World Spider Catalog

Araneidae Clerck, 1757

Family | ACCEPTED

N.B.: family name and authorship as per ICZN, 2009 Opinion 2224, after application by Kluge, 2007; Arkyinae L. Koch, 1872 (genera Arkys Walckenaer, 1837 and Demadiana Strand, 1929) elevated to Arkyidae by Dimitrov et al., 2017: 244; monophyly of Araneidae stated by Kallal et al., 2018: 138; Nephilinae Simon, 1894 [urn:lsid:nmbe.ch:spiderfam:0111] elevated to family and relimited by Kuntner, 2006: 24, re-synonymized with Araneidae by Dimitrov et al., 2017: 244; Nephilinae and Phonognathinae Simon, 1894 (Simon, 1894a: 746 as Phonognatheae), the valid name for Zygiellinae Simon, 1929 sensu Wunderlich, 2004f, elevated to family by Kuntner et al., 2019: 563, not accepted by Scharff et al., 2020 and Kallal et al., 2020, and confirmed as family by Kuntner et al., 2023: 969-970 along with the separation of Paraplectanoididae Kuntner, Coddington, Agnarsson & Bond, 2023 (with the single genus Paraplectanoides Keyserling, 1886); Nephilinae and Phonognathinae downgraded to subfamily and Paraplectanoididae considered a synonym of Araneidae by Hormiga et al., 2023: 803; Oarcinae (Gnolus and Oarces) transferred from the Mimitidae to the Araneidae by Dimitrov et al., 2012: Suppl. 1, p. 15, re-transfer to the Mimetidae by Eskov & Marusik, 2023: 340 not accepted here (by majority vote of the WSC boards).

Gen. Eustala Simon, 1895 urn:lsid:nmbe.ch:spidergen:01480

Transferred to other genera:

Eustala aethiopica Roewer, 1961 -- see Araneus

Eustala cucurbitoria Yin, Wang, Xie & Peng, 1990 -- see Cyclosa

Eustala decemtuberculata Caporiacco, 1955 -- see Parawixia

Eustala hirsuta Mello-Leitão, 1942 -- see Ocrepeira

Eustala isosceles Mello-Leitão, 1939 -- see Kapogea

Eustala lactea Mello-Leitão, 1944 -- see Araneus

Eustala marginata Mello-Leitão, 1942 -- see Ocrepeira

Eustala seditiosa (Keyserling, 1893) -- see Metepeira

Eustala tuceps Chamberlin, 1925 -- see Metazygia

In synonymy:

Eustala abdita Chickering, 1955 = Eustala californiensis Keyserling, 1885 (Levi, 1977a: 104)

Eustala anastera buliafera Chamberlin, 1924 = Eustala conchlea McCook, 1888 (Levi, 1977a: 122)

Eustala anastera leuca Chamberlin, 1924 = Eustala conchlea McCook, 1888 (Levi, 1977a: 122)

Eustala anastera vermiformis Franganillo, 1931 = Eustala fuscovittata Keyserling, 1864 (Levi & Santos, 2013: 222)

Eustala apotroga (Walckenaer, 1841, T from Epeira) = Eustala anastera Walckenaer, 1841 (Petrunkevitch, 1911: 340)

Eustala arkansana Archer, 1951 = Eustala emertoni Banks, 1904 (Levi, 1977a: 120)

Eustala bombycinaria (Hentz, 1847) = Eustala anastera Walckenaer, 1841 (Marx, 1890a: 544, sub Epeira eustala)

Eustala bucardia (McCook, 1894, T from Epeira) = Eustala californiensis Keyserling, 1885 (Banks, 1901c: 189, sub Cyrtophora)

Eustala cambridgei (Keyserling, 1893, T from Epeira) = Eustala illicita O. Pickard-Cambridge, 1889 (F. O. Pickard-Cambridge, 1904: 506)

Eustala caudata (Banks, 1898, T from Epeira) = Eustala fuscovittata Keyserling, 1864 (F. O. Pickard-Cambridge, 1904: 505)

Eustala decolorata (Walckenaer, 1841, T from Epeira) = Eustala anastera Walckenaer, 1841 (Petrunkevitch, 1911: 340)

Eustala diegensis (Schenkel, 1950, T from Araneus) = Eustala californiensis Keyserling, 1885 (Levi, 1977a: 104)

Eustala eustala (Walckenaer, 1841, T from Epeira) = Eustala anastera Walckenaer, 1841 (Petrunkevitch, 1911: 340)

