Theridiidae Sundevall, 1833

Family |   ACCEPTED
N.B.: considered a senior synonym of Hadrotarsidae Thorell, 1881 (removed from the synonymy of Oonopidae by Roewer, 1963a: 122, contra Roewer, 1942a: 287) by Forster, Platnick & Coddington, 1990: 111 (after Wunderlich, 1978a: 25 and Wunderlich, 1986: 95, contra Baert, 1984a: 615 and Davies, 1985: 67), who recognized an expanded Hadrotarsinae as well as a separate subfamily Spintharinae.

In synonymy:

Deliana Keyserling, 1886 = Lasaeola Simon, 1881 (Wunderlich, 1988: 148, by T of the type species)

Pachydactylus Menge, 1868 = Lasaeola Simon, 1881 (Simon, 1881a: 136, preoccupied, replacement name)

Pselothorax Chamberlin, 1949 = Lasaeola Simon, 1881 (Yoshida, 2002a: 13, contra Ivie, 1967: 126, who placed that name as a junior synonym of Dipoena Thorell, 1869)

Homonym replaced:

Pachydactylus Menge, 1868 -- see Lasaeola Simon, 1881 - in Simon, 1881a

Gen. Lasaeola Simon, 1881 urn:lsid:nmbe.ch:spidergen:00701
N.B.: replacement name for Pachydactylus Menge, 1868: 177 [urn:lsid:nmbe.ch:spidergen:05260], preoccupied in Reptilia by Wiegman, 1834; removed (together with its junior synonym Deliana Keyserling, 1886b: 35 [urn:lsid:nmbe.ch:spidergen:05261] (type D. spinithorax Keyserling, 1886)) from the synonymy of Dipoena Thorell, 1869 by Wunderlich, 1988: 148 but only taxa specifically mentioned by Wunderlich are shown as transfers here; considered by Yoshida, 2002a: 12 to include "the species of the D. prona group designated by Levi (1953)" but again only taxa explicitly transferred by him are listed here, until a wider, worldwide sample of taxa can be studied.

Transferred to other genera:

Lasaeola proxima O. Pickard-Cambridge, 1896 -- see Dipoena

In synonymy:

Lasaeola daltoni (Levi, 1953, T from Dipoena) = Lasaeola atopa Chamberlin, 1949 (Ivie, 1967: 126, sub Dipoena)

Lasaeola hamata (Tullgren, 1949, T from Dipoena) = Lasaeola prona Menge, 1868 (Levi, 1963a: 145, sub Dipoena)

Lasaeola tibiale (Hahn, 1831, T from Theridion) = Lasaeola tristis Hahn, 1833 (Prpic, 2015: 88)

Lasaeola trapezoidalis (Levy & Amitai, 1981, T from Dipoena) = Lasaeola convexa Blackwall, 1870 (Vanuytven, Van Keer & Poot, 1994: 2, sub Dipoena)

