Lycosidae Sundevall, 1833

Family |   ACCEPTED
N.B.: according to Brignoli, 1983c: 432, "Guy (1966, 1969) has proposed many synonymies and has downgraded many genera to subgenus rank; as this author based his conclusions more on nomenclature than on the study of material (or even of the original descriptions)", Platnick (WSC 15.0) did not accept Guy's conclusions; the changes by Guy, 1966 are noted (except where more recent work has superseded them); Guy, 1969 is merely a literature compilation that reflects those changes and is not cited separately; several genera of Lycosidae have been found not to be monophyletic (Murphy et al., 2006). Piacentini & Ramírez, 2019: 233-237 confirm ten valid subfamilies of Lycosidae: Venoniinae Lehtinen & Hippa, 1979, Zoicinae Lehtinen & Hippa, 1979 (= Piratinae Zyuzin, 1993), Evippinae Zyuzin, 1985, Sosippinae Dondale, 1986 (= Hygrolycosinae He & Song, 1996), Artoriinae Framenau, 2007, Tricassinae Alderweireldt & Jocqué, 1993 (= Arctosinae He & Song, 1996), Hippasinae Simon, 1898, Allocosinae Dondale, 1986, Pardosinae Simon, 1898 (= Wadicosinae Zyuzin, 1985) and Lycosinae Sundevall, 1833; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 518, f. 17-18.

In synonymy:

Artoriella Roewer, 1960 = Artoria Thorell, 1877 (Framenau, 2002: 210)

Lycosula Roewer, 1960 = Artoria Thorell, 1877 (Framenau, 2007a: 5)

Trabaeola Roewer, 1960 = Artoria Thorell, 1877 (Framenau, 2002: 210)

Gen. Artoria Thorell, 1877 urn:lsid:nmbe.ch:spidergen:01598
N.B.: considered a senior synonym of Artoriella Roewer, 1960d: 563 [urn:lsid:nmbe.ch:spidergen:05555] (type Artoria flavimana Simon, 1909) and of Trabaeola Roewer, 1960d: 582 [urn:lsid:nmbe.ch:spidergen:05556] (type Trabea lineata L. Koch, 1877) by Framenau, 2002: 210, and of Lycosula Roewer, 1960d: 889 [urn:lsid:nmbe.ch:spidergen:01631] (type Lycosella thorelli Berland, 1929) by Framenau, 2007a: 5; Guy, 1966: 63 considered Artoriella to be a subgenus of Artoria; see also note under Artoriellula.

Transferred to other genera:

Artoria fascicularis Purcell, 1903 -- see Proevippa

Artoria maura (Urquhart, 1892) -- see Anoteropsis

In synonymy:

Artoria ambrymiana (Berland, 1938, T from Lycosa) = Artoria berenice L. Koch, 1877 (Framenau, 2005: 272)

Artoria luwamata Barrion & Litsinger, 1995 = Artoria parvula Thorell, 1877 (Framenau, 2002: 223)

Artoria naeviella (Roewer, 1951, T from Hogna) = Artoria berenice L. Koch, 1877 (Framenau, 2005: 272)

Artoria neboissi (McKay, 1976, T from Lycosa) = Artoria flavimana Simon, 1909 (Framenau, 2002: 215)

Artoria versicolor (L. Koch, 1877) = Artoria berenice L. Koch, 1877 (Framenau, 2005: 272)

Nomina dubia:

Artoria australiensis (L. Koch, 1877a: 968, pl. 84, f. 6, j, Australia/New South Wales, originally in Trabea) -- Russell-Smith, 1982: 88, confirmed by Framenau, 2002: 211

Artoria pruinosa (L. Koch, 1877a: 925, pl. 80, f. 2, m, Australia/New South Wales, originally in Lycosa) -- Framenau & Baehr, 2018: 221