Eustala gertschi Chickering, 1955 = Eustala conformans Chamberlin, 1925 (Levi & Santos, 2013: 222)

Eustala hebes (Hentz, 1847, T from Epeira) = Eustala anastera Walckenaer, 1841 (Petrunkevitch, 1911: 340)

Eustala hirta (Taczanowski, 1873, T from Nephilengys) = Eustala fuscovittata Keyserling, 1864 (Kuntner & Levi, 2007: 446)

Eustala illustrata (Walckenaer, 1841, T from Epeira) = Eustala anastera Walckenaer, 1841 (Petrunkevitch, 1911: 340)

Eustala longembola Chickering, 1955 = Eustala ingenua Chickering, 1955 (Levi & Santos, 2013: 223)

Eustala mexicana Chickering, 1955 = Eustala californiensis Keyserling, 1885 (Levi, 1977a: 104)

Eustala minima Chickering, 1955 = Eustala tantula Chickering, 1955 (Levi & Santos, 2013: 223)

Eustala montana Chickering, 1955 = Eustala montivaga Chickering, 1955 (Levi & Santos, 2013: 223)

Eustala nava (O. Pickard-Cambridge, 1899, T from Epeira) = Eustala scutigera O. Pickard-Cambridge, 1898 (F. O. Pickard-Cambridge, 1904: 508)

Eustala nigromaculata (O. Pickard-Cambridge, 1895, T from Amamra) = Eustala clavispina O. Pickard-Cambridge, 1889 (F. O. Pickard-Cambridge, 1904: 509)

Eustala orina (Chamberlin, 1916) = Eustala guianensis Taczanowski, 1873 (Levi & Santos, 2013: 222)

Eustala panamana Chickering, 1955 = Eustala bucolica Chickering, 1955 (Levi & Santos, 2013: 222)

Eustala parvula (Keyserling, 1864) = Eustala cepina Walckenaer, 1841 (Levi, 1977a: 118)

Eustala procurva Franganillo, 1936 = Eustala fuscovittata Keyserling, 1864 (Levi & Santos, 2013: 222)

Eustala prompta (Hentz, 1847, T from Epeira) = Eustala anastera Walckenaer, 1841 (F. O. Pickard-Cambridge, 1904: 506)

Eustala richardsi Mello-Leitão, 1939 = Eustala fuscovittata Keyserling, 1864 (Levi & Santos, 2013: 222)

Eustala sanguinosa (Keyserling, 1893) = Eustala albiventer Keyserling, 1884 (Poeta, Marques & Buckup, 2010a: 156)

Eustala spatulata (Walckenaer, 1841, T from Epeira) = Eustala anastera Walckenaer, 1841 (Petrunkevitch, 1911: 340)

Eustala subfusca (Walckenaer, 1841, T from Epeira) = Eustala anastera Walckenaer, 1841 (Petrunkevitch, 1911: 340)

Eustala thorelli (McCook, 1894, T from Cyclosa) = Eustala fuscovittata Keyserling, 1864 (F. O. Pickard-Cambridge, 1904: 505)

Eustala triflex (Walckenaer, 1841, T from Epeira) = Eustala anastera Walckenaer, 1841 (Petrunkevitch, 1911: 340)

Eustala trinotata (Walckenaer, 1841, T from Epeira) = Eustala anastera Walckenaer, 1841 (Petrunkevitch, 1911: 340)

Eustala tumida Chickering, 1955 = Eustala nasuta Mello-Leitão, 1939 (Levi & Santos, 2013: 222)

Eustala venusta Chickering, 1955 = Eustala histrio Mello-Leitão, 1948 (Levi & Santos, 2013: 222)

Eustala vivida (Walckenaer, 1841, T from Epeira) = Eustala anastera Walckenaer, 1841 (Petrunkevitch, 1911: 340)

Homonyms replaced:

Eustala mucronata (L. Koch, 1871) -- see Eustala mucronatella Roewer, 1942 Roewer, 1942a

Eustala viridipes (Taczanowski, 1878) -- see Eustala viridipedata Roewer, 1942 Roewer, 1942a

Nomina dubia:

Eustala albicans Caporiacco, 1954: 88, f. 16 (f, French Guiana) -- Poeta & Teixeira, 2017: 119

Eustala andina Chamberlin, 1916: 245 (j, Peru) -- Poeta & Teixeira, 2017: 116

Eustala essequibensis (Hingston, 1932: 366, m, Guyana, originally in Epeira) -- Poeta & Teixeira, 2017: 116

Eustala nigerrima Mello-Leitão, 1940d: 205 (f, Brazil) -- Poeta & Teixeira, 2017: 119