Lasaeola algarvensis Wunderlich, 2011 | | Portugal urn:lsid:nmbe.ch:spidersp:044106
Lasaeola algarvensis Wunderlich, 2011: 257, f. 68-73 (Dmf) Original description
Lasaeola armona Wunderlich, 2015 | | Portugal, Spain urn:lsid:nmbe.ch:spidersp:047926
Lasaeola armona Wunderlich, 2015a: 442, f. 8-10 (Dm) Original description
Lasaeola armona Wunderlich, 2023: 31, f. 44 (Df)
Lasaeola atopa (Chamberlin, 1949) | | USA urn:lsid:nmbe.ch:spidersp:007346
Pselothorax atopus Chamberlin, 1949: 541 (Dm) Original description
Dipoena daltoni Levi, 1953b: 35, f. 65-71, 116-117 (Dmf)
Dipoena atopa Ivie, 1967: 126 (S)
Dipoena atopa Levi & Randolph, 1975: 37, f. 44 (m)
Dipoena atopa Roth, 1982: 47-9, f. 44 (m)
Dipoena atopa Roth, 1985: B43-9, f. 44 (m)
Dipoena atopa Roth, 1994: 183, f. 44 (m)
Lasaeola atopa Yoshida, 2002a: 13 (implicitly Tmf from Dipoena by synonymy of generic name)
Lasaeola bequaerti (Chickering, 1948) | | Panama urn:lsid:nmbe.ch:spidersp:007791
Dipoena bequaerti Chickering, 1948: 331, f. 1-3 (Dm) Original description
Dipoena bequaerti Levi, 1963a: 178, f. 266-267 (m)
Dipoena becquaerti Brignoli, 1983c: 401 (lapsus)
Lasaeola bequaerti Wunderlich, 1988: 154 (Tm from Dipoena)
Lasaeola canariensis (Wunderlich, 1987) | | Canary Is. urn:lsid:nmbe.ch:spidersp:007518
Dipoena canariensis Wunderlich, 1987a: 210, f. 552-556 (Dmf) Original description
Dipoenata canariensis Wunderlich, 1988: 154 (Tmf from Dipoena)
Lasaeola canariensis Wunderlich, 2011: 256 (Tmf from Dipoenata)
Lasaeola convexa (Blackwall, 1870) | | Mediterranean urn:lsid:nmbe.ch:spidersp:007367
Theridion convexum Blackwall, 1870b: 426 (Df) Original description
Euryopis pilula Simon, 1873a: 119 (Dmf)
Steatoda convexa Thorell, 1875c: 58
Lasaeola convexa Simon, 1881a: 148 (m)
Lasaeola convexa Chyzer & Kulczyński, 1894: 26, pl. 1, f. 23 (mf)
Dipoena convexa Simon, 1894a: 526, f. 572
Dipoena convexa Simon, 1914: 274, 302, f. 528 (mf)
Dipoena convexa Brignoli, 1967e: 183, f. 3A-C (f)
Dipoena trapezoidalis Levy & Amitai, 1981a: 184, f. 23-27 (Dm)
Dipoena convexa Vanuytven, Van Keer & Poot, 1994: 2, f. 1-5 (mf, S)
Dipoena convexa Levy, 1998b: 154, f. 287-293 (mf)
Dipoena convexa Knoflach & Pfaller, 2004: 142, f. 24f (f)
Lasaeola convexa Wunderlich, 2011: 256, f. 85-87 (Tmf from Dipoena)
Dipoena convexa Le Peru, 2011: 439, f. 675 (mf)
Lasaeola convexa IJland, van Helsdingen & Miller, 2012: 14, f. 18a-b (m)
Lasaeola convexa Gaymard & Lecigne, 2018: 39, f. 16H-I, 17G-H (f)
Lasaeola convexa Wunderlich, 2020a: 11, f. 15 (prosoma)
Lasaeola convexa Iorio, 2023: 30, f. 24-25 (m)
Lasaeola coracina (C. L. Koch, 1837) | | Western Europe to Ukraine, Iran urn:lsid:nmbe.ch:spidersp:007368
Theridium coracinum C. L. Koch, 1837b: 8 (D) Original description
