World Spider Catalog

Pholcidae C. L. Koch, 1850

Family | ACCEPTED

N.B.: For comments on the monophyly and relationships of many genera see Huber, 1998d and Huber, 2000. The division of Pholcidae into five subfamilies, Ninetinae Simon, 1890, Arteminae Simon, 1893, Modisiminae Simon, 1893, Smeringopinae Simon, 1893 and Pholcinae, is confirmed by Huber, Eberle & Dimitrov, 2018.

In synonymy:

Coroia González-Sponga, 2005 = Artema Walckenaer, 1837 (Huber, Colmenares & Ramírez, 2014: 416)

Pholciella Roewer, 1960 = Artema Walckenaer, 1837 (Huber, 2009a: 67)

Gen. Artema Walckenaer, 1837 urn:lsid:nmbe.ch:spidergen:00370

N.B.: considered a senior synonym of Pholciella Roewer, 1960b: 42 [urn:lsid:nmbe.ch:spidergen:00409] (type P. ziaretana Roewer, 1960) by Huber, 2009a: 67 and of Coroia González-Sponga, 2005: 102 [urn:lsid:nmbe.ch:spidergen:04197] (type C. magna González-Sponga, 2005 = Artema atlanta Walckenaer, 1837) by Huber, Colmenares & Ramírez, 2014: 416.

In synonymy:

Artema magna (González-Sponga, 2005, T from Coroia = Artema) = Artema atlanta Walckenaer, 1837 (Huber, Colmenares & Ramírez, 2014: 417)

Artema mauriciana Walckenaer, 1837 = Artema atlanta Walckenaer, 1837 (Caporiacco, 1948a: 626, after F. O. Pickard-Cambridge, 1902a: 366)

Artema sexsignata (Franganillo, 1926, T from Crossopriza) = Artema atlanta Walckenaer, 1837 (Pérez, 1996: 432)

Artema sisyphoides (Doleschall, 1857) = Artema atlanta Walckenaer, 1837 (Millot, 1941: 3, sub A. mauriciana)

Artema atlanta Walckenaer, 1837 | | North Africa, Middle East. Introduced to both Americas, tropical Africa, Britain, Belgium, Greece, Turkey, tropical Asia, China, Japan, Taiwan, Australia, Pacific Is. urn:lsid:nmbe.ch:spidersp:003327

Artema atlanta Walckenaer, 1837: 656 (Dmf) Original description

Artema mauriciana Walckenaer, 1837: 657, pl. 15, f. 1 (Dmf) [urn:lsid:nmbe.ch:spidersp:056687]

Pholcus sisyphoides Doleschall, 1857: 408 (D) [urn:lsid:nmbe.ch:spidersp:056689]

Artema convexa Blackwall, 1858b: 332 (Dj)

Pholcus borbonicus Vinson, 1863: 132, pl. 3, f. 4 (Df)

Artema mauricia Vinson, 1863: 141, 307 (Dm)

Artema convexa Blackwall, 1866: 459 (Df)

Pholcus rotundatus Karsch, 1879h: 106 (Df)

Artema mauricia Simon, 1885h: 19

Artema sisyphoides Karsch, 1892: 276

Artema mauricia Simon, 1893a: 463, f. 441, 451-454 (mf)

Artema atlanta Pocock, 1900a: 239, f. 81 (Df)

Artema mauriciana Kulczyński, 1901a: 20, pl. 1, f. 15 (f)

Artema atlanta F. O. Pickard-Cambridge, 1902a: 366, pl. 34, f. 15-16 (mf)

Artema mauriciana Tullgren, 1910: 119, pl. 2, f. 35 (f)

Crossopriza sex-signata Franganillo, 1926a: 49 (Dmf) [urn:lsid:nmbe.ch:spidersp:056688]

Artema atlanta Petrunkevitch, 1929b: 119, f. 102-104 (mf)

Artema atlanta Dyal, 1935: 170, pl. 15, f. 97-103 (mf)

Artema mauriciana Millot, 1941: 3, f. 1F (m, S of Artema kochi and A. sisyphoides; S of A. kochi rejected by Aharon, Huber & Gavish-Regev, 2017: 26; other figs. see A. bunkpurugu)

Artema mauriciana Millot, 1946a: 129, f. 1 (f)

Artema atlanta Caporiacco, 1948a: 626 (S of Artema mauriciana)

Artema atlanta Chrysanthus, 1967b: 92, f. 15-19 (mf)

Artema atlanta Tikader & Biswas, 1981: 18, f. 12 (f)

Artema atlanta Brignoli, 1981c: 92, f. 1-7 (mf)

Artema atlanta Pérez, 1996: 432 (S of Crossopriza sexsignata)

Artema atlanta Huber, 2000: 12, f. 12-13, 48, 56-57, 99, 121, 145, 169, 195 (m)

Artema atlanta Van Keer & Van Keer, 2001: 83, f. 1.1-7 (mf)

Artema atlanta Saaristo, 2001b: 15, f. 16-22 (mf)

Artema atlanta Lee, 2005: 7, f. a-d (m)

Coroia magna González-Sponga, 2005: 102, pl. 2, f. 1-11 (Dmf) [urn:lsid:nmbe.ch:spidersp:047256]

Tibiosa coreana González-Sponga, 2006: 17, pl. 3, f. 8-9 (f) [misidentified per Huber, 2009a: 65]

Artema atlanta Beatty, Berry & Huber, 2008: 3, f. 15-16, 42-43 (mf)

Artema atlanta Colmenares-García, 2008: 87, f. 1A-C (m)

Artema atlanta Irie, 2009: 106, f. 1-2 (f)

Artema atlanta Saaristo, 2010: 159, f. 25.1-5 (mf)

