World Spider Catalog

Anyphaenidae Bertkau, 1878

Family | ACCEPTED

N.B.: considered a senior synonym of Amaurobioididae by Platnick, 1974: 211; transfer of the monotypic subfamily Malenellinae Ramírez, 1995 to the Amaurobiidae by Wheeler et al., 2017: 605; mentioning of "Anyphaeniens" by Simon, 1864: 94, but the French ending -iens was not accepted as taxon at familiar level by Bonnet and Platnick, and this continued here.

In synonymy:

Gayenella Berland, 1913 = Josa Keyserling, 1891 (Ramírez, 2003: 108)

Haptisus Simon, 1897 = Josa Keyserling, 1891 (Ramírez, 2003: 108)

Olbophthalmus Simon, 1904 = Josa Keyserling, 1891 (Ramírez, 2003: 108)

Pelayo O. Pickard-Cambridge, 1896 = Josa Keyserling, 1891 (Brescovit, 1993a: 129)

Tetromma Keyserling, 1878 = Josa Keyserling, 1891 (Ramírez, 2003: 108, preoccupied, earliest available name)

Gen. Josa Keyserling, 1891 urn:lsid:nmbe.ch:spidergen:02139

N.B.: considered a senior synonym of Pelayo O. Pickard-Cambridge, 1896a: 194 [urn:lsid:nmbe.ch:spidergen:05025] (type P. laetus O. Pickard-Cambridge, 1896) by Brescovit, 1993a: 129; considered a junior synonym of Tetromma Keyserling, 1878a: 607 [urn:lsid:nmbe.ch:spidergen:02160] (type T. lutea Keyserling, 1878) (preoccupied in Coleoptera by Déjean, 1834) and a senior synonym of Haptisus Simon, 1897a: 100 [urn:lsid:nmbe.ch:spidergen:02131] (type H. nicoleti Simon, 1897), Olbophthalmus Simon, 1904e: 98 [urn:lsid:nmbe.ch:spidergen:02148] (type Olbus personatus Simon, 1897) and of Gayenella Berland, 1913b: 102 [urn:lsid:nmbe.ch:spidergen:02129] (type G. riveti Berland, 1913) by Ramírez, 2003: 108.

Transferred to other genera:

Josa alba Mello-Leitão, 1945 -- see Wulfila

Josa fusca (Caporiacco, 1947) -- see Rhoicinus

Josa lutescens Schmidt, 1971 -- see Elaver

Josa montana (Chickering, 1940) -- see Lepajan

In synonymy:

Josa insignis (Banks, 1909) = Josa nigrifrons Simon, 1897 (Ramírez, 2003: 117)

Josa nicoleti (Simon, 1897, T from Haptisus) = Josa lutea Keyserling, 1878 (Ramírez, 2003: 109)

Josa pilosa (Keyserling, 1879) = Josa lutea Keyserling, 1878 (Ramírez, 2003: 109)

Josa riveti (Berland, 1913, T from Gayenna) = Josa lutea Keyserling, 1878 (Ramírez, 2003: 109)

Nomen dubium:

Josa nigricans (Nicolet, 1849: 447, f, Chile, originally in Clubiona) -- Ramírez, 2003: 109

Josa analis (Simon, 1897) | | Venezuela urn:lsid:nmbe.ch:spidersp:023859

Haptisus analis Simon, 1897b: 504 (Dmf) Original description

Josa analis Dupérré, 2023a: 136, f. 13A-B, 14A-C (mf)

Josa andesiana (Berland, 1913) | | Ecuador urn:lsid:nmbe.ch:spidersp:023805

Gayenna andesiana Berland, 1913b: 101, pl. 10, f. 75-76 (Dmf) Original description

Josa andesiana Ramírez, 2003: 109 (Tmf from Gayenna)

Josa andesiana Dupérré, 2023a: 139, f. 15A-B, 16A-C (mf)

Josa andesiana Poy et al., 2023b: 6, f. 3e-f (m)

Josa bryantae (Caporiacco, 1955) | | Venezuela urn:lsid:nmbe.ch:spidersp:023906

Acanthoceto bryantae Caporiacco, 1955: 383, f. 49 (Df) Original description

Josa bryantae Ramírez, 1997: 190 (Tf from Clubionidae incertae sedis)

Josa calilegua Ramírez, 2003 | | Argentina urn:lsid:nmbe.ch:spidersp:038818

Josa calilegua Ramírez, 2003: 114, f. 54A-D, 58A-D (Dmf) Original description

Josa calilegua Poy et al., 2023b: 6, f. 3c-d, 7c-d (m)

