silvestris Tullgren, 1946: 105, f. 31B, pl. 17, f. 217-220 (mf)

Drassodes silvestris Locket & Millidge, 1951: 103, f. 53B, E, 54D (mf)

Filistata dubia Roewer, 1955c: 1510 (nomen dubium)

Haplodrassus silvestris Brændegaard, 1966: 96, f. 83-84 (mf)

Haplodrassus silvestris Miller, 1971: 82, pl. VII, f. 2-4 (mf)

Haplodrassus silvestris Miller & Buchar, 1977: 168, pl. III, f. 5-7 (mf)

Haplodrassus silvestris Grimm, 1985: 150, f. 18b, 153, 166-167 (mf)

Haplodrassus silvestris Roberts, 1985: 66, f. 24b (mf)

Haplodrassus silvestris Izmailova, 1989: 107, f. 91 (f)

Haplodrassus silvestris Heimer & Nentwig, 1991: 428, f. 1131 (mf)

Haplodrassus silvestris Noordam, 1992: 6 (mf)

Haplodrassus silvestris Roberts, 1995: 107, f. (mf)

Haplodrassus silvestris Roberts, 1998: 110, f. (mf)

Haplodrassus silvestris Almquist, 2006: 411, f. 355a-f (mf)

Haplodrassus silvestris Seyyar, 2011: 1248, f. 7-9 (m)

Haplodrassus silvestris Kovblyuk, Kastrygina & Omelko, 2012: 66, f. 17-21 (mf)

Haplodrassus silvestris Ponomarev & Dvadnenko, 2013: 52, f. 1c-d, 2c-d, 3d (mf)

Haplodrassus silvestris Zonstein & Marusik, 2019b: 83 (S of Filistata dubia, removed from nomen dubium)

Haplodrassus silvestris Seropian et al., 2023: 242, f. 24-25 (f)

Haplodrassus soerenseni (Strand, 1900) | | Europe, Russia (Europe to Far East), Kazakhstan, China, Turkey? urn:lsid:nmbe.ch:spidersp:027362

Drassus soerenseni Strand, 1900a: 98, f. (Df) Original description

Haplodrassus lapponicus Holm, 1939b: 4, f. 1a-d, 2 (Dmf)

Haplodrassus soerenseni Lohmander, 1942: 101

Haplodrassus soerenseni Tullgren, 1942: 221, 230, pl. 1, f. 3 (mf)

Haplodrassus soerenseni Palmgren, 1943: 97, f

Haplodrassus soerenseni Tullgren, 1946: 102, f. 30C, pl. 17, f. 209-213 (mf)

Haplodrassus sörenseni Locket & Millidge, 1953: 406, f. 243A-C (mf)

Haplodrassus soerenseni Miller, 1971: 84, pl. VII, f. 8-10 (mf)

Haplodrassus sörenseni Merrett, 1972: 180, f. 1e (m)

Haplodrassus soerenseni Locket, Millidge & Merrett, 1974: 6, f. 3E (m)

Haplodrassus soerenseni Miller & Buchar, 1977: 168, pl. II, f. 11-12, pl. III, f. 1 (mf)

Haplodrassus soerenseni Grimm, 1985: 153, f. 151, 160-161 (mf)

Haplodrassus soerenseni Roberts, 1985: 66, f. 24d (mf)

Haplodrassus soerenseni Ovtsharenko & Marusik, 1988: 214, f. 48, 51-52 (mf)

Haplodrassus soerenseni Heimer & Nentwig, 1991: 428, f. 1127 (mf)

Haplodrassus soerenseni Roberts, 1995: 107, f. (mf)

Haplodrassus soerenseni Ovtsharenko & Marusik, 1996: 121, pl. 2, f. 9-10 (mf)

Haplodrassus soerenseni Roberts, 1998: 111, f. (mf)

Haplodrassus soerenseni Song, Zhu & Zhang, 2004: 141, f. 83A-C (f)

Haplodrassus soerenseni Almquist, 2006: 412, f. 356a-e (mf)

Haplodrassus soerenseni Ponomarev & Shmatko, 2017: 11, f. 7, 10 (m)

Haplodrassus stuxbergi (L. Koch, 1879) | | North America, Russia (Europe to Far East) urn:lsid:nmbe.ch:spidersp:027363