Eustala petasata (Walckenaer, 1841: 70, f, USA, originally in Epeira) -- Levi, 1977a: 120

Eustala tridentata (C. L. Koch, 1838: 41, f. 363-364, mf, French Guiana, Brazil, originally in Argyrodes) -- Poeta & Teixeira, 2017: 116

Eustala tristis (Blackwall, 1862b: 436, f, Brazil, originally in Epeira) -- Poeta & Teixeira, 2017: 116

Eustala uncicurva Franganillo, 1936c: 80 (f, Cuba) -- Poeta & Teixeira, 2017: 118

Eustala albiventer (Keyserling, 1884) | | Brazil urn:lsid:nmbe.ch:spidersp:015910

Epeira albiventer Keyserling, 1884b: 651, pl. 21, f. 3 (Df) Original description

Epeira albiventer Keyserling, 1892: 157, pl. 8, f. 115 (f)

Epeira sanguinosa Keyserling, 1893: 225, pl. 11, f. 167 (Df) [urn:lsid:nmbe.ch:spidersp:015977]

Araneus albiventer Petrunkevitch, 1911: 278

Araneus sanguinosus Petrunkevitch, 1911: 314

Eustala sanguinosa Roewer, 1942a: 766

Metazygia albiventer Mello-Leitão, 1943a: 186 (T from Araneus)

Eustala albiventer Levi, 1991a: 178 (T from Metazygia)

Eustala albiventer Poeta, Marques & Buckup, 2010a: 155, f. 20-24 (Dm, f, S of Eustala sanguinosa)

Eustala anastera (Walckenaer, 1841) | | North, Central America urn:lsid:nmbe.ch:spidersp:015911

Epeira anastera Walckenaer, 1841: 33 (Df) Original description

Epeira eustala Walckenaer, 1841: 37 (Df) [urn:lsid:nmbe.ch:spidersp:061418]

Epeira apotroga Walckenaer, 1841: 43 (Df) [urn:lsid:nmbe.ch:spidersp:061419]

Epeira spatulata Walckenaer, 1841: 44 (Df) [urn:lsid:nmbe.ch:spidersp:061420]

Epeira illustrata Walckenaer, 1841: 45 (Dm) [urn:lsid:nmbe.ch:spidersp:061421]

Epeira decolorata Walckenaer, 1841: 49 (Dmf) [urn:lsid:nmbe.ch:spidersp:061422]

Epeira vivida Walckenaer, 1841: 54 (Df) [urn:lsid:nmbe.ch:spidersp:061423]

Epeira triflex Walckenaer, 1841: 60 (Dm) [urn:lsid:nmbe.ch:spidersp:061424]

Epeira trinotata Walckenaer, 1841: 75 (Df) [urn:lsid:nmbe.ch:spidersp:061425]

Epeira subfusca Walckenaer, 1841: 76 (Df) [urn:lsid:nmbe.ch:spidersp:061426]

Epeira prompta Hentz, 1847: 473, pl. 31, f. 4 (Df) [urn:lsid:nmbe.ch:spidersp:061427]

Epeira hebes Hentz, 1847: 473, pl. 31, f. 7 (Df) [urn:lsid:nmbe.ch:spidersp:061428]

Epeira bombycinaria Hentz, 1847: 476, pl. 31, f. 16 (Df) [urn:lsid:nmbe.ch:spidersp:061429]

Epeira eustala Marx, 1890a: 544 (S of Epeira bombycinaria and E. parvula; S of E. parvula rejected by Levi, 1977a: 118, see Eustala cepina)

Epeira bombycinaria Keyserling, 1892: 145, pl. 7, f. 107 (mf)

Epeira anastera McCook, 1894: 172, pl. 8, f. 1-4 (mf, S of Eustala cepina; revalidated by Chamberlin & Ivie, 1944: 103)

Eustala anastera Simon, 1895a: 795, f. 860 (f)

Eustala anastera F. O. Pickard-Cambridge, 1904: 506, pl. 48, f. 7-8 (mf, S of Epeira prompta)

Eustala anastera Petrunkevitch, 1911: 340 (S of Eperia apotroga, E. decolorata, E. eustala, E. hebes, E. illustrata, E. spatulata, E. subfusca, E. triflex, E. trinotata and E. vivida)

Eustala anastera Petrunkevitch, 1930a: 331, f. 215 (f)

Eustala anastera Comstock, 1940: 524, f. 549-554 (f)

Eustala anastera Muma, 1943: 92, pl. XIII, f. 18 (f)

