Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Berinda Roewer, 1928 urn:lsid:nmbe.ch:spidergen:02404

Transferred to other genera:

Berinda aculeata (Charitonov, 1946) -- see Anagraphis

Berinda bifurcata Zamani, Chatzaki, Esyunin & Marusik, 2021 -- see Zagrotes

In synonymy:

Berinda citipes (Simon, 1893, T from Drassodes) = Berinda infumata O. Pickard-Cambridge, 1872 (Levy, 1998c: 145, sub Zelotes)

Berinda grazianoi (Caporiacco, 1948) = Berinda ensiger O. Pickard-Cambridge, 1874 (Bosmans & Chatzaki, 2005: 78, confirmed by Panayiotou et al., 2010: 47)

Berinda idae Lissner, 2016 = Berinda infumata O. Pickard-Cambridge, 1872 (Zamani et al., 2021a: 70)

Berinda reimoseri (Bristowe, 1935, T from Drassodes) = Berinda ensiger O. Pickard-Cambridge, 1874 (Chatzaki, Thaler & Mylonas, 2002a: 573)

Berinda aegilia Chatzaki, 2002 | | Greece urn:lsid:nmbe.ch:spidersp:038043
Berinda aegilia Chatzaki in Chatzaki, Thaler & Mylonas, 2002b: 606, f. 9-12 (Dmf) Original description
Berinda aegilia Panayiotou et al., 2010: 47, f. 1-2, 5-6 (mf)
Berinda amabilis Roewer, 1928 | | Greece (Crete) urn:lsid:nmbe.ch:spidersp:026679
Berinda amabilis Roewer, 1928b: 106, pl. 1, f. 11-13 (Dm) Original description
Berinda amabilis Chatzaki, Thaler & Mylonas, 2002a: 570, f. 12-13, 16-17 (m, Df, S of Talanites aculeatus, but moved to S of Anagraphis pallens by Khasayeva & Huseynov, 2017: 126)
Berinda amabilis Murphy, 2007: 37, f. 178-179 (mf)
Berinda amabilis Panayiotou et al., 2010: 47, f. 3-4, 7-78 (mf)
Berinda amabilis Wunderlich, 2011: 36, f. 16 (m)
Berinda cooki Logunov, 2012 | | Turkey urn:lsid:nmbe.ch:spidersp:045680
Berinda cooki Logunov, 2012b: 375, f. 1-2 (Dm) Original description
Berinda cooki Lecigne, 2021a: 17, f. 18a-b (m)
Berinda cypria Chatzaki & Panayiotou, 2010 | | Cyprus urn:lsid:nmbe.ch:spidersp:042983
Berinda cypria Chatzaki & Panayiotou in Panayiotou et al., 2010: 50, f. 13-14, 17, 20-21 (Dmf) Original description
Berinda ensiger (O. Pickard-Cambridge, 1874) | | Greece (incl. Crete), Cyprus, Turkey urn:lsid:nmbe.ch:spidersp:026890
Drassus ensiger O. Pickard-Cambridge, 1874a: 389, pl. 51, f. 14 (Dmf; for the ending of the specific epithet see ICZN Article 31.2.2) Original description
Drassodes reimoseri Bristowe, 1935: 779, f. 4-6 (Df)
Drassodes reimoseri Hadjissarantos, 1940: 77, f. 22 (Dm)
Haplodrassus grazianoi Caporiacco, 1948c: 59, f. 10 (Df) [urn:lsid:nmbe.ch:spidersp:027336]
Berinda ensigera Chatzaki, Thaler & Mylonas, 2002a: 573, f. 14-15, 18-19 (mf, S of Drassodes reimoseri)
Berinda ensigera Bosmans & Chatzaki, 2005: 78 (S of Haplodrassus grazianoi)
Berinda ensiger Kovblyuk et al., 2009: 170, f. 1-11 (mf)
Berinda ensigera Panayiotou et al., 2010: 47, f. 9-12 (mf, S of Haplodrassus grazianoi confirmed)
Berinda ensigera Lecigne, 2011: 18, f. pp. 19-20 (m)
Berinda ensigera Lecigne, 2016a: 109, f. 5D-J (mf)
Berinda ensigera Lecigne, 2021a: 31, f. 29 (m)
Berinda ensigera Tutar & Yağmur, 2023: 205, f. 9A-B (f)
Berinda hakani Chatzaki & Seyyar, 2010 | | Turkey urn:lsid:nmbe.ch:spidersp:042984
Berinda hakani Chatzaki & Seyyar in Panayiotou et al., 2010: 52, f. 15-16, 18-19, 22-23 (Dmf) Original description
Berinda hakani Tutar & Yağmur, 2023: 205, f. 10A-B (f)
Berinda hoerwegi Zamani, Chatzaki, Esyunin & Marusik, 2021 | | Iran urn:lsid:nmbe.ch:spidersp:053224
Berinda hoerwegi Zamani et al., 2021a: 72, f. 2A-L (Dmf) Original description
Berinda infumata (O. Pickard-Cambridge, 1872) | | Greece, Cyprus, Tanzania, Egypt, Israel. Introduced to Japan urn:lsid:nmbe.ch:spidersp:028252
Drassus infumatus O. Pickard-Cambridge, 1872: 238, pl. 15, f. 16 (Dmf) Original description
Drassodes citipes Simon, 1893a: 362 (Df)
Drassodes lutorius Tullgren, 1910: 104, pl. 1, f. 15b (f, misidentified, not f. 15a)
Camillina lutoria Berland, 1920c: 108, f. 125-127 (f, misidentified)
Zelotes infumatus Levy, 1998c: 145, f. 112-115 (mf, T from Drassodes, S of Drassodes citipes)
Heser infumatus Tuneva, 2004: 323 (T from Zelotes)
Zelotes infumatus FitzPatrick, 2007: 124, f. 97-100 (mf)
Berinda idae Lissner in Lissner & Chatzaki, 2016a: 9, f. 1a-c, 2a-b, 3a-b, 4a-b, 5a-b, 6-7 (Dmf) [urn:lsid:nmbe.ch:spidersp:048205]
Heser infumatus Kamura, 2019: 7, f. 1A-E (f)
Berinda idae Bosmans et al., 2019b: 12, f. 4a-g (mf)
Berinda infumatus Zamani et al., 2021a: 70 (T from Heser, S of Berinda idae)

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