Drassus stuxbergi L. Koch, 1879c: 87, pl. 3, f. 6 (Df) Original description

Drassus hiemalis Emerton, 1909: 218, pl. 9, f. 1 (Dmf) [urn:lsid:nmbe.ch:spidersp:027338]

Drassodes stuxbergi Kulczyński, 1916: 2, pl. 1, f. 1 (f)

Haplodrassus hiemalis Chamberlin, 1922: 161

Haplodrassus altanus Chamberlin, 1933a: 5, f. 8-9 (Dm; f see Haplodrassus signifer) [urn:lsid:nmbe.ch:spidersp:058450]

Haplodrassus hiemalis Kaston, 1948: 350, f. 1173-1175, 1187 (mf)

Haplodrassus stuxbergi Holm, 1973: 103 (T from Drassodes)

Haplodrassus hiemalis Ubick & Roth, 1973a: 4 (S of Haplodrassus altanus)

Haplodrassus hiemalis Platnick & Shadab, 1975b: 7, f. 1-8 (mf)

Haplodrassus hiemalis Ovtsharenko & Marusik, 1988: 211, f. 54-55, 61 (mf)

Haplodrassus hiemalis Platnick & Dondale, 1992: 217, f. 326-329 (mf)

Haplodrassus hiemalis Marusik & Logunov, 1995b: 192, f. 71-75 (mf)

Haplodrassus hiemalis Ovtsharenko & Marusik, 1996: 120, pl. 2, f. 3-4 (mf)

Haplodrassus hiemalis Paquin & Dupérré, 2003: 78, f. 676-678 (mf)

Haplodrassus hiemalis Kamura, 2007: 98, f. 13-16, 21-22 (mf)

Haplodrassus hiemalis Murphy, 2007: 58, f. 480-481 (mf)

Haplodrassus hiemalis Omelko & Marusik, 2012: 340, f. 25 (m)

Haplodrassus hiemalis Azevedo, Griswold & Santos, 2018a: 681, f. 4C (m)

Haplodrassus hiemalis Azevedo, Griswold & Santos, 2018b: 609, f. 23b, S1b, S6d (m)

Haplodrassus stuxbergi Mikhailov & Marusik, 2023: 226 (S of Haplodrassus hiemalis)

Haplodrassus taepaikensis Paik, 1992 | | Russia (South Siberia, Far East), Korea urn:lsid:nmbe.ch:spidersp:027364

Haplodrassus taepaikensis Paik, 1992j: 86, f. 1-8 (Df) Original description

Haplodrassus taepaikensis Jung et al., 2005: 184, f. 25, 67, 91 (f)

Haplodrassus taepaikensis Marusik et al., 2007: 43, f. 1-4, 11-18 (f, Dm)

Haplodrassus taepaikensis Marusik & Kovblyuk, 2011: 156, f. 12, 16 (mf)

Haplodrassus taepaikensis Kim & Lee, 2013: 116, f. 80A-B (f)

Haplodrassus taibo (Chamberlin, 1919) | | USA urn:lsid:nmbe.ch:spidersp:027365

Zelotes taibo Chamberlin, 1919b: 6, pl. 2, f. 5 (Df) Original description

Haplodrassus taibo Chamberlin, 1922: 161

Haplodrassus taibo Platnick & Shadab, 1975b: 29, f. 71-74 (f, Dm)

Haplodrassus tegulatus (Schenkel, 1963) | | Russia (South Siberia), China urn:lsid:nmbe.ch:spidersp:028043

Drassodes tegulatus Schenkel, 1963: 40, f. 18 (Df) Original description

Tuvadrassus tegulatus Marusik & Logunov, 1995b: 194, f. 83-87 (Tf from Drassodes, Dm)

Tuvadrassus tegulatus Song, Zhu & Zhang, 2004: 238, f. 139A-D (mf)

Haplodrassus tegulatus Murphy, 2007: 58, f. 486-487 (f)

Haplodrassus tegulatus Omelko & Marusik, 2012: 341, f. 29 (m)

Haplodrassus tehriensis Tikader & Gajbe, 1977 | | India urn:lsid:nmbe.ch:spidersp:027366

Haplodrassus tehriensis Tikader & Gajbe, 1977c: 68, f. 3A-E (Dmf) Original description

Haplodrassus tehriensis Tikader, 1982b: 464, f. 379-384 (mf)