Eustala anastera Chamberlin & Ivie, 1944: 102 (S of Eustala conchlea; revalidated by Levi, 1977a: 122)

Eustala triflex Chamberlin & Ivie, 1944: 103 (removed from S of Eustala anastera, contra Petrunkevitch, 1911: 340; S confirmed by Levi, 1977a: 114)

Eustala anastera Kaston, 1948: 233, f. 706-709, 727 (f)

Eustala anastera Archer, 1951a: 19, f. 46 (m)

Eustala anastera Chickering, 1955: 414, f. 24-29 (mf, S of Eustala anastera buliafera and E. a. leuca, rejected by Levi, 1977a: 122, see Eustala conchlea; S of Eustala prompta confirmed)

Eustala anastera Levi, 1977a: 114, f. 205-232, 280-285, 298-302, 314-315 (mf, S of Eustala triflex confirmed, contra Chamberlin & Ivie, 1944: 103)

Eustala anastera Coddington, 1990: 14, f. 60 (m)

Eustala anastera Breene et al., 1993: 104, f. 155A-C (mf)

Eustala anastera Levi, 2002: 533, f. 13-15, 208-210 (mf)

Eustala anastera Paquin & Dupérré, 2003: 46, f. 308-310 (mf)

Eustala anastera Dondale et al., 2003: 267, f. 600-611 (mf)

Eustala bacelarae Caporiacco, 1955 | | Venezuela urn:lsid:nmbe.ch:spidersp:015914

Eustala bacelarae Caporiacco, 1955: 340, f. 27a-d (Dmf) Original description

Eustala banksi Chickering, 1955 | | Mexico, Costa Rica urn:lsid:nmbe.ch:spidersp:015915

Eustala banksi Chickering, 1955: 417, f. 30-34 (Dm) Original description

Eustala belissima Poeta, Marques & Buckup, 2010 | | Brazil, Uruguay urn:lsid:nmbe.ch:spidersp:044011

Eustala belissima Poeta, Marques & Buckup, 2010b: 267, f. 4-9 (Dmf) Original description

Eustala bifida F. O. Pickard-Cambridge, 1904 | | USA to Panama urn:lsid:nmbe.ch:spidersp:015916

Eustala bifida F. O. Pickard-Cambridge, 1904: 507, pl. 48, f. 9-10 (Dmf) Original description

Epeira bifida Banks, 1909a: 212

Eustala bifida Chickering, 1955: 421, f. 35-40 (mf)

Eustala bifida Levi, 1977a: 108, f. 167-175, 178 (mf)

Eustala bisetosa Bryant, 1945 | | Hispaniola urn:lsid:nmbe.ch:spidersp:015917

Eustala bisetosa Bryant, 1945a: 370, f. 5-6 (Dmf) Original description

Eustala brevispina Gertsch & Davis, 1936 | | USA, Mexico urn:lsid:nmbe.ch:spidersp:015918

Eustala brevispina Gertsch & Davis, 1936: 12, f. 9-10 (Dmf) Original description

Eustala brevispina Levi, 1977a: 106, f. 149-158 (mf)

Eustala bucolica Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015919

Eustala bucolica Chickering, 1955: 425, f. 41-44 (Dm) Original description

Eustala panamana Chickering, 1955: 480, f. 103-105 (Df) [urn:lsid:nmbe.ch:spidersp:015965]

Eustala bucolica Levi & Santos, 2013: 222 (S of Eustala panamana)

Eustala californiensis (Keyserling, 1885) | | USA, Mexico urn:lsid:nmbe.ch:spidersp:015920

Cyrtophora californiensis Keyserling, 1885: 525, pl. 13, f. 24 (Df) Original description

Cyrtophora californiensis Keyserling, 1893: 263, pl. 13, f. 106 (f)

Epeira bucardia McCook, 1894: 183, pl. 9, f. 4 (Df) [urn:lsid:nmbe.ch:spidersp:061410]

Cyrtophora californiensis Banks, 1901c: 189 (S of Epeira bucardia)

Araneus diegensis Schenkel, 1950b: 67, f. 23a-c (Df) [urn:lsid:nmbe.ch:spidersp:061406]

Eustala abdita Chickering, 1955: 410, f. 19-23 (Dm) [urn:lsid:nmbe.ch:spidersp:061402]

Eustala mexicana Chickering, 1955: 465, f. 88-89 (Df) [urn:lsid:nmbe.ch:spidersp:061407]

Eustala californiensis Levi, 1977a: 104, f. 138-148 (mf, T from Cyrtophora, S of Araneus diegensis, Eustala abdita and E. mexicana)