Theridium coracinum C. L. Koch, 1841a: 84, f. 655 (Df)
Steatoda coracina C. L. Koch, 1850: 18
Steatoda coracina O. Pickard-Cambridge, 1879d: 98 (Dm)
Lasaeola nigrina Simon, 1881a: 144 (Dmf)
Lasaeola nigrina Chyzer & Kulczyński, 1894: 26, pl. 1, f. 22 (mf)
Lasaeola nigrina Bösenberg, 1902: 123, pl. 11, f. 156 (mf)
Dipoena nigrina Lessert, 1910b: 117
Dipoena coracina Simon, 1914: 277, 301, f. 528 (mf)
Dipoena coracina Miller, 1947: 62, pl. IV, f. 3-4 (mf)
Dipoena coracina Locket & Millidge, 1953: 48, f. 33B, E (mf)
Dipoena coracina Miller, 1967: 290, pl. XIII, f. 10-15 (mf)
Dipoena coracina Miller, 1971: 184, pl. XXXI, f. 16-17 (mf)
Dipoena coracina Palmgren, 1974b: 11, f. 3.10-11 (mf)
Dipoena coracina Roberts, 1985: 176, f. 78e (mf)
Dipoena coracina Heimer & Nentwig, 1991: 286, f. 769 (mf)
Dipoena coracina Roberts, 1998: 284, f. (mf)
Dipoena coracina Le Peru, 2011: 439, f. 676 (mf)
Lasaeola coracina Wunderlich, 2015a: 439, f. 1-7 (Tmf from Dipoena)
Lasaeola coracina Lecigne, 2018b: 77, f. 11C (m)
Lasaeola coracina Zamani et al., 2021b: 289, f. 10K-L (m)
Lasaeola dbari Kovblyuk, Marusik & Omelko, 2012 | | Turkey, Russia (Europe), Georgia urn:lsid:nmbe.ch:spidersp:045528
Lasaeola dbari Kovblyuk, Marusik & Omelko, 2012: 248, f. 1-18 (Dmf) Original description
Lasaeola dbari Ponomarev & Shmatko, 2020b: 307, f. 30-33 (mf)
Lasaeola dbari Demircan Aksan & Topçu, 2022b: 37, f. 1A-D (f)
Lasaeola donaldi (Chickering, 1943) | | Panama, Venezuela urn:lsid:nmbe.ch:spidersp:007792
Dipoena donaldi Chickering, 1943b: 347, f. 24-26 (Dm) Original description
Dipoena donaldi Levi, 1963a: 165, f. 163-168 (m, Df)
Lasaeola donaldi Wunderlich, 1988: 150 (Tmf from Dipoena)
Lasaeola fastigata Zhang, Liu & Zhang, 2011 | | China urn:lsid:nmbe.ch:spidersp:044259
Lasaeola fastigata Zhang, Liu & Zhang, 2011b: 6, f. 4-6 (Dm) Original description
Lasaeola flavitarsis (Wunderlich, 1992) | | Canary Is. urn:lsid:nmbe.ch:spidersp:007520
Dipoenata flavitarsis Wunderlich, 1992a: 409, f. 572-573 (Df) Original description
Lasaeola flavitarsis Wunderlich, 2011: 254 (Tf from Dipoenata)
Lasaeola grancanariensis (Wunderlich, 1987) | | Canary Is. urn:lsid:nmbe.ch:spidersp:007390
Dipoena grancanariensis Wunderlich, 1987a: 212, f. 564-565 (Df) Original description
Lasaeola grancanariensis Wunderlich, 2011: 256 (Tf from Dipoena, may be a S of L. striata)
Lasaeola lunata Zhang, Liu & Zhang, 2011 | | China urn:lsid:nmbe.ch:spidersp:044258
Lasaeola lunata Zhang, Liu & Zhang, 2011b: 4, f. 1-3 (Dm) Original description
Lasaeola minutissima Wunderlich, 2011 | | Portugal, Spain urn:lsid:nmbe.ch:spidersp:044107
Lasaeola minutissima Wunderlich, 2011: 258, f. 79-84 (Dmf) Original description
Lasaeola minutissima Barrientos et al., 2016: 96, f. 26-30 (mf)
