Thomisidae Sundevall, 1833

Family |   ACCEPTED
N.B.: Includes Aphantochilidae as "a subfamily related to the Strophiinae" (Ono, 1988c: 217-218, after Homann, 1975: 184; see also Benjamin, 2011: 8 and Teixeira, Campos & Lise, 2014: 73). Separation of Borboropactidae Wunderlich, 2004 (Wunderlich, 2004m: 1738, Murphy & Roberts, 2015: viii) not sufficiently justified and is not accepted here. Lehtinen, 2016 splits the family into five families, i.e. Thomisidae s. str., Borboropactidae Wunderlich, 2004, Stephanopidae O. Pickard-Cambridge, 1871, Stiphropidae Simon, 1886 [sub Stiphropodidae; for nomenclatural author and year see Simon, 1886b: 170] and Bomidae Simon, 1886 [for nomenclatural author and year see Simon, 1886b: 173]; because of lack of a phylogenetic analysis and partly unclear limitations this splitting is not accepted; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 518, f. 17-18.

In synonymy:

Regillulus Strand, 1942 = Borboropactus Simon, 1884 (Roewer, 1955c: 752)

Regillus O. Pickard-Cambridge, 1884 = Borboropactus Simon, 1884 (Roewer, 1955c: 752, synonymized the replacement name Regillulus)

Gen. Borboropactus Simon, 1884 urn:lsid:nmbe.ch:spidergen:02641
N.B.: the older name Regillus O. Pickard-Cambridge, 1884a: 203 [urn:lsid:nmbe.ch:spidergen:04925] (type R. asper O. Pickard-Cambridge, 1884) is preoccupied in Aves by MacGillivray, 1839; Regillus was replaced with Regillulus [urn:lsid:nmbe.ch:spidergen:04926] by Strand, 1942: 399; synonmy of Borboropactus with Regillus considered by Simon, 1895a: 1055; synonymy of Regillulus with Borboropactus considered by Roewer, 1955c: 752 (see also Bonnet, 1955: 902 and 1958: 3848); Borboropactus was originally described in Bradystichidae (now a synonym of Pisauridae); placed in Borborpactidae fam. nov. by Wunderlich, 2004m: 1738, family status in fact rejected by Benjamin et al., 2008: 717 (including the Borboropactus clade in the Thomisidae), confirmed by Benjamin, 2011: 8; as Borboropactinae by Wheeler et al., 2017: 588.

In synonymy:

Borboropactus bangkongeus Barrion & Litsinger, 1995 = Borboropactus cinerascens Doleschall, 1859 (Benjamin et al., 2008: 722)

Borboropactus divergens (Hogg, 1914) = Borboropactus bituberculatus Simon, 1884 (Benjamin, 2011: 11)

Borboropactus hainanus Song, 1993 = Borboropactus bituberculatus Simon, 1884 (Benjamin, 2011: 11)

Borboropactus mindoroensis Barrion & Litsinger, 1995 = Borboropactus cinerascens Doleschall, 1859 (Benjamin et al., 2008: 722)

Borboropactus umaasaeus Barrion & Litsinger, 1995 = Borboropactus cinerascens Doleschall, 1859 (Benjamin et al., 2008: 722)

Nomen dubium:

Borboropactus cinerascens sumatrae (Strand, 1907m: 429, f, Indonesia/Sumatra, originally in Regillus) -- Nentwig et al., 2019: 47