Eustala cameronensis Gertsch & Davis, 1936 | | USA urn:lsid:nmbe.ch:spidersp:015921

Eustala cameronensis Gertsch & Davis, 1936: 13, f. 13 (Dm) Original description

Eustala cameronensis Levi, 1977a: 112, f. 189-191 (m)

Eustala catarina Poeta, 2014 | | Brazil urn:lsid:nmbe.ch:spidersp:047223

Eustala catarina Poeta, 2014: 443, f. 10-18 (Dmf) Original description

Eustala cazieri Levi, 1977 | | USA, Bahamas, Turks & Caicos urn:lsid:nmbe.ch:spidersp:015922

Eustala cazieri Levi, 1977a: 102, f. 128-137 (Dmf) Original description

Eustala cepina (Walckenaer, 1841) | | Canada, USA, Mexico urn:lsid:nmbe.ch:spidersp:015923

Epeira cepina Walckenaer, 1841: 37 (Df) Original description

Epeira parvula Keyserling, 1864: 131, pl. 6, f. 9-10 (Df) [urn:lsid:nmbe.ch:spidersp:061409]

Epeira parvula Emerton, 1884: 317, pl. 34, f. 12, pl. 37, f. 1, 2 (f, Dm)

Epeira parvula Emerton, 1902: 179, f. 416-418 (mf)

Eustala cepina Chamberlin & Ivie, 1944: 103 (removed from S of Eustala anastera, rejecting McCook, 1894: 172)

Eustala arkansana Archer, 1951a: 19, f. 47 (f; misidentified per Levi, 1977a: 118)

Eustala cepina Levi, 1977a: 118, f. 233-252, 286-290, 303-308, 316 (Dm, f, S of Eustala parvula, E. parvula removed from S of E. anastera, rejecting Marx, 1890a: 544)

Eustala cepina Breene et al., 1993: 104, f. 156A-C (mf)

Eustala cepina Dondale et al., 2003: 271, f. 622-631 (mf)

Eustala cepina Paquin et al., 2008c: 29, f. 6-8 (mf)

Eustala cidae Poeta, 2014 | | Brazil urn:lsid:nmbe.ch:spidersp:047225

Eustala cidae Poeta, 2014: 446, f. 29-37 (Dmf) Original description

Eustala clavispina (O. Pickard-Cambridge, 1889) | | USA to El Salvador urn:lsid:nmbe.ch:spidersp:015924

Epeira clavispina O. Pickard-Cambridge, 1889d: 37, pl. 7, f. 11 (Df) Original description

Epeira clavispina Keyserling, 1892: 102, pl. 5, f. 75 (Dm)

Amamra nigromaculata O. Pickard-Cambridge, 1895a: 155, pl. 19, f. 5 (Df) [urn:lsid:nmbe.ch:spidersp:061411]

Eustala clavispina F. O. Pickard-Cambridge, 1904: 509, pl. 48, f. 19 (f, S of Amamra nigromaculata)

Eustala rosae Gertsch & Davis, 1936: 14, f. 12 (m; misidentified per Levi, 1977a: 112)

Eustala clavispina Chickering, 1955: 428, f. 45-48 (f)

Eustala clavispina Kraus, 1955b: 21, f. 51-52 (f)

Eustala clavispina Levi, 1977a: 106, f. 159-166, 176-177 (f, Dm)

Eustala conchlea (McCook, 1888) | | USA, Mexico urn:lsid:nmbe.ch:spidersp:015925

Epeira parvula conchlea McCook, 1888b: 199, f. 6 (Df) Original description

Epeira anastera conchlea McCook, 1894: 174, pl. 8, f. 1n (f)

Eustala anastera buliafera Chamberlin, 1924b: 650 (Df) [urn:lsid:nmbe.ch:spidersp:061403]

Eustala anastera leuca Chamberlin, 1924b: 650 (Df) [urn:lsid:nmbe.ch:spidersp:061404]

Eustala conchlea Levi, 1977a: 122, f. 269-279, 296, 312, 318 (mf, elevated from subspecies, removed from S of Eustala anastera contra Chamberlin & Ivie, 1944: 102; S of Eustala anastera buliafera and E. a. leuca, both removed from S of Eustala anastera contra Chickering, 1955: 414)

Eustala conformans Chamberlin, 1925 | | Panama, Colombia urn:lsid:nmbe.ch:spidersp:015926

Eustala conformans Chamberlin, 1925d: 216 (Df) Original description

Eustala conformans Chickering, 1955: 430, f. 49-50 (f)

