World Spider Catalog

Pholcidae C. L. Koch, 1850

Family | ACCEPTED

N.B.: For comments on the monophyly and relationships of many genera see Huber, 1998d and Huber, 2000. The division of Pholcidae into five subfamilies, Ninetinae Simon, 1890, Arteminae Simon, 1893, Modisiminae Simon, 1893, Smeringopinae Simon, 1893 and Pholcinae, is confirmed by Huber, Eberle & Dimitrov, 2018.

In synonymy:

Ceratopholcus Spassky, 1934 = Crossopriza Simon, 1893 (Huber, Colmenares & Ramírez, 2014: 420)

Tibiosa González-Sponga, 2006 = Crossopriza Simon, 1893 (Huber, 2009a: 65)

Gen. Crossopriza Simon, 1893 urn:lsid:nmbe.ch:spidergen:00382

N.B.: considered a senior synonym of Tibiosa González-Sponga, 2006: 10 [urn:lsid:nmbe.ch:spidergen:03802] (type T. caracensis González-Sponga, 2006) by Huber, 2009a: 65 and of Ceratopholcus Spassky, 1934a: 361 [urn:lsid:nmbe.ch:spidergen:00378] (type C. maculipes Spassky, 1934) by Huber, Colmenares & Ramírez, 2014: 420.

Transferred to other genera:

Crossopriza cylindrogaster Simon, 1907 -- see Smeringopus

Crossopriza saltensis Mello-Leitão, 1941 -- see Priscula

Crossopriza sexsignata Franganillo, 1926 -- see Artema

In synonymy:

Crossopriza brasiliensis Mello-Leitão, 1935 = Crossopriza lyoni Blackwall, 1867 (Huber, Deeleman-Reinhold & Pérez, 1999: 2)

Crossopriza caracensis (González-Sponga, 2006) = Crossopriza lyoni Blackwall, 1867 (Huber, 2009a: 65)

Crossopriza casanaimensis (González-Sponga, 2006) = Crossopriza lyoni Blackwall, 1867 (Huber, 2009a: 65)

Crossopriza coreana (González-Sponga, 2006) = Crossopriza lyoni Blackwall, 1867 (Huber, 2009a: 65)

Crossopriza francoisi Millot, 1946 = Crossopriza lyoni Blackwall, 1867 (Huber, Deeleman-Reinhold & Pérez, 1999: 2)

Crossopriza guayanesa (González-Sponga, 2006) = Crossopriza lyoni Blackwall, 1867 (Huber, 2009a: 65)

Crossopriza moraensis (González-Sponga, 2006) = Crossopriza lyoni Blackwall, 1867 (Huber, 2009a: 65)

Crossopriza mucronata Mello-Leitão, 1942 = Crossopriza lyoni Blackwall, 1867 (Huber, Deeleman-Reinhold & Pérez, 1999: 2)

Crossopriza nigrescens Millot, 1946 = Crossopriza lyoni Blackwall, 1867 (Huber, 2022: 193)

Crossopriza stridulans Millot, 1946 = Crossopriza lyoni Blackwall, 1867 (Huber, Deeleman-Reinhold & Pérez, 1999: 2)

Crossopriza dhofar Huber, 2022 | | Oman urn:lsid:nmbe.ch:spidersp:053790

Crossopriza dhofar Huber, 2022: 122, f. 396-398, 450-474 (Dmf) Original description

Crossopriza ghul Huber, 2022 | | Oman urn:lsid:nmbe.ch:spidersp:053793

Crossopriza ghul Huber, 2022: 137, f. 481-482, 526-537 (Dmf) Original description

Crossopriza ibnsinai Huber, 2022 | | Kazakhstan, Uzbekistan, Turkmenistan, Tajikistan, Afghanistan urn:lsid:nmbe.ch:spidersp:053794

Crossopriza ibnsinai Huber, 2022: 139, f. 538-554 (Dmf) Original description

Crossopriza illizi Huber, 2022 | | Algeria urn:lsid:nmbe.ch:spidersp:053787

Crossopriza illizi Huber, 2022: 107, f. 373-390 (Dmf) Original description

Crossopriza johncloudsleyi Deeleman-Reinhold & van Harten, 2001 | | Yemen, Kenya urn:lsid:nmbe.ch:spidersp:037485

Crossopriza johncloudsleyi Deeleman-Reinhold & van Harten, 2001: 197, f. 11-16 (Dmf) Original description

Crossopriza johncloudsleyi Huber & Warui, 2012: 18, f. 28-31, 62-63, 70-74 (mf)

Crossopriza kandahar Huber, 2022 | | Afghanistan urn:lsid:nmbe.ch:spidersp:053798

