Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.

In synonymy:

Paramyrmecion Bryant, 1940 = Litopyllus Chamberlin, 1922 (Platnick & Shadab, 1980a: 16)

Gen. Litopyllus Chamberlin, 1922 urn:lsid:nmbe.ch:spidergen:02444
N.B.: considered a senior synonym of Paramyrmecion Bryant, 1940: 394 [urn:lsid:nmbe.ch:spidergen:05696] (type P. cubanum Bryant, 1940) by Platnick & Shadab, 1980a: 16.

Transferred to other genera:

Litopyllus inconspicuus Bryant, 1940 -- see Cubanopyllus

Litopyllus paludis Chamberlin & Gertsch, 1940 -- see Synaphosus

In synonymy:

Litopyllus ambiguus Fox, 1938 = Litopyllus temporarius Chamberlin, 1922 (Platnick & Shadab, 1980a: 17)

Litopyllus barrowsi Roewer, 1951 = Litopyllus temporarius Chamberlin, 1922 (Platnick & Shadab, 1980a: 18)

Litopyllus liber Chamberlin & Gertsch, 1940 = Litopyllus temporarius Chamberlin, 1922 (Platnick & Shadab, 1980a: 17)

Litopyllus luteus (Barrows, 1919) = Litopyllus temporarius Chamberlin, 1922 (Platnick & Shadab, 1980a: 17)

Litopyllus rupicolens Chamberlin, 1922 = Litopyllus temporarius Chamberlin, 1922 (Ubick & Roth, 1973a: 5)

Litopyllus cubanus (Bryant, 1940) | | USA, Bahamas, Cuba urn:lsid:nmbe.ch:spidersp:027470
Paramyrmecion cubanum Bryant, 1940: 395, pl. 13, f. 173, 175 (Df) Original description
Litopyllus cubanus Platnick & Shadab, 1980a: 21, f. 36-39 (Dm, f, T from Paramyrmecion)
Litopyllus realisticus (Chamberlin, 1924) | | Mexico urn:lsid:nmbe.ch:spidersp:027471
Nodocion realisticus Chamberlin, 1924b: 613, f. 49 (Df) Original description
Litopyllus realisticus Platnick & Shadab, 1980a: 20, f. 34-35 (f, T from Nodocion)
Litopyllus temporarius Chamberlin, 1922 | | USA urn:lsid:nmbe.ch:spidersp:027472
Prosthesima lutea Barrows, 1919b: 356, pl. 15, f. 5 (Dm; preoccupied by F. O. Pickard-Cambridge, 1899, see Urozelotes rusticus) [urn:lsid:nmbe.ch:spidersp:058973]
Litopyllus temporarius Chamberlin, 1922: 155 (Dm) Original description
Litopyllus rupicolens Chamberlin, 1922: 155 (Df) [urn:lsid:nmbe.ch:spidersp:058972]
Litopyllus luteus Chamberlin, 1922: 155
Litopyllus ambiguus Fox, 1938a: 235, pl. 1, f. 4 (Dm) [urn:lsid:nmbe.ch:spidersp:058970]
Litopyllus liber Chamberlin & Gertsch, 1940: 2, f. 11-12 (Dmf) [urn:lsid:nmbe.ch:spidersp:058971]
Litopyllus rupicolens Muma, 1944: 10
Litopyllus rupicolens Kaston, 1948: 365, f. 1268-1272 (mf)
Litopyllus barrowsi Roewer, 1951: 443 (replacement name) [urn:lsid:nmbe.ch:spidersp:058974]
Litopyllus temporarius Ubick & Roth, 1973a: 5 (S of Litopyllus rupicolens)
Litopyllus temporarius Platnick & Shadab, 1980a: 17, f. 27-30 (mf, S of Litopyllus ambiguus, L. barrowsi, L. liber and L. luteus)
Litopyllus temporarius Heiss & Allen, 1986: 17, f. 30-31 (mf)
Litopyllus temporarius Platnick & Dondale, 1992: 225, f. 15, 338-341 (mf)
Litopyllus temporarius Murphy, 2007: 50, f. 360-361 (mf)
Litopyllus temporarius Azevedo, Griswold & Santos, 2018a: 679, f. 2B, 6B (mf)
Litopyllus temporarius Azevedo, Griswold & Santos, 2018b: 607, f. 7f,i, 21f, S6f (m)

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