Lasaeola oceanica Simon, 1883 | | Azores urn:lsid:nmbe.ch:spidersp:007793
Lasaeola testaceomarginata oceanica Simon, 1883: 271 (Dmf) Original description
Lasaeola oceanica Wunderlich, 1992a: 412, f. 574-575 (mf, elevated from subspecies)
Lasaeola octoginta Wunderlich, 2020 | | Portugal urn:lsid:nmbe.ch:spidersp:051653
Lasaeola octoginta Wunderlich, 2020a: 12, f. 18-20 (Dm) Original description
Lasaeola okinawana (Yoshida & Ono, 2000) | | China, Japan (Ryukyu Is.) urn:lsid:nmbe.ch:spidersp:037045
Dipoena amamiensis Yoshida, 1991e: 35, f. 5-6 (m, misidentified)
Dipoena amamiensis Zhu, 1998: 242, f. 159E (m, misidentified)
Dipoena amaniensis Song, Zhu & Chen, 1999: 110, f. 53M (m, misidentified)
Dipoena okinawana Yoshida & Ono, 2000: 146, f. 33-36 (Dm) Original description
Lasaeola okinawana Yoshida, 2002a: 13 (Tm from Dipoena)
Lasaeola okinawana Yoshida, 2003a: 172, f. 472-475 (m)
Lasaeola okinawana Yoshida, 2009b: 389, f. 328 (m)
Lasaeola pinna Tang, F. Liu, Z. Y. Liu, Yang & Peng, 2024 | | China urn:lsid:nmbe.ch:spidersp:060694
Lasaeola pinna Tang et al., 2024: 380, f. 1A-F, 2A-E (Dmf) Original description
Lasaeola prona (Menge, 1868) | | North America, Europe, Caucasus, Russia (Europe to South Siberia), Kazakhstan, Iran, Japan urn:lsid:nmbe.ch:spidersp:007461
Pachydactylus pronus Menge, 1868: 177, pl. 33, f. 80 (Dmf) Original description
Theridion pronum O. Pickard-Cambridge, 1873c: 535
Steatoda prona O. Pickard-Cambridge, 1874d: 324
Euryopis prona O. Pickard-Cambridge, 1881a: 481
Lasaeola prona Simon, 1881a: 145
Lasaeola prona Chyzer & Kulczyński, 1894: 25, pl. 1, f. 20 (mf)
Lasaeola prona Bösenberg, 1902: 124, pl. 11, f. 159 (f)
Lasaeola jucunda O. Pickard-Cambridge, 1904a: 152, 162, pl. A, f. 3 (Dmf)
Lasaeola jucunda O. Pickard-Cambridge, 1908a: 125, pl. A, f. 13-15 (mf)
Dipoena prona Simon, 1914: 277, 302, f. 528 (mf)
Dipoena prona Wiehle, 1937: 186, f. 184-185 (mf)
Dipoena prona Tullgren, 1949: 50, f. 9 (f)
Dipoena hamata Tullgren, 1949: 50, f. 10, 11a-b (Dm)
Dipoena prona Locket & Millidge, 1953: 46, f. 31A-B, 32A (mf)
Dipoena hamata Levi, 1953b: 30, f. 50-59, 105-106 (mf)
Dipoena prona Lehtinen & Kleemola, 1962: 110, f. 14 (f)
Dipoena prona Levi & Levi, 1962: 50 (m)
Dipoena prona Levi, 1963a: 145 (S)
Dipoena prona Miller, 1967: 290, pl. XIII, f. 16-19 (mf)
Dipoena prona Tystshenko, 1971: 143, f. 372, 375 (mf)
Dipoena prona Miller, 1971: 185, pl. XXXI, f. 14-15 (mf)
Dipoena prona Palmgren, 1974b: 11, f. 2.27-29 (mf)
Dipoena hamata Palmgren, 1974b: 11, f. 3.4-5 (mf, after Levi, 1953b)
Dipoena prona Roberts, 1985: 176, f. 78b (mf)
Dipoena prona Heimer & Nentwig, 1991: 284, f. 765 (mf)
Dipoena prona Roberts, 1995: 269, f. (mf)
Dipoena prona Roberts, 1998: 284, f. (mf)
Dipoena prona Yoshida & Ono, 2000: 149, f. 40-47 (mf)