Crossopriza kandahar Huber, 2022: 168, f. 664-680 (Dmf) Original description

Crossopriza khayyami Huber, 2022 | | Turkey, Iraq, Iran, Afghanistan urn:lsid:nmbe.ch:spidersp:053795

Crossopriza khayyami Huber, 2022: 144, f. 555-573 (Dmf) Original description

Crossopriza kittan Huber, 2022 | | Oman urn:lsid:nmbe.ch:spidersp:053792

Crossopriza kittan Huber, 2022: 133, f. 479-480, 509-525 (Dmf) Original description

Crossopriza lyoni (Blackwall, 1867) | | Probably native to Africa and/or Asia. Introduced to the Americas, Belgium, Germany, Australia, Micronesia urn:lsid:nmbe.ch:spidersp:003389

Pholcus lyoni Blackwall, 1867a: 392 (Dmf) Original description

Smeringopus lyoni Thorell, 1895b: 70

Crossopriza lyoni Pocock, 1900a: 240

Crossopriza lyoni Dyal, 1935: 168, pl. 15, f. 90-96 (mf)

Crossopriza brasiliensis Mello-Leitão, 1935a: 94, f. 13 (Dmf)

Crossopriza mucronata Mello-Leitão, 1942b: 389, f. 1-2 (Df)

Crossopriza francoisi Millot, 1946a: 154, f. 29, 30B (Dm)

Crossopriza stridulans Millot, 1946a: 156, f. 31A-B (Df)

Crossopriza nigrescens Millot, 1946a: 155, f. 30A (Dj) [urn:lsid:nmbe.ch:spidersp:003390]

Crossopriza lyoni Zhu & Wang, 1963a: 462, f. 4, 7-8 (mf)

Crossopriza lyoni Chrysanthus, 1967b: 96, f. 20-24 (mf)

Crossopriza lyoni Tikader & Biswas, 1981: 18, f. 13-15 (f)

Crossopriza lyoni Yaginuma, 1986a: 31, f. 17.5 (mf)

Crossopriza lyoni Kim, 1988: 35, f. 1-6 (mf)

Crossopriza lyoni Chikuni, 1989b: 29, f. 7 (mf)

Crossopriza lyoni Chen & Zhang, 1991: 72, f. 62.1-3 (mf)

Crossopriza lyoni Huber, Deeleman-Reinhold & Pérez, 1999: 2, f. 1-12 (mf, S)

Crossopriza lyoni Song, Zhu & Chen, 1999: 52, f. 22H-O (mf)

Crossopriza lyoni Huber, 2000: 30, f. 98

Crossopriza lyoni Deeleman-Reinhold & van Harten, 2001: 195, f. 5-6 (mf)

Crossopriza lyoni Irie, 2001: 7, f. 1-8 (mf)

Crossopriza lyoni Van Keer & Van Keer, 2001: 83, f. 2.1-8 (mf)

Tibiosa caracensis González-Sponga, 2006: 11, pl. 1, f. 1-10 (Dmf) [urn:lsid:nmbe.ch:spidersp:041418]

Tibiosa casanaimensis González-Sponga, 2006: 14, pl. 2, f. 1-9 (Dmf) [urn:lsid:nmbe.ch:spidersp:041419]

Tibiosa coreana González-Sponga, 2006: 17, pl. 3, f. 1-7 (Dm) [urn:lsid:nmbe.ch:spidersp:041420] [f, f. 8-9: misidentified Artema atlanta]

Tibiosa guayanesa González-Sponga, 2006: 20, pl. 4, f. 1-9 (Dmf) [urn:lsid:nmbe.ch:spidersp:041421]

Tibiosa moraensis González-Sponga, 2006: 23, pl. 5, f. 1-9 (Dmf) [urn:lsid:nmbe.ch:spidersp:041422]

Crossopriza lyoni Beatty, Berry & Huber, 2008: 9, f. 17-18, 47-48 (mf)

Crossopriza lyoni Colmenares-García, 2008: 88, f. 2A-C (m)

Crossopriza lyoni Irie, 2009: 106, f. 3-5 (mf)

Crossopriza lyoni Huber, 2009a: 65 (S)

Crossopriza lyoni Yin et al., 2012: 163, f. 30a-h (mf)

Crossopriza lyoni Sen et al., 2015: 88, f. 514-518, pl. 19 (f)

Crossopriza lyoni Bauer et al., 2016: 4, f. 1-7 (mf)

Crossopriza lyoni Huber, 2022: 193, f. 702-705, 783-805 (mf, S of Crossopriza nigrescens)

Crossopriza lyoni Bourque et al., 2022: 28, f. 1-11 (mf)

Crossopriza lyoni Suzuki, 2022b: 193, f. 16-17 (f)

Crossopriza maculipes (Spassky, 1934) | | Iraq, Kazakhstan, Uzbekistan, Turkmenistan, Tajikistan, Afghanistan, Pakistan urn:lsid:nmbe.ch:spidersp:003356