Eustala gertschi Chickering, 1955: 440, f. 59-62 (Dm) [urn:lsid:nmbe.ch:spidersp:015937]

Eustala conformans Levi & Santos, 2013: 222 (S of Eustala gertschi)

Eustala crista Poeta, Marques & Buckup, 2010 | | Brazil urn:lsid:nmbe.ch:spidersp:044012

Eustala crista Poeta, Marques & Buckup, 2010b: 268, f. 10-15 (Dmf) Original description

Eustala cuia Poeta, 2014 | | Brazil urn:lsid:nmbe.ch:spidersp:047227

Eustala cuia Poeta, 2014: 450, f. 47-55 (Dmf) Original description

Eustala davalosae Pett & Pai-Gibson, 2024 | | Paraguay urn:lsid:nmbe.ch:spidersp:060052

Eustala davalosae Pett & Pai-Gibson, 2024: 998, f. 1-6 (Df) Original description

Eustala delasmata Bryant, 1945 | | Mexico, Dominican Rep. urn:lsid:nmbe.ch:spidersp:015928

Eustala delasmata Bryant, 1945a: 372, f. 11 (Dm) Original description

Eustala delasmata Poeta & Teixeira, 2017: 115, f. 10-16 (m, Df)

Eustala delecta Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015929

Eustala delecta Chickering, 1955: 431, f. 51-54 (Dm) Original description

Eustala devia (Gertsch & Mulaik, 1936) | | USA to Panama, Caribbean urn:lsid:nmbe.ch:spidersp:015930

Neosconella devia Gertsch & Mulaik, 1936b: 16, f. 38 (Df) Original description

Eustala devia Levi, 1973: 487 (T from Araneus)

Eustala devia Levi, 1977a: 101, f. 118-127 (f, Dm, S of Eustala minima, rejected by Levi & Santos, 2013: 223, see Eustala tantula)

Eustala eldorado Poeta, 2014 | | Brazil urn:lsid:nmbe.ch:spidersp:047229

Eustala eldorado Poeta, 2014: 452, f. 65-68 (Dm) Original description

Eustala eleuthera Levi, 1977 | | USA, Bahama Is., Jamaica urn:lsid:nmbe.ch:spidersp:015931

Eustala eleuthera Levi, 1977a: 111, f. 179-188 (Dmf) Original description

Eustala emertoni (Banks, 1904) | | USA, Mexico urn:lsid:nmbe.ch:spidersp:015932

Epeira emertoni Banks, 1904a: 111 (Df; preoccupied by Scudder, 1890 by a fossil species [now in Araneus]; the two species are not currently considered congeneric, but Banks' name is a primary homonym) Original description

Araneus emertoni Petrunkevitch, 1911: 291

Eustala anastera emertoni Archer, 1940: 55 (T from Araneus, placed as subspecies)

Eustala arkansana Archer, 1951a: 19, f. 44 (Dm; f f. 47 see Eustala cepina) [urn:lsid:nmbe.ch:spidersp:061405]

Eustala triflex Archer, 1951a: 19, f. 45 (m; misidentified per Levi, 1977a: 120)

Eustala emertoni Levi, 1977a: 120, f. 253-268, 291-295, 309-311, 317 (mf, elevated from subspecies of Eustala anastera, S of Eustala arkansana)

Eustala emertoni Dondale et al., 2003: 269, f. 612-621 (mf)

Eustala emertoni Jiménez et al., 2018: 4, f. 4-7 (f)

Eustala ericae Poeta, 2014 | | Brazil urn:lsid:nmbe.ch:spidersp:047224

Eustala ericae Poeta, 2014: 445, f. 19-28 (Dmf) Original description

Eustala exigua Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015934

Eustala exigua Chickering, 1955: 435, f. 55-58 (Dm) Original description

Eustala farroupilha Poeta, 2014 | | Brazil urn:lsid:nmbe.ch:spidersp:047228

Eustala farroupilha Poeta, 2014: 451, f. 56-64 (Dmf) Original description

Eustala fragilis (O. Pickard-Cambridge, 1889) | | Guatemala, Panama urn:lsid:nmbe.ch:spidersp:015935

Epeira fragilis O. Pickard-Cambridge, 1889d: 23, pl. 4, f. 18 (Df) Original description

Epeira fragilis Keyserling, 1892: 186, pl. 9, f. 137 (f)

Eustala fragilis F. O. Pickard-Cambridge, 1904: 509, pl. 48, f. 18 (f)