Borboropactus asper (O. Pickard-Cambridge, 1884) | | Sri Lanka urn:lsid:nmbe.ch:spidersp:030180
Regillus asper O. Pickard-Cambridge, 1884a: 204, pl. 15, f. 5 (Dj) Original description
Regillus asper Simon, 1895a: 1055, f. 1094-1096
Regillulus asper Strand, 1942: 399 (superfluous generic replacement name)
Borboropactus asper Roewer, 1955c: 752 (T from Regillulus)
Borboropactus asper Marusik, Omelko & Benjamin, 2013: 197, f. 1-2, 4-8 (Df)
Borboropactus australis (Lawrence, 1937) | | South Africa urn:lsid:nmbe.ch:spidersp:030181
Regillus australis Lawrence, 1937: 237, f. 13 (Df) Original description
Borboropactus australis Roewer, 1955c: 752 (T from Regillulus)
Borboropactus australis Dippenaar-Schoeman et al., 2020u: 14, 2 f. (f)
Borboropactus biprocessus Tang, Yin & Peng, 2012 | | China urn:lsid:nmbe.ch:spidersp:046219
Borboropactus biprocessus Tang, Yin & Peng in Yin et al., 2012: 1257, f. 675a-c (Dm) Original description
Borboropactus bituberculatus Simon, 1884 | | India (Nicobar Is.), China (Hainan), Indonesia (Moluccas, New Guinea), Papua New Guinea urn:lsid:nmbe.ch:spidersp:030183
Borboropactus bituberculatus Simon, 1884e: 301 (Df) Original description
Regillus bituberculatus Simon, 1895a: 1049, f. 1093
Regillus divergens Hogg, 1914a: 57 (Df) [urn:lsid:nmbe.ch:spidersp:030186]
Regillus divergens Hogg, 1915a: 461, f. 29 (f)
Borboropactus bituberculatus Roewer, 1955c: 753 (T from Regillulus)
Borboropactus divergens Roewer, 1955c: 753 (T from Regillulus)
Borboropactus divergens Chrysanthus, 1964: 89, f. 12-14 (f)
Borboropactus hainanus Song, 1993: 89, f. 1a-c (Dm) [urn:lsid:nmbe.ch:spidersp:030188]
Borboropactus hainanus Song, 1994b: 119, f. 1A-E (Df, misidentified per Tang & Li, 2010a: 17)
Borboropactus hainanus Song & Zhu, 1997: 21, f. 8A-E, 9A-C (mf)
Borboropactus hainanus Song, Zhu & Chen, 1999: 480, f. 274G, 276J, 277I (mf)
Borboropactus hainanus Wunderlich, 2004m: 1744, f. 3 (m)
Borboropactus hainanus Wunderlich, 2008c: 483, f. 23 (f)
Borboropactus hainanus Tang & Li, 2010a: 16, f. 11A-B, 12A-D, 13A-D, 14A-D (m, Df)
Borboropactus bituberculatus Benjamin, 2011: 11, f. 5A, 20A-B, D (mf, S of Borboropactus divergens and B. hainanus)
Borboropactus bituberculatus Ramírez, 2014: 257, f. 172D (f)
Borboropactus bituberculatus Dash & Sivaperuman, 2021b: 5, f. 3a-f (f)
Borboropactus brevidens Tang & Li, 2010 | | China urn:lsid:nmbe.ch:spidersp:043014
Borboropactus brevidens Tang & Li, 2010a: 8, f. 3A-C, 4A-C, 5A-D, 6A-D (Dmf) Original description
Borboropactus cinerascens (Doleschall, 1859) | | Malaysia, Indonesia (Java, Sumatra), New Guinea, Philippines urn:lsid:nmbe.ch:spidersp:030184
Thomisus cinerascens Doleschall, 1859: 58, pl. 12, f. 4 (Df) Original description
Borboropactus cinerascens Simon, 1884e: 300
Regillus cinerascens Thorell, 1890d: 318
Regillus cinerascens Thorell, 1892a: 6 (Df)
Borboropactus cinerascens Roewer, 1955c: 753 (T from Regillulus)
Borboropactus mindoroensis Barrion & Litsinger, 1995: 203, f. 116a-g (Df) [urn:lsid:nmbe.ch:spidersp:030190]
Borboropactus umaasaeus Barrion & Litsinger, 1995: 203, f. 117a-i (Dm) [urn:lsid:nmbe.ch:spidersp:030194]
Borboropactus bangkongeus Barrion & Litsinger, 1995: 206, f. 118a-i (Dm) [urn:lsid:nmbe.ch:spidersp:030182]
Borboropactus cinerascens Benjamin et al., 2008: 722, f. 8A-E (mf, S of Borboropactus bangkongeus, B. mindoroensis and B. umaasaeus)
Borboropactus cinerascens Benjamin, 2011: 12, f. 18A-E (mf)
Borboropactus cinerascens Marusik, Omelko & Benjamin, 2013: 198, f. 9-10 (f)
