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N.B.: family name and authorship as per ICZN, 2009 Opinion 2224, after application by Kluge, 2007; Arkyinae L. Koch, 1872 (genera Arkys Walckenaer, 1837 and Demadiana Strand, 1929) elevated to Arkyidae by Dimitrov et al., 2017: 244; monophyly of Araneidae stated by Kallal et al., 2018: 138; Nephilinae Simon, 1894 [urn:lsid:nmbe.ch:spiderfam:0111] elevated to family and relimited by Kuntner, 2006: 24, re-synonymized with Araneidae by Dimitrov et al., 2017: 244; Nephilinae and Phonognathinae Simon, 1894 (Simon, 1894a: 746 as Phonognatheae), the valid name for Zygiellinae Simon, 1929 sensu Wunderlich, 2004f, elevated to family by Kuntner et al., 2019: 563, not accepted by Scharff et al., 2020 and Kallal et al., 2020, and confirmed as family by Kuntner et al., 2023: 969-970 along with the separation of Paraplectanoididae Kuntner, Coddington, Agnarsson & Bond, 2023; Nephilinae and Phonognathinae downgraded to subfamily and Paraplectanoididae considered a synonym of Araneidae by Hormiga et al., 2023: 803; family status of Paraplectanoididae, Nephilidae and Phonognathidae revalidated by Kuntner et al., 2025: 17, 23, but not accepted here for reasons of stability; Oarcinae (Gnolus and Oarces) transferred from the Mimetidae to the Araneidae by Dimitrov et al., 2012: Suppl. 1, p. 15, re-transfer to the Mimetidae by Eskov & Marusik, 2023: 340 not accepted here (by majority vote of the WSC boards); Eskov & Marusik, 2023: 241 and Eskov & Marusik, 2024: 52 consider Zygiellidae as the valid name instead of Phonognathidae and accept the family status of Nephilidae; the status of the Nephilinae and Phonognathinae (= Zygiellidae) as subfamilies is accepted by majority vote of the WSC boards; Eskov & Marusik, 2024: 52 list ten subfamilies and tribes as valid (besides Nephilidae and Zygiellidae): Araneinae, Argiopinae Simon, 1890a: 81 (sub Argiopidae), Cyrtarachninae Simon, 1895a: 876 (sub Cyrtarachneae), Сyrtophorinae Simon, 1895a: 876 (sub Cyrtophoreae), Gasteracanthinae O. Pickard-Cambridge, 1871e: 217 (sub Gasteracanthides), Micratheninae Simon, 1895a: 848 (sub Micratheneae), Guizygiellinae Zhu, Song & Zhang, 2003: 42, Caerostrini Simon, 1895a: 831 (sub Cerostreae), Hypognathini Simon, 1895a: 871 (sub Hypognatheae) and Poltyini Simon, 1895a: 888 (sub Poltyeae).
Transferred to other genera:
Eustala aethiopica Roewer, 1961 -- see Araneus
Eustala cucurbitoria Yin, Wang, Xie & Peng, 1990 -- see Cyclosa
Eustala decemtuberculata Caporiacco, 1955 -- see Parawixia
Eustala hirsuta Mello-Leitão, 1942 -- see Ocrepeira
Eustala isosceles Mello-Leitão, 1939 -- see Kapogea
Eustala lactea Mello-Leitão, 1944 -- see Araneus
Eustala marginata Mello-Leitão, 1942 -- see Ocrepeira
Eustala seditiosa (Keyserling, 1893) -- see Metepeira
Eustala tuceps Chamberlin, 1925 -- see Metazygia
In synonymy:
Eustala abdita Chickering, 1955 = Eustala californiensis (Keyserling, 1885) (Levi, 1977a: 104)
Eustala anastera buliafera Chamberlin, 1924 = Eustala conchlea (McCook, 1888) (Levi, 1977a: 122)
Eustala anastera leuca Chamberlin, 1924 = Eustala conchlea (McCook, 1888) (Levi, 1977a: 122)
Eustala anastera vermiformis Franganillo, 1931 = Eustala fuscovittata (Keyserling, 1864) (Levi & Santos, 2013: 222)
Eustala apotroga (Walckenaer, 1841, T from Epeira) = Eustala anastera (Walckenaer, 1841) (Petrunkevitch, 1911: 340)
Eustala arkansana Archer, 1951 = Eustala emertoni (Banks, 1904) (Levi, 1977a: 120)
Eustala bombycinaria (Hentz, 1847) = Eustala anastera (Walckenaer, 1841) (Marx, 1890a: 544, sub Epeira eustala)
Eustala bucardia (McCook, 1894, T from Epeira) = Eustala californiensis (Keyserling, 1885) (Banks, 1901c: 189, sub Cyrtophora)
Eustala cambridgei (Keyserling, 1893, T from Epeira) = Eustala illicita (O. Pickard-Cambridge, 1889) (F. O. Pickard-Cambridge, 1904: 506)
Eustala caudata (Banks, 1898, T from Epeira) = Eustala fuscovittata (Keyserling, 1864) (F. O. Pickard-Cambridge, 1904: 505)
Eustala decolorata (Walckenaer, 1841, T from Epeira) = Eustala anastera (Walckenaer, 1841) (Petrunkevitch, 1911: 340)