Eustala fuscovittata (Keyserling, 1864) | | USA, Cuba, Mexico, Guatemala, Honduras, Costa Rica, Nicaragua, Panama, Colombia, Guyana urn:lsid:nmbe.ch:spidersp:015936

Epeira fusco-vittata Keyserling, 1864: 129, pl. 6, f. 7-8 (Df) Original description

Nephila hirta Taczanowski, 1873: 149 (Df) [urn:lsid:nmbe.ch:spidersp:013991]

Epeira fuscovittata Keyserling, 1893: 251, pl. 13, f. 187 (f, Dm)

Cyclosa thorelli McCook, 1894: 228, pl. 19, f. 11 (Df) [urn:lsid:nmbe.ch:spidersp:061412]

Epeira caudata Banks, 1898b: 255, pl. 15, f. 5 (Df; preoccupied by Hentz, 1850, see Cyclosa turbinata) [urn:lsid:nmbe.ch:spidersp:061413]

Eustala fuscovittata F. O. Pickard-Cambridge, 1904: 505, pl. 48, f. 3-4 (mf, S of Cyclosa thorelli and Epeira caudata Banks)

Eustala anastera vermiformis Franganillo, 1931d: 44 (Df) [urn:lsid:nmbe.ch:spidersp:015912]

Eustala procurva Franganillo, 1936c: 80, f. 37a-b (Df) [urn:lsid:nmbe.ch:spidersp:015969]

Eustala richardsi Mello-Leitão, 1939g: 105, f. 1-4 (Dmf) [urn:lsid:nmbe.ch:spidersp:015971]

Nephilengys hirta Roewer, 1942a: 934

Eustala fusco-vittata Chickering, 1955: 398, f. A-B, 1-5 (mf)

Eustala fuscovittata Grasshoff, 1970b: 216, f. 2a-c (mf)

Eustala fuscovittata Kuntner & Levi, 2007: 446, f. 1-3 (f, S of Nephilengys hirta)

Eustala fuscovittata Levi & Santos, 2013: 222 (S of Eustala anastera vermiformis, E. procurva and E. richardsi)

Eustala fuscovittata Galvis, 2020c: 263, f. 1-11 (mf)

Eustala gonygaster (C. L. Koch, 1838) | | Brazil, Guyana urn:lsid:nmbe.ch:spidersp:015938

Argyopes gonygaster C. L. Koch, 1838: 43, f. 365 (Df) Original description

Eustala gonygaster Simon, 1895a: 789

Eustala gonygaster Caporiacco, 1948a: 660, f. 68-69 (Dm)

Eustala guarani Poeta, 2014 | | Brazil urn:lsid:nmbe.ch:spidersp:047222

Eustala guarani Poeta, 2014: 442, f. 1-9 (Dmf) Original description

Eustala guianensis (Taczanowski, 1873) | | Peru, French Guiana urn:lsid:nmbe.ch:spidersp:015939

Singa guianensis Taczanowski, 1873: 124 (Df) Original description

Epeira guianensis Taczanowski, 1878b: 160, pl. 2, f. 11 (f, Dm)

Eustala guianensis Simon, 1895a: 789

Aranea orina Chamberlin, 1916: 248, pl. 19, f. 3 (Dm) [urn:lsid:nmbe.ch:spidersp:015963]

Eustala orina Levi, 1991a: 177 (T from Araneus)

Eustala guianensis Levi & Santos, 2013: 222 (S of Eustala orina)

Eustala guttata F. O. Pickard-Cambridge, 1904 | | Mexico to Brazil urn:lsid:nmbe.ch:spidersp:015940

Eustala guttata F. O. Pickard-Cambridge, 1904: 508, pl. 48, f. 13-14 (Dmf) Original description

Eustala guttata Camargo, 1953: 309, f. 6, 8, 31-34, 48 (m)

Eustala guttata Chickering, 1955: 442, f. 63-68 (mf)

Eustala guttata Kraus, 1955b: 22, f. 53 (f)

Eustala histrio Mello-Leitão, 1948 | | Panama, Guyana urn:lsid:nmbe.ch:spidersp:015941

Eustala histrio Mello-Leitão, 1948: 165, f. 8 (Df) Original description

Eustala venusta Chickering, 1955: 515, f. 139-140 (Df) [urn:lsid:nmbe.ch:spidersp:015996]

Eustala histrio Levi & Santos, 2013: 222 (S of Eustala venusta)

Eustala illicita (O. Pickard-Cambridge, 1889) | | Mexico to Brazil urn:lsid:nmbe.ch:spidersp:015942