Haplodrassus triangularis Bosmans, 2018 | | Morocco, Tunisia urn:lsid:nmbe.ch:spidersp:050465

Haplodrassus triangularis Bosmans in Bosmans et al., 2018: 44, f. 172-180, 190-193 (Dmf) Original description

Haplodrassus typhon (Simon, 1878) | | Algeria, Tunisia, Portugal, Spain, France, Italy (Sardinia) urn:lsid:nmbe.ch:spidersp:050466

Drassus typhon Simon, 1878a: 128, pl. 16, f. 20 (Dmf) Original description

Haplodrassus typhon Bosmans et al., 2018: 49, f. 181-189, 194-197 (mf, T from Drassus/Drassodes, removed from S of H. macellinus, contra Simon, 191 4: 209)

Haplodrassus umbratilis (L. Koch, 1866) | | Europe to Kazakhstan urn:lsid:nmbe.ch:spidersp:027367

Drassus umbratilis L. Koch, 1866: 113, pl. 5, f. 71 (Df) Original description

Haplodrassus umbratilis Merrett, 1972: 179, f. 1a-c (mf)

Drassus microps Menge, 1875: 380, pl. 65, f. 217 (Dmf)

Haplodrassus umbratilis Locket, Millidge & Merrett, 1974: 7, f. 3A-C (mf)

Drassus umbratilis Chyzer & Kulczyński, 1897: 217, pl. 8, f. 59, 62 (mf)

Haplodrassus umbratilis Miller & Buchar, 1977: 168, pl. III, f. 2-4 (mf)

Drassus umbratilis Bösenberg, 1902: 299, pl. 28, f. 441 (mf)

Haplodrassus umbratilis Grimm, 1985: 156, f. 150, 158-159 (mf)

Drassodes umbratilis Simon, 1914: 138, 209, f. 240-247 (mf)

Haplodrassus umbratilis Roberts, 1985: 66, f. 25a (mf)

Drassus umbratilis Spassky, 1925: 35, f. 24 (m)

Haplodrassus umbratilis Heimer & Nentwig, 1991: 428, f. 1128 (mf)

Haplodrassus umbratilis Reimoser, 1937a: 19, f. 43-44 (mf)

Haplodrassus umbratilis Roberts, 1995: 108, f. (mf)

Haplodrassus umbratilis Holm, 1939b: 5, f. 1e (m)

Haplodrassus umbratilis Roberts, 1998: 112, f. (mf)

Haplodrassus umbratilis Palmgren, 1943: 96, f

Haplodrassus umbratilis Almquist, 2006: 413, f. 357a-f (mf)

Haplodrassus umbratilis Tullgren, 1946: 101, f. 30B, pl. 16, f. 204-208 (mf)

Haplodrassus umbratilis Kovblyuk, Kastrygina & Omelko, 2012: 84, f. 79-83 (mf)

Haplodrassus umbratilis Brændegaard, 1966: 100, f. 87 (f)

Haplodrassus umbratilis Ponomarev & Shmatko, 2017: 11, f. 2, 5-6, 9, 13-16 (mf)

Haplodrassus umbratilis Roșca, 1968: 86, f. 20a-b (mf)

Haplodrassus umbratilis Tystshenko, 1971: 97, f. 178 (m)

Haplodrassus umbratilis Miller, 1971: 84, pl. VII, f. 11-13 (mf)

Haplodrassus umbratilis gothicus Lohmander, 1942 | | Sweden urn:lsid:nmbe.ch:spidersp:027368

Haplodrassus umbratilis gothicus Lohmander, 1942: 101 (Df) Original description

Haplodrassus vastus (Hu, 1989) | | China urn:lsid:nmbe.ch:spidersp:028434

Zelotes vastus Hu, 1989: 99, f. 3-4 (Df) Original description

Zelotes vastus Hu & Wu, 1989: 295, f. 238.1-2 (f)

Haplodrassus vastus Song, Zhu & Zhang, 2004: 143, f. 84A-B (Tf from Zelotes)

Haplodrassus yinae Liu, 2022 | | China urn:lsid:nmbe.ch:spidersp:054501

Haplodrassus montanus Yin et al., 2012: 1177, f. 627a-f (mf; misidentified per Liu et al., 2022d: 192)

Haplodrassus yinae Liu in Liu et al., 2022d: 192, f. 1A-F, 2A-H, 3A-D (Dmf) Original description