Artema atlanta Gao & Li, 2010b: 11, f. 1A-D, 2A-D, 3A-C (mf)

Artema atlanta Le Peru, 2011: 152, f. 179 (mf)

Artema atlanta Tong, 2013: 55, f. 71A-D, 72A-D, 73A-C (mf)

Artema atlanta Huber, Colmenares & Ramírez, 2014: 417 (S of Coroia magna)

Artema atlanta Aharon, Huber & Gavish-Regev, 2017: 8, f. 3, 15-34, 201, 208 (mf)

Artema atlanta Al-Abbad, Al-Mayah & Al-Abody, 2019: 499, f. 2a-d, 3a-e, 4a-b, 5a-b (mf)

Artema bahla Huber, 2019 | | Oman urn:lsid:nmbe.ch:spidersp:050889

Artema bahla Huber in Huber & Carvalho, 2019: 6, f. 1-2, 7-8, 13-14, 19-20, 25, 30, 33-34 (Dmf) Original description

Artema bunkpurugu Huber & Kwapong, 2013 | | West Africa urn:lsid:nmbe.ch:spidersp:046602

Artema mauriciana Millot, 1941: 3, f. 1A-E, G-I (mf) [misidentified per Huber & Kwapong, 2013: 24]

Artema bunkpurugu Huber & Kwapong, 2013: 24, f. 14-15, 48-62 (Dmf) Original description

Artema bunkpurugu Aharon, Huber & Gavish-Regev, 2017: 50, f. 202, 209 (f)

Artema dhofar Huber, 2019 | | Oman urn:lsid:nmbe.ch:spidersp:050891

Artema dhofar Huber in Huber & Carvalho, 2019: 14, f. 5-6, 11-12, 17-18, 23-24, 27-28, 32, 37-38 (Dmf) Original description

Artema doriae (Thorell, 1881) | | Turkey, Israel, United Arab Emirates, Iraq, Iran, Afghanistan. Introduced to Japan urn:lsid:nmbe.ch:spidersp:003328

Pholcus doriae Thorell, 1881: 180 (Df, patronym for Doria, Latin genitive Doriae ICZN 31.1.1; Artema doriai is an unjustified emendation) Original description

Artema doriai Brignoli, 1981c: 92 (removed f from S of A. mauriciana, possible senior S of A. transcaspica and A. magna)

Artema doriai Tabrizi, Rad & Hedayati, 2014: 35, f. 1M-N, 3L-M (f)

Artema doriae Aharon, Huber & Gavish-Regev, 2017: 30, f. 90-120, 205, 212 (Dm, f)

Artema doriae Al-Khazali & Najim, 2018: 179, pl. 1A-D, 2A-D, 3 (mf)

Artema ghubrat Huber, 2019 | | Oman urn:lsid:nmbe.ch:spidersp:050890

Artema sp. a Aharon, Huber & Gavish-Regev, 2017: 44, f. 170 (f)

Artema ghubrat Huber in Huber & Carvalho, 2019: 11, f. 3-4, 9-10, 15-16, 21-22, 26, 29, 31, 35-36 (Dmf) Original description

Artema kochi Kulczyński, 1901 | | Eritrea, Ethiopia, Sudan, Yemen, Egypt? urn:lsid:nmbe.ch:spidersp:049789

Pholcus borbonicus L. Koch, 1875b: 25, pl. 3, f. 1 (f) [misidentified per Aharon, Huber & Gavish-Regev, 2017: 26]

Artema kochii Kulczyński, 1901a: 20, pl. 1, f. 14 (Df) Original description

Artema kochi Siyam, Dunlop & Jäger, 2017: 319, f. 1-3 (m)

Artema kochi Aharon, Huber & Gavish-Regev, 2017: 26, f. 76-89, 204, 211 (mf) [removed from S of A. atlanta, contra Simon, 1908a: 426 and Millot, 1941: 3, both sub A. mauriciana]

Artema magna Roewer, 1960 | | Afghanistan, Pakistan? urn:lsid:nmbe.ch:spidersp:003329

Artema magna Roewer, 1960b: 43, f. 14a-h (Df) Original description

Artema magna Aharon, Huber & Gavish-Regev, 2017: 41, f. 155-169 (Dm, f) [see also note at A. ziaretana]

Artema martensi Huber, 2021 | | Morocco urn:lsid:nmbe.ch:spidersp:053252

Artema martensi Huber, 2021: 326, f. 1-24 (Dmf) Original description

Artema nephilit Aharon, Huber & Gavish-Regev, 2017 | | Greece, Turkey, Cyprus, Israel, Jordan, Yemen?, United Arab Emirates? urn:lsid:nmbe.ch:spidersp:049788

Artema nephilit Aharon, Huber & Gavish-Regev, 2017: 15, f. 4-6, 9-14, 35-75, 203, 210 (Dmf) Original description

Artema transcaspica Spassky, 1934 | | Iran, Tajikistan, Turkmenistan, Uzbekistan urn:lsid:nmbe.ch:spidersp:003330

Artema transcaspica Spassky, 1934a: 369, f. 8-10 (Dmf) Original description

Artema transcaspica Aharon, Huber & Gavish-Regev, 2017: 35, f. 121-154, 206, 213 (mf)

Artema transcaspica Zamani et al., 2020: 585, f. 10A-B (m)

Artema ziaretana (Roewer, 1960) | j | Afghanistan urn:lsid:nmbe.ch:spidersp:003658

Pholciella ziaretana Roewer, 1960b: 42, f. 13a-d (Df) Original description

Artema ziaretana Huber, 2009a: 67 (T from Pholciella, type is j not f)

Artema ziaretana Aharon, Huber & Gavish-Regev, 2017: 44 (might be a synonym of A. magna)