Artoria albopedipalpis Framenau, 2002 | | Australia (Victoria) urn:lsid:nmbe.ch:spidersp:037706
Artoria albopedipalpis Framenau, 2002: 211, f. 1A-F (Dmf) Original description
Artoria albopilata (Urquhart, 1893) | | Australia (Queensland to South Australia and Tasmania) urn:lsid:nmbe.ch:spidersp:018179
Lycosa albo-pilata Urquhart, 1893b: 123 (Dmf) Original description
Artoria albopilata Framenau, 2005: 266, f. 1A-E (mf, T from Lycosa)
Artoria albopilata Framenau & Baehr, 2018: 175, f. 3A-H, 46F (mf)
Artoria alta Framenau, 2004 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:038675
Artoria alta Framenau, 2004: 28, f. 1A-D (Dm) Original description
Artoria alta Framenau & Baehr, 2018: 175, f. 5A-H, 46F (m, Df [putative])
Artoria amoena (Roewer, 1960) | | Congo urn:lsid:nmbe.ch:spidersp:017704
Artoriella amoena Roewer, 1960d: 566, f. 329a-c (Dm) Original description
Artoria avona Framenau, 2002 | | Australia (South Australia, Victoria) urn:lsid:nmbe.ch:spidersp:037707
Artoria avona Framenau, 2002: 213, f. 2A-F (Dmf) Original description
Artoria barringtonensis Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050801
Artoria barringtonensis Framenau & Baehr, 2018: 177, f. 2B, 6A-D, 47A (Dm) Original description
Artoria beaury Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050802
Artoria beaury Framenau & Baehr, 2018: 179, f. 7A-H, 47B (Dmf) Original description
Artoria belfordensis Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050803
Artoria belfordensis Framenau & Baehr, 2018: 183, f. 2D, 9A-H, 47C (Dmf) Original description
Artoria berenice (L. Koch, 1877) | | Australia (Queensland to Tasmania), New Caledonia, Vanuatu urn:lsid:nmbe.ch:spidersp:019179
Lycosa berenice L. Koch, 1877a: 937, pl. 81, f. 3 (Df) Original description
Pardosa versicolor L. Koch, 1877a: 966, pl. 84, f. 4-5 (Dmf) [urn:lsid:nmbe.ch:spidersp:019001]
Lycosa naevia L. Koch, 1878a: 978, pl. 85, f. 5 (Df; preoccupied by L. Koch, 1875)
Lycosa ambrymiana Berland, 1938: 184, f. 153-156 (Dmf) [urn:lsid:nmbe.ch:spidersp:018182]
Tarentula naeviella Roewer, 1951: 442 (replacement name) [urn:lsid:nmbe.ch:spidersp:018063]
Avicosa berenice Roewer, 1955c: 236
Hogna naeviella Roewer, 1955c: 253
Artoria versicolor Framenau, 2002: 230, f. 27A-F (mf, T from Pardosa)
Artoria berenice Framenau, 2005: 272, f. 3 (f, T from Avicosa=Schizocosa, S of Artoria versicolor, Hogna naeviella and Lycosa ambrymiana)
Artoria berenice Framenau & Baehr, 2018: 183, f. 10C-D,G-H, 11A-H, 46H (mf)
Artoria bondi Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050804
Artoria bondi Framenau & Baehr, 2018: 188, f. 6E-H, 47D (Dm) Original description
Artoria booderee Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050805
Artoria booderee Framenau & Baehr, 2018: 190, f. 13A-H, 47E (Dmf) Original description
Artoria cingulipes Simon, 1909 | | Australia (Western Australia, South Australia) urn:lsid:nmbe.ch:spidersp:017705
Artoria cingulipes Simon, 1909d: 193, f. 10 (Df) Original description
Artoriella cingulipes Roewer, 1955c: 233
Artoria cingulipes Framenau, 2002: 214, f. 5A-B (f)
Artoria cingulipes Framenau, 2005: 273, f. 5A-E (f, Dm)
Artoria comleroi Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050806
Artoria comleroi Framenau & Baehr, 2018: 190, f. 16A-H, 47F (Dmf) Original description
Artoria corowa Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050807
Artoria corowa Framenau & Baehr, 2018: 192, f. 14A-H, 47G (Dmf) Original description