Borboropactus cinerascens Lehtinen, 2016: 160, f. 5a-h (mf)
Borboropactus cinerascens Machado & Teixeira, 2021: 302, f. 12a, f, 13e, 18f (f)
Borboropactus edentatus Tang & Li, 2010 | | China urn:lsid:nmbe.ch:spidersp:043015
Borboropactus edentatus Tang & Li, 2010a: 12, f. 7A-C. 8A-C, 9A-D, 10A-D (Dmf) Original description
Borboropactus elephantus (Tikader, 1966) | | India urn:lsid:nmbe.ch:spidersp:030187
Regillus elephantus Tikader, 1966g: 54, f. 2a-b (Df) Original description
Regillus elephantus Tikader, 1968b: 114, f. 37-38 (f)
Regillus elephantus Tikader, 1971a: 64, f. 17G-H (f)
Regillus elephantus Tikader, 1980a: 167, f. 232-233 (f)
Borboropactus elephantus Brignoli, 1983c: 605
Borboropactus javanicola (Strand, 1913) | | Indonesia (Java) urn:lsid:nmbe.ch:spidersp:030189
Regillus javanicola Strand, 1913a: 397 (Df) Original description
Borboropactus javanicola Roewer, 1955c: 752 (T from Regillulus)
Borboropactus jiangyong Yin, Peng, Yan & Kim, 2004 | | China urn:lsid:nmbe.ch:spidersp:040711
Borboropactus jiangyong Yin et al., 2004b: 27, f. 1-5 (Df) Original description
Borboropactus jiangyong Yin et al., 2012: 1259, f. 676a-e (f)
Borboropactus jiangyong Meng et al., 2019: 140, f. 1A-H, 2A-G, 3A-H, 4A-G, 5A-C (Dmf)
Borboropactus longidens Tang & Li, 2010 | | China (Hainan), Vietnam urn:lsid:nmbe.ch:spidersp:043016
Borboropactus longidens Tang & Li, 2010a: 21, f. 15A-D, 16A-B (Df) Original description
Borboropactus longidens Li et al., 2023d: 172, f. 1A-J, 2A-G, 3A-J (Dmf)
Borboropactus nanda Lin & Li, 2023 | | China urn:lsid:nmbe.ch:spidersp:057158
Borboropactus nanda Lin & Li in Lin, Li & Pham, 2023: 66, f. 58A-C, 59A-B, 60A-B, 69A-B (Dmf) Original description
Borboropactus noditarsis (Simon, 1903) | j | West Africa urn:lsid:nmbe.ch:spidersp:030191
Regillus noditarsis Simon, 1903e: 101 (Dj) Original description
Borboropactus noditarsis Roewer, 1955c: 752 (T from Regillulus)
Borboropactus nyerere Benjamin, 2011 | | Tanzania urn:lsid:nmbe.ch:spidersp:044387
Borboropactus nyerere Benjamin, 2011: 12, f. 6A-B, 19A-E, 22A-F, 23A-F, 24A-F, 25A-F, 26A-D (Dmf) Original description
Borboropactus nyerere Machado, Teixeira & Lise, 2017: 454, f. S3B, S5E, S12A, S13C, S14A,C (m)
Borboropactus nyerere Machado & Teixeira, 2021: 298, f. 8h, 19e (m)
Borboropactus semenchenkoi Omelko & Marusik, 2022 | | Malaysia (Borneo) urn:lsid:nmbe.ch:spidersp:054533
Borboropactus semenchenkoi Omelko & Marusik, 2022a: 28, f. 1-15 (Dm) Original description
Borboropactus silvicola (Lawrence, 1938) | | South Africa urn:lsid:nmbe.ch:spidersp:030192
Regillus silvicolus Lawrence, 1938a: 488, f. 19 (Dmf) Original description
Borboropactus silvicolus Roewer, 1955c: 752 (T from Regillulus)
Borboropactus silvicola Dippenaar-Schoeman et al., 2020u: 15, 13: 2 f., 15: 5 f. (mf)
Borboropactus squalidus Simon, 1884 | | Zambia, South Africa urn:lsid:nmbe.ch:spidersp:030193
Borboropactus squalidus Simon, 1884e: 301 (Df) Original description
Regillus squalidus Simon, 1895a: 1055
Borboropactus squalidus Roewer, 1955c: 752 (T from Regillulus)
Borboropactus squalidus Benjamin, 2011: 53, f. 5B, 20C (f)
Borboropactus squalidus Dippenaar-Schoeman et al., 2020u: 16, 3 f. (f)
Borboropactus vulcanicus (Doleschall, 1859) | | Indonesia (Java) urn:lsid:nmbe.ch:spidersp:030195
Thomisus vulcanicus Doleschall, 1859: 57, pl. 13, f. 6 (Df) Original description
Regillus vulcanicus Simon, 1895a: 1055
Borboropactus vulcanicus Roewer, 1955c: 752 (T from Regillulus)

Sponsors