Ceratopholcus maculipes Spassky, 1934a: 361, f. 1-3 (Dmf) Original description

Crossopriza maculipes Huber, Colmenares & Ramírez, 2014: 420 (T from Ceratopholcus=Crossopriza)

Crossopriza maculipes Ali, Zahid & Butt, 2016: 250, f. 3A-C (m)

Crossopriza maculipes Najim & Al-Khazali, 2019: 549, f. 2A-I, 3A-D (mf)

Crossopriza maculipes Huber, 2022: 208, f. 823-840, 854-865 (mf)

Crossopriza malegaon Huber, 2022 | | India urn:lsid:nmbe.ch:spidersp:053799

Crossopriza malegaon Huber, 2022: 172, f. 681-697 (Dmf) Original description

Crossopriza manakhah Huber, 2022 | | Yemen urn:lsid:nmbe.ch:spidersp:053796

Crossopriza manakhah Huber, 2022: 160, f. 625-642 (Dmf) Original description

Crossopriza miskin Huber, 2022 | | Oman urn:lsid:nmbe.ch:spidersp:053800

Crossopriza miskin Huber, 2022: 175, f. 698-701, 706-725 (Dmf) Original description

Crossopriza moqal Huber, 2022 | | Oman urn:lsid:nmbe.ch:spidersp:053791

Crossopriza moqal Huber, 2022: 127, f. 475-478, 483-508 (Dmf) Original description

Crossopriza parsa Huber, 2022 | | Iran urn:lsid:nmbe.ch:spidersp:053801

Crossopriza parsa Huber, 2022: 180, f. 726-744 (Dmf) Original description

Crossopriza pristina (Simon, 1890) | | Sudan, Eritrea, Ethiopia, Yemen urn:lsid:nmbe.ch:spidersp:003391

Artema pristina Simon, 1890a: 93 (Dmf) Original description

Crossopriza pristina Simon, 1893a: 476, f. 465 (f)

Crossopriza pristina Wiehle, 1933: 242, f. 3a (m)

Crossopriza pristina Timm, 1976: 70, f. 5-6 (m, misidentified)

Crossopriza pristina Siyam, Dunlop & El-Hennawy, 2015: 265, f. 2-6 (mf)

Crossopriza pristina Huber, 2022: 155, f. 608-624 (mf)

Crossopriza sahtan Huber, 2022 | | Oman urn:lsid:nmbe.ch:spidersp:053788

Crossopriza sahtan Huber, 2022: 110, f. 391-392, 399-432 (Dmf) Original description

Crossopriza sanaa Huber, 2022 | | Yemen urn:lsid:nmbe.ch:spidersp:053797

Crossopriza pristina Deeleman-Reinhold & van Harten, 2001: 195, f. 1-4, 7-10 (mf; misidentified per Huber, 2022: 164)

Crossopriza sanaa Huber, 2022: 164, f. 643-663 (Dmf) Original description

Crossopriza semicaudata (O. Pickard-Cambridge, 1876) | | Egypt, Chad, Sudan urn:lsid:nmbe.ch:spidersp:003392

Pholcus semicaudatus O. Pickard-Cambridge, 1876b: 565 (Dmf) Original description

Holocnemus semicaudatus Simon, 1907e: 5

Crossopriza semicaudata Wiehle, 1933: 242, f. 3b (m)

Crossopriza semicaudata Denis, 1945b: 48, f. 9 (m)

Crossopriza semicaudata Huber, 2022: 149, f. 574-607 (mf)

Crossopriza sengleti Huber, 2022 | | Iran urn:lsid:nmbe.ch:spidersp:053803

Crossopriza sengleti Huber, 2022: 189, f. 765-782 (Dmf) Original description

Crossopriza soudanensis Millot, 1941 | | Mali, Burkina Faso urn:lsid:nmbe.ch:spidersp:003393

Crossopriza soudanensis Millot, 1941: 28, f. 11A-L (Dmf) Original description

Crossopriza soudanensis Huber, 2022: 103, f. 354-372 (mf)

Crossopriza srinagar Huber, 2022 | | Pakistan, India urn:lsid:nmbe.ch:spidersp:053802

Crossopriza srinagar Huber, 2022: 185, f. 745-764 (Dmf) Original description

Crossopriza surobi Huber, 2022 | | Afghanistan urn:lsid:nmbe.ch:spidersp:053804

Crossopriza surobi Huber, 2022: 204, f. 806-822, 841-853 (Dmf) Original description

Crossopriza tiwi Huber, 2022 | | Oman urn:lsid:nmbe.ch:spidersp:053789

Crossopriza tiwi Huber, 2022: 118, f. 393-395, 433-449 (Dmf) Original description