Josa chazaliae (Simon, 1897) | | Colombia urn:lsid:nmbe.ch:spidersp:024066

Tomopisthes chazaliae Simon, 1897c: 11 (Df) Original description

Josa chazaliae Ramírez, 2003: 109 (Tf from Tomopisthes)

Josa gounellei (Simon, 1897) | | Brazil urn:lsid:nmbe.ch:spidersp:023951

Olbus gounellei Simon, 1897b: 506 (Dmf) Original description

Olbophthalmus gounellei Simon, 1904e: 98

Josa keyserlingi (L. Koch, 1866) | | Colombia, Brazil urn:lsid:nmbe.ch:spidersp:023826

Anyphaena keyserlingi L. Koch, 1866: 203, pl. 8, f. 136-138 (Dmf) Original description

Samuza keyserlingi Keyserling, 1891: 137

Josa keyserlingi Ramírez, 2003: 109 (Tmf from Gayenna)

Josa laeta (O. Pickard-Cambridge, 1896) | | Costa Rica urn:lsid:nmbe.ch:spidersp:023908

Pelayo laetus O. Pickard-Cambridge, 1896a: 194, pl. 24, f. 7 (Dm) Original description

Josa lojensis (Berland, 1913) | | Ecuador urn:lsid:nmbe.ch:spidersp:023952

Olbophthalmus lojensis Berland, 1913b: 104, pl. 11, f. 86-90 (Dmf) Original description

Josa lutea (Keyserling, 1878) | | Colombia, Ecuador urn:lsid:nmbe.ch:spidersp:024025

Anyphaena citrina L. Koch, 1866: 199, pl. 8, f. 133-135 (mf; misidentified per Ramírez, 2003: 109)

Tetromma lutea Keyserling, 1878a: 608 (Df) Original description

Anyphaena pilosa Keyserling, 1879: 327, pl. 4, f. 24 (Dmf) [urn:lsid:nmbe.ch:spidersp:023909]

Josa pilosa Keyserling, 1891: 83

Haptisus nicoleti Simon, 1897a: 92, 95, 100, f. 89 (Dm; p. 92 sub Anyphaena) [urn:lsid:nmbe.ch:spidersp:023861]

Gayenna riveti Berland, 1913b: 100, pl. 10, f. 73 (Df) [urn:lsid:nmbe.ch:spidersp:023843]

Haptisus nicoleti Berland, 1913b: 104, pl. 11, f. 83-85 (m, Df)

Josa lutea Ramírez, 2003: 109, f. 55A-D (mf, S of Gayenna riveti, Haptisus nicoleti and Josa pilosa)

Josa maura (Simon, 1897) | j | Venezuela urn:lsid:nmbe.ch:spidersp:023860

Haptisus maurus Simon, 1897b: 504 (Dj) Original description

Josa maura Dupérré, 2023a: 142, f. 17A-B (j)

Josa nigrifrons (Simon, 1897) | | Mexico to Bolivia urn:lsid:nmbe.ch:spidersp:023863

Haptisus nigrifrons Simon, 1897b: 505 (Df) Original description

Pelayo insignis Banks, 1909a: 199, pl. 6, f. 28 (Df) [urn:lsid:nmbe.ch:spidersp:023907]

Pelayo insignis Banks, 1914a: 677, pl. 28, f. 14 (f)

Josa nigrifrons Ramírez, 2003: 117, f. 59A-G, 60A-E (f, Dm, S of Josa insignis)

Josa personata (Simon, 1897) | | Ecuador urn:lsid:nmbe.ch:spidersp:023953

Olbus personatus Simon, 1897b: 506 (Df) Original description

Olbophthalmus personatus Simon, 1904e: 98

Josa personata Ramírez, 2003: 114, f. 57A-F (f)

Josa riveti (Berland, 1913) | | Ecuador, Bolivia urn:lsid:nmbe.ch:spidersp:023858

Gayenella riveti Berland, 1913b: 102, pl. 10, f. 77-79 (Df) Original description

Josa riveti Ramírez, 2003: 111, f. 56A-J (f, Dm, T from Gayenella)

Josa simoni (Berland, 1913) | | Ecuador urn:lsid:nmbe.ch:spidersp:023847

Gayenna simoni Berland, 1913b: 101, pl. 10, f. 74 (Dmf) Original description

Josa simoni Ramírez, 2003: 109 (Tmf from Gayenna)

Josa simoni Dupérré, 2023a: 144, f. 18A-B, 19A-C (mf)