Lasaeola prona Yoshida, 2002a: 13, f. 3-5 (Tmf from Dipoena)
Dipoena prona Paquin & Dupérré, 2003: 215, f. 2402-2404 (mf)
Lasaeola prona Yoshida, 2003a: 171, f. 466-471 (mf)
Dipoena prona Almquist, 2005: 62, f. 88a-f (mf)
Lasaeola prona Yoshida, 2009b: 389, f. 326-327 (mf)
Dipoena prona Le Peru, 2011: 442, f. 687 (mf)
Lasaeola prona Kovblyuk, Marusik & Omelko, 2012: 250, f. 19-22 (m)
Lasaeola prona Barrientos, Uribarri & García-Sarrión, 2015: 66, f. 7a-b (m)
Lasaeola prona Lecigne, 2020: 50, pl. 1K (f)
Lasaeola prona Vanuytven, 2021: 172, f. B.161e, B.164 (m)
Lasaeola prona Seropian et al., 2023: 285, f. 130 (m)
Lasaeola spinithorax (Keyserling, 1886) | | Peru urn:lsid:nmbe.ch:spidersp:007794
Deliana spinithorax Keyserling, 1886b: 35, pl. 11, f. 153 (Df) Original description
Deliana spinithorax Levi & Levi, 1962: 19, f. 58-59 (f)
Dipoena spinithorax Levi, 1963a: 170, f. 205-208 (f)
Lasaeola spinithorax Wunderlich, 1988: 148 (Tf from Dipoena)
Lasaeola striata (Wunderlich, 1987) | | Canary Is. urn:lsid:nmbe.ch:spidersp:007491
Dipoena striata Wunderlich, 1987a: 211, f. 557-563 (Dmf) Original description
Lasaeola striata Wunderlich, 2011: 256, f. 74-75 (Tmf from Dipoena)
Lasaeola superba (Chickering, 1948) | | Mexico, Panama urn:lsid:nmbe.ch:spidersp:007795
Dipoena superba Chickering, 1948: 337, f. 8-10 (Dm) Original description
Dipoena superba Levi, 1963a: 167, f. 183-184 (m)
Lasaeola superba Wunderlich, 1988: 152 (Tm from Dipoena)
Lasaeola tengchongensis Tang, F. Liu, Z. Y. Liu, Yang & Peng, 2024 | | China urn:lsid:nmbe.ch:spidersp:060695
Lasaeola tengchongensis Tang et al., 2024: 383, f. 3A-F, 4A-E (Dmf) Original description
Lasaeola testaceomarginata Simon, 1881 | | Mediterranean urn:lsid:nmbe.ch:spidersp:007796
Lasaeola testaceomarginata Simon, 1881a: 142 (Dmf) Original description
Dipoena testaceomarginata Simon, 1914: 276, 301, f. 528 (mf)
Dipoena testaceomarginata Brignoli, 1968d: 97, f. 7-8, 11 (mf)
Lasaeola testaceomarginata Wunderlich, 1992a: 412, f. 576 (Tmf from Dipoena)
Lasaeola testaceomarginata Trotta, 2005: 176, f. 476-477 (mf)
Lasaeola testaceomarginata Wunderlich, 2011: 256, f. 76-78 (mf)
Dipoena testaceomarginata Le Peru, 2011: 442, f. 690 (mf)
Lasaeola testaceomarginata Lecigne, 2017: 35, f. 11D-F (f)
Lasaeola testaceomarginata Wunderlich, 2021: 11, f. 31-32 (m)
Lasaeola testaceomarginata Febrer & Barrientos, 2022: 140, f. 4A-C (m)
Lasaeola tristis (Hahn, 1833) | | Europe, Turkey, Russia (Europe to South Siberia), Kazakhstan, Iran, Central Asia urn:lsid:nmbe.ch:spidersp:007797
Theridion tibiale Hahn, 1831b: 1, pl. 24, f. dD4 (Df; nomen oblitum per Prpic, 2015: 87) [urn:lsid:nmbe.ch:spidersp:048037]
Theridion triste Hahn, 1833a: 89, f. 67 (Dmf) Original description