Epeira illicita O. Pickard-Cambridge, 1889d: 17, pl. 6, f. 16 (Df) Original description

Epeira cambridgii Keyserling, 1893: 255, pl. 13, f. 190 (Df) [urn:lsid:nmbe.ch:spidersp:061414]

Eustala illicita F. O. Pickard-Cambridge, 1904: 506, pl. 48, f. 5, 6 (Dm, f, S of Epeira cambridgei)

Eustala illicita Chickering, 1955: 401, f. 6-11 (mf)

Eustala inconstans Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015943

Eustala inconstans Chickering, 1955: 446, f. 69-72 (Dm) Original description

Eustala ingenua Chickering, 1955 | | Guatemala, Panama urn:lsid:nmbe.ch:spidersp:015944

Eustala ingenua Chickering, 1955: 449, f. 73 (Df) Original description

Eustala longembola Chickering, 1955: 458, f. 81-85 (Dm) [urn:lsid:nmbe.ch:spidersp:015950]

Eustala ingenua Levi & Santos, 2013: 223 (S of Eustala longembola)

Eustala innoxia Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015945

Eustala innoxia Chickering, 1955: 452, f. 74-76 (Df) Original description

Eustala itapocuensis Strand, 1916 | | Brazil urn:lsid:nmbe.ch:spidersp:015947

Eustala itapocuensis Strand, 1916b: 107 (Dm) Original description

Eustala itapocuensis Poeta, Marques & Buckup, 2010b: 270, f. 16-21 (m, Df)

Eustala lata Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015948

Eustala lata Chickering, 1955: 454, f. 77 (Df) Original description

Eustala latebricola (O. Pickard-Cambridge, 1889) | | Guatemala to Panama urn:lsid:nmbe.ch:spidersp:015949

Epeira latebricola O. Pickard-Cambridge, 1889d: 31, pl. 8, f. 4 (Df) Original description

Epeira latebricola Keyserling, 1892: 150, pl. 7, f. 110 (f)

Eustala latebricola F. O. Pickard-Cambridge, 1904: 508, pl. 48, f. 15 (f)

Eustala latebricola Chickering, 1955: 457, f. 78-80 (f)

Eustala levii Poeta, Marques & Buckup, 2010 | | Brazil urn:lsid:nmbe.ch:spidersp:043573

Eustala levii Poeta, Marques & Buckup, 2010a: 151, f. 8-13 (Dmf) Original description

Eustala lisei Poeta, 2014 | | Brazil, Uruguay urn:lsid:nmbe.ch:spidersp:047226

Eustala lisei Poeta, 2014: 448, f. 38-46 (Dmf) Original description

Eustala lunulifera Mello-Leitão, 1939 | | French Guiana, Guyana urn:lsid:nmbe.ch:spidersp:015951

Eustala lunulifera Mello-Leitão, 1939g: 108, f. 7-8 (Df) Original description

Eustala lunulifera Caporiacco, 1954: 90, f. 17 (Dm)

Eustala maxima Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015952

Eustala maxima Chickering, 1955: 461, f. 86-87 (Df) Original description

Eustala meridionalis Baert, 2014 | | Ecuador (Galapagos Is.) urn:lsid:nmbe.ch:spidersp:046986

Eustala meridionalis Baert, 2014b: 12, f. 4A-C, 5B, 6B (Dmf) Original description

Eustala mimica Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015953

Eustala mimica Chickering, 1955: 467, f. 90-93 (Dm) Original description

Eustala minuscula (Keyserling, 1892) | | Brazil urn:lsid:nmbe.ch:spidersp:015954

Epeira minuscula Keyserling, 1892: 140, pl. 7, f. 103 (Dmf) Original description

Araneus minusculus Petrunkevitch, 1911: 304

Eustala minuscula Mello-Leitão, 1919: 478

Mangora minuscula Roewer, 1942a: 774

Eustala minuscula Mello-Leitão, 1943a: 179

Eustala monticola Chamberlin, 1916 | | Peru urn:lsid:nmbe.ch:spidersp:015956

Eustala monticola Chamberlin, 1916: 246, pl. 18, f. 7 (Df) Original description

Eustala montivaga Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015957

Eustala montivaga Chickering, 1955: 478, f. 101-102 (Df) Original description

Eustala montana Chickering, 1955: 474, f. 97-100 (Dm) [urn:lsid:nmbe.ch:spidersp:015955]

Eustala montivaga Levi & Santos, 2013: 223 (S of Eustala montana)

Eustala mourei Mello-Leitão, 1947 | | Brazil urn:lsid:nmbe.