Artoria equipalus Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050808
Artoria equipalus Framenau & Baehr, 2018: 194, f. 17A-H, 47H (Dmf) Original description
Artoria extraordinaria Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050809
Artoria extraordinaria Framenau & Baehr, 2018: 196, f. 2E, 18A-H, 47I (Dmf) Original description
Artoria flavimana Simon, 1909 | | Australia (Western Australia to New South Wales, Victoria and Tasmania) urn:lsid:nmbe.ch:spidersp:017706
Artoria flavimanus Simon, 1909d: 192, f. 9 (Df) Original description
Artoriella flavimanus Roewer, 1955c: 233
Lycosa neboissi McKay, 1976a: 415, f. 1e-i (Dmf) [urn:lsid:nmbe.ch:spidersp:018333]
Lycosa naboissi Brignoli, 1983c: 450 (lapsus)
Artoria flavimanus Framenau, 2002: 215, f. 7A-F (mf, S of Lycosa neboissi)
Artoria flavimanus Vink, Mitchell & Paterson, 2002: 234, f. 8 (m)
Artoria flavimana Framenau & Baehr, 2018: 198, f. 1C, 19A-H, 46D (mf)
Artoria gloriosa (Rainbow, 1920) | | Australia (Lord Howe Is.) urn:lsid:nmbe.ch:spidersp:018258
Lycosa gloriosa Rainbow, 1920b: 261, pl. 30, f. 94-95 (Df) Original description
Lycosa gloriosa McKay, 1979c: 238, f. 1F-K (f, Dm)
Artoria gloriosa Framenau, 2005: 279, f. 7A-F (Tmf from Lycosa)
Artoria gloriosa Framenau & Baehr, 2018: 199, f. 20A-H, 46E (mf)
Artoria grahammilledgei Framenau & Baehr, 2018 | | Australia (Queensland, New South Wales) urn:lsid:nmbe.ch:spidersp:050810
Artoria grahammilledgei Framenau & Baehr, 2018: 201, f. 22A-H, 47J (Dmf) Original description
Artoria hamata Wang, Framenau & Zhang, 2021 | | China urn:lsid:nmbe.ch:spidersp:053064
Artoria hamata Wang, Framenau & Zhang, 2021: 577, f. 5A-E, 6A-J (Dmf) Original description
Artoria hebridisiana (Berland, 1938) | | Vanuatu urn:lsid:nmbe.ch:spidersp:018444
Lycosella hebridisiana Berland, 1938: 180, f. 143-144 (Dm) Original description
Lycosula hebridisiana Roewer, 1955c: 277
Artoria helensmithae Framenau & Baehr, 2018 | | Australia (New South Wales, Victoria) urn:lsid:nmbe.ch:spidersp:050811
Artoria helensmithae Framenau & Baehr, 2018: 205, f. 24A-H, 47K (Dmf) Original description
Artoria hospita Vink, 2002 | | New Zealand urn:lsid:nmbe.ch:spidersp:038676
Artoria hospita Vink, 2002: 35, f. 28, 62, 89 (Dmf) Original description
Artoria howquaensis Framenau, 2002 | | Australia (South Australia, New South Wales, Victoria) urn:lsid:nmbe.ch:spidersp:037708
Artoria howquaensis Framenau, 2002: 217, f. 9A-G (Dmf) Original description
Artoria howquaensis Framenau, Gotch & Austin, 2006: 28, f. 59-61 (mf)
Artoria howquaensis Framenau & Baehr, 2018: 208, f. 25A-H, 46J (mf)
Artoria impedita (Simon, 1909) | | Australia (Western Australia) urn:lsid:nmbe.ch:spidersp:019291
Lycosa impedita Simon, 1909d: 187, f. 6 (Df) Original description
Allocosa impedita Roewer, 1955c: 206
Trochosa expolita impedita McKay, 1979e: 288 (T from Allocosa, reduced to subspecies)
Artoria impedita Framenau, 2005: 283, f. 9A-F (mf, T from Trochosa, elevated to species)
Artoria kanangra Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050812
Artoria kanangra Framenau & Baehr, 2018: 208, f. 2C, 26A-D, 47L (Dm) Original description
Artoria kerewong Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050813
Artoria kerewong Framenau & Baehr, 2018: 209, f. 26E-H, 48A (Dm) Original description
Artoria ligulacea (Qu, Peng & Yin, 2009) | | China urn:lsid:nmbe.ch:spidersp:042846
Hygrolycosa ligulacea Qu, Peng & Yin, 2009: 71, f. 1a-c, 2a-d (Dmf) Original description
Artoria ligulacea Li, Framenau & Zhang, 2012: 40, f. 4A-I, 5A-E (Tmf from Hygrolycosa)
Artoria ligulacea Wang, Framenau & Zhang, 2021: 578, f. 7A-E, 8A-H (mf)
Artoria lineata (L. Koch, 1877) | | Australia (South Australia, New South Wales to Tasmania) urn:lsid:nmbe.ch:spidersp:037404
Trabea lineata L. Koch, 1877a: 970, pl. 84, f. 7 (Dm) Original description
Trabaeosa lineata Roewer, 1955c: 297
Trabaeola lineata Roewer, 1960d: 582
Trabaeola lineata Russell-Smith, 1982: 89 (cannot be identified and species inquirenda = nomen dubium)
Artoria lineata Framenau, 2002: 218, f. 11A-F (m, Df, T from Trabaeola, removed from nomen dubium)
Artoria lineata Framenau & Baehr, 2018: 211, f. 10A-B,E-F, 28A-H, 46A (mf)
Artoria linnaei Framenau, 2008 | | Australia (Western Australia) urn:lsid:nmbe.ch:spidersp:042066
Artoria linnaei Framenau, 2008c: 364, f. 1A-B, 2A-D (Dmf) Original description
Artoria lycosimorpha Strand, 1909 | j | South Africa urn:lsid:nmbe.ch:spidersp:017707
Artoria lycosimorpha Strand, 1909g: 589 (Dj) Original description
Artoriella lycosimorpha Roewer, 1955c: 233
Artoriella lycosimorpha Roewer, 1960d: 564, f. 327 (j)
Artoria maculatipes (Roewer, 1960) | | Namibia urn:lsid:nmbe.ch:spidersp:017708
Artroiella maculatipes Roewer, 1960d: 565, f. 328a-c (Dm, lapsus) Original description
Artoria maroota Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050814
Artoria maroota Framenau & Baehr, 2018: 214, f. 29A-D, 48B (Dm) Original description
Artoria mckayi Framenau, 2002 | | Australia (Queensland and South Australia to Tasmania) urn:lsid:nmbe.ch:spidersp:037709
Artoria mckayi Framenau, 2002: 220, f. 13A-F (Dmf) Original description
Artoria mckayi Framenau & Baehr, 2018: 214, f. 1A-B, 31A-H, 46G (mf)
Artoria minima (Berland, 1938) | | Vanuatu urn:lsid:nmbe.ch:spidersp:018445
Lycosella minima Berland, 1938: 182, f. 145-146 (Dm) Original description
Lycosula minima Roewer, 1955c: 277
Artoria mungo Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050815
Artoria mungo Framenau & Baehr, 2018: 216, f. 32A-H, 48C (Dmf) Original description
Artoria munmorah Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050816
Artoria munmorah Framenau & Baehr, 2018: 218, f. 33A-H, 48D (Dmf) Original description
Artoria myallensis Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050817
Artoria myallensis Framenau & Baehr, 2018: 219, f. 29E-H (Dm) Original description
Artoria palustris Dahl, 1908 | | New Guinea (Indonesia, Papua New Guinea/Bismarck Arch.) urn:lsid:nmbe.ch:spidersp:017702
Artoria palustris Dahl, 1908: 234, f. 16-17 (Dmf) Original description
Artoria parvula Dahl, 1908: 233, f. 14 (mf; misidentified per Framenau, 2002: 222)
Artoria parvula Chrysanthus, 1967a: 423, f. 68-69 (f; misidentified per Framenau, 2002: 222)
Artoria palustris Framenau, 2002: 222, f. 15A-F (mf)
Artoria parvula Thorell, 1877 | | China, Malaysia (peninsula, Borneo), Philippines, Indonesia (Sulawesi), Australia (Northern Territory) urn:lsid:nmbe.ch:spidersp:017703
Artoria parvula Thorell, 1877b: 531 (Dm) Original description
Artoria luwamata Barrion & Litsinger, 1995: 364, f. 213a-e (Dm) [urn:lsid:nmbe.ch:spidersp:017701]
Artoria parvula Framenau, 2002: 233, f. 17A-D (m, S of Artoria luwamata)
Artoria parvula Framenau, 2005: 286, f. 11A-B (m, Df)
Artoria parvula Li, Framenau & Zhang, 2012: 36, f. 1A-I, 2A-E, 3A-E (mf)
Artoria parvula Wang, Zhang & Peng, 2019: 394, f. 3A-H (mf)
Artoria parvula Wang, Framenau & Zhang, 2021: 579, f. 9A-E, 10A-H (mf)
Artoria quadrata Framenau, 2002 | | Australia (Queensland, New South Wales, Victoria) urn:lsid:nmbe.ch:spidersp:037710
Artoria quadrata Framenau, 2002: 224, f. 19A-F (Dmf) Original description
Artoria quadrata Framenau & Baehr, 2018: 221, f. 35A-H, 46C (mf)
Artoria schizocoides Framenau & Hebets, 2007 | | Australia (Western Australia) urn:lsid:nmbe.ch:spidersp:040063
Artoria schizocoides Framenau & Hebets, 2007: 96, f. 4-10 (Dmf) Original description
Artoria segrega Vink, 2002 | | New Zealand urn:lsid:nmbe.ch:spidersp:038677
Artoria segrega Vink, 2002: 35, f. 29, 63, 90 (Dmf) Original description
Artoria separata Vink, 2002 | | New Zealand urn:lsid:nmbe.ch:spidersp:038678
Artoria separata Vink, 2002: 36, f. 4, 30, 37, 64, 91 (Dmf) Original description
Artoria slatyeri Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050818
Artoria slatyeri Framenau & Baehr, 2018: 36, f. 36A-D, 48E (Dm) Original description
Artoria strepera Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050819
Artoria strepera Framenau & Baehr, 2018: 224, f. 2A, 36E-H, 48F (Dm) Original description
Artoria subhamata Wang & Zhang, 2022 | | China urn:lsid:nmbe.ch:spidersp:056657
Artoria subhamata Wang & Zhang, 2022c: 109, f. 1A-E, 2A-J (Dmf) Original description
Artoria taeniifera Simon, 1909 | | Australia (Western Australia, South Australia, New South Wales) urn:lsid:nmbe.ch:spidersp:017710
Artoria taeniifera Simon, 1909d: 193, f. 11 (Df) Original description
Artoriella taeniifera Roewer, 1955c: 233
Artoria taeniifera Framenau, 2002: 226, f. 21A-B (f)
Artoria taeniifera Framenau, 2005: 288, f. 13A-B (f)
Artoria taeniifera Framenau & Baehr, 2018: 225, f. 1D, 38A-H, 48G (Dm, f)
Artoria terania Framenau & Baehr, 2018 | | Australia (Queensland, New South Wales) urn:lsid:nmbe.ch:spidersp:050820
Artoria terania Framenau & Baehr, 2018: 227, f. 39A-H, 48H (Dmf) Original description
Artoria thorelli (Berland, 1929) | | Samoa, Marquesas Is. urn:lsid:nmbe.ch:spidersp:018446
Lycosella thorelli Berland, 1929b: 68, f. 65-68 (Df) Original description
Lycosella thorelli Berland, 1934b: 334, f. 18 (f)
Lycosella thorelli Marples, 1955a: 472, pl. 57, f. 4 (Dm)
Lycosula thorelli Roewer, 1955c: 277
Artoria triangularis Framenau, 2002 | | Australia (South Australia and Queensland to Tasmania) urn:lsid:nmbe.ch:spidersp:037711
Artoria triangularis Framenau, 2002: 227, f. 23A-E (Dmf) Original description
Artoria triangularis Framenau & Baehr, 2018: 230, f. 40A-H, 46K (mf)
Artoria ulrichi Framenau, 2002 | | Australia (New South Wales, Victoria) urn:lsid:nmbe.ch:spidersp:037712
Artoria ulrichi Framenau, 2002: 228, f. 25A-F (Dmf) Original description
Artoria ulrichi Framenau & Baehr, 2018: 230, f. 42A-H, 46B (mf)
Artoria victoriensis Framenau, Gotch & Austin, 2006 | | Australia (South Australia and Queensland to Tasmania) urn:lsid:nmbe.ch:spidersp:040064
Artoria victoriensis Framenau, Gotch & Austin, 2006: 28, f. 63-69 (Dmf) Original description
Artoria victoriensis Framenau & Baehr, 2018: 232, f. 43A-H, 46I (mf)
Artoria weiweii Wang, Zhang & Peng, 2019 | | Malaysia (Borneo) urn:lsid:nmbe.ch:spidersp:051380
Artoria weiweii Wang, Zhang & Peng, 2019: 392, f. 1A-D, 2A-G (Dmf) Original description
Artoria wilkiei Framenau & Baehr, 2018 | | Australia (New South Wales) urn:lsid:nmbe.ch:spidersp:050821
Artoria wilkiei Framenau & Baehr, 2018: 235, f. 45A-H, 48I (Dmf) Original description

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