Theridium triste C. L. Koch, 1841a: vol. 8: 83, f. 653-654 (mf)
Steatoda tristis C. L. Koch, 1850: 18
Euryopis tristis Menge, 1868: 176, pl. 33, f. 79 (j)
Lasaeola tristis Simon, 1881a: 138
Lasaeola tristis Becker, 1896: (2): 110, pl. 24, f. 2 (mf)
Lasaeola tristis Bösenberg, 1902: 122, pl. 11, f. 155 (mf)
Dipoena tristis Simon, 1914: 276, 301
Dipoena tristis Wiehle, 1937: 182, f. 175-179 (mf)
Dipoena tristis Tullgren, 1949: 48, f. 7a, 8a-b (mf)
Dipoena tristis Locket & Millidge, 1953: 48, f. 33A, D (mf)
Dipoena tristis Levi, 1953b: 7, f. 35, 89 (mf)
Theridion tibiale Roewer, 1955c: 1685 (nicht zu deuten = nomen dubium)
Dipoena tristis Miller, 1967: 289, pl. XIII, f. 1-4 (mf)
Dipoena tristis Miller, 1971: 184, pl. XXXI, f. 12-13 (mf)
Dipoena tristis Palmgren, 1974b: 13, f. 2.23-26 (mf)
Dipoena tristis Roberts, 1985: 176, f. 78d (mf)
Lasaeola tristis Wunderlich, 1988: 148 (T from Dipoena; mistakenly as type species of Lasaeola)
Dipoena tristis Heimer & Nentwig, 1991: 286, f. 770 (mf)
Dipoena tristis Roberts, 1995: 269, f. (mf)
Dipoena tristis Roberts, 1998: 284, f. (mf)
Lasaeola tristis Knoflach & Pfaller, 2004: 137, f. 20f (m)
Lasaeola tristis Trotta, 2005: 177, f. 478-479 (mf)
Lasaeola tristis Almquist, 2005: 75, f. 98a-f (mf)
Lasaeola tristis Agnarsson, Coddington & Knoflach, 2007: 382, f. 173 (m)
Dipoena tristis Le Peru, 2011: 443, f. 692 (mf)
Lasaeola tristis Kovblyuk, Marusik & Omelko, 2012: 250, f. 23-26 (m)
Lasaeola tristis Prpic, 2015: 88 (S of Theridion tibiale, removed from nomen dubium)
Lasaeola tristis Aakra et al., 2016: 32, f. 27 (m)
Lasaeola tristis hissariensis (Charitonov, 1951) | | Russia (South Siberia) urn:lsid:nmbe.ch:spidersp:007798
Dipoena tristis hissariensis Charitonov, 1951: 210, f. 1 (Df) Original description
Lasaeola yona (Yoshida & Ono, 2000) | | Japan (Ryukyu Is.) urn:lsid:nmbe.ch:spidersp:037046
Dipoena yona Yoshida & Ono, 2000: 153, f. 54-56 (Dm) Original description
Lasaeola yona Yoshida, 2002a: 13 (Tm from Dipoena)
Lasaeola yona Yoshida, 2003a: 173, f. 476-478 (m)
Lasaeola yona Yoshida, 2009b: 391, f. 329 (m)
Lasaeola yoshidai (Ono, 1991) | | Russia (Far East), China, Korea, Japan urn:lsid:nmbe.ch:spidersp:007513
Dipoena yoshidai Ono in Ono et al., 1991: 91, f. 2-8 (Dmf) Original description
Dipoena yoshidai Yoshida & Ono, 2000: 152, f. 48-53 (mf)
Lasaeola yoshidai Yoshida, 2002a: 13 (Tmf from Dipoena)
Lasaeola yoshidai Yoshida, 2003a: 174, f. 479-482 (mf)
Lasaeola yoshidai Yoshida, 2009b: 391, f. 330-331 (mf)
Lasaeola yoshidai Seo, 2010a: 175, f. 11-14 (m)
Lasaeola yoshidai Suzuki, 2018b: 58, 1a-b (m)
Lasaeola yoshidai Kim, 2021: 77, f. 31A-E (f)
Lasaeola yoshidai Hidaka, 2022c: 172, f. 1-2 (f)

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