Eustala diegensis (Schenkel, 1950, T from Araneus) = Eustala californiensis (Keyserling, 1885) (Levi, 1977a: 104)
Eustala eustala (Walckenaer, 1841, T from Epeira) = Eustala anastera (Walckenaer, 1841) (Petrunkevitch, 1911: 340)
Eustala gertschi Chickering, 1955 = Eustala conformans Chamberlin, 1925 (Levi & Santos, 2013: 222)
Eustala hebes (Hentz, 1847, T from Epeira) = Eustala anastera (Walckenaer, 1841) (Petrunkevitch, 1911: 340)
Eustala hirta (Taczanowski, 1873, T from Nephilengys) = Eustala fuscovittata (Keyserling, 1864) (Kuntner & Levi, 2007: 446)
Eustala illustrata (Walckenaer, 1841, T from Epeira) = Eustala anastera (Walckenaer, 1841) (Petrunkevitch, 1911: 340)
Eustala longembola Chickering, 1955 = Eustala ingenua Chickering, 1955 (Levi & Santos, 2013: 223)
Eustala mexicana Chickering, 1955 = Eustala californiensis (Keyserling, 1885) (Levi, 1977a: 104)
Eustala minima Chickering, 1955 = Eustala tantula Chickering, 1955 (Levi & Santos, 2013: 223)
Eustala montana Chickering, 1955 = Eustala montivaga Chickering, 1955 (Levi & Santos, 2013: 223)
Eustala nava (O. Pickard-Cambridge, 1899, T from Epeira) = Eustala scutigera (O. Pickard-Cambridge, 1898) (F. O. Pickard-Cambridge, 1904: 508)
Eustala nigromaculata (O. Pickard-Cambridge, 1895, T from Amamra) = Eustala clavispina (O. Pickard-Cambridge, 1889) (F. O. Pickard-Cambridge, 1904: 509)
Eustala orina (Chamberlin, 1916) = Eustala guianensis (Taczanowski, 1873) (Levi & Santos, 2013: 222)
Eustala panamana Chickering, 1955 = Eustala bucolica Chickering, 1955 (Levi & Santos, 2013: 222)
Eustala parvula (Keyserling, 1864) = Eustala cepina (Walckenaer, 1841) (Levi, 1977a: 118)
Eustala procurva Franganillo, 1936 = Eustala fuscovittata (Keyserling, 1864) (Levi & Santos, 2013: 222)
Eustala prompta (Hentz, 1847, T from Epeira) = Eustala anastera (Walckenaer, 1841) (F. O. Pickard-Cambridge, 1904: 506)
Eustala richardsi Mello-Leitão, 1939 = Eustala fuscovittata (Keyserling, 1864) (Levi & Santos, 2013: 222)
Eustala sanguinosa (Keyserling, 1893) = Eustala albiventer (Keyserling, 1884) (Poeta, Marques & Buckup, 2010a: 156)
Eustala spatulata (Walckenaer, 1841, T from Epeira) = Eustala anastera (Walckenaer, 1841) (Petrunkevitch, 1911: 340)
Eustala subfusca (Walckenaer, 1841, T from Epeira) = Eustala anastera (Walckenaer, 1841) (Petrunkevitch, 1911: 340)
Eustala thorelli (McCook, 1894, T from Cyclosa) = Eustala fuscovittata (Keyserling, 1864) (F. O. Pickard-Cambridge, 1904: 505)
Eustala triflex (Walckenaer, 1841, T from Epeira) = Eustala anastera (Walckenaer, 1841) (Petrunkevitch, 1911: 340)
Eustala trinotata (Walckenaer, 1841, T from Epeira) = Eustala anastera (Walckenaer, 1841) (Petrunkevitch, 1911: 340)
Eustala tumida Chickering, 1955 = Eustala nasuta Mello-Leitão, 1939 (Levi & Santos, 2013: 222)
Eustala venusta Chickering, 1955 = Eustala histrio Mello-Leitão, 1948 (Levi & Santos, 2013: 222)
Eustala vivida (Walckenaer, 1841, T from Epeira) = Eustala anastera (Walckenaer, 1841) (Petrunkevitch, 1911: 340)
Homonyms replaced:
Eustala mucronata (L. Koch, 1871) -- see Eustala mucronatella Roewer, 1942 Roewer, 1942a
Eustala viridipes (Taczanowski, 1878) -- see Eustala viridipedata Roewer, 1942 Roewer, 1942a
Nomina dubia:
Eustala albicans Caporiacco, 1954: 88, f. 16 (f, French Guiana) -- Poeta & Teixeira, 2017: 119
Eustala andina Chamberlin, 1916: 245 (j, Peru) -- Poeta & Teixeira, 2017: 116
Eustala essequibensis (Hingston, 1932: 366, m, Guyana, originally in Epeira) -- Poeta & Teixeira, 2017: 116
Eustala nigerrima Mello-Leitão, 1940d: 205 (f, Brazil) -- Poeta & Teixeira, 2017: 119
Eustala petasata (Walckenaer, 1841: 70, f, USA, originally in Epeira) -- Levi, 1977a: 120
Eustala tridentata (C. L. Koch, 1838: 41, f. 363-364, mf, French Guiana, Brazil, originally in Argyrodes) -- Poeta & Teixeira, 2017: 116
Eustala tristis (Blackwall, 1862b: 436, f, Brazil, originally in Epeira) -- Poeta & Teixeira, 2017: 116
Eustala uncicurva Franganillo, 1936c: 80 (f, Cuba) -- Poeta & Teixeira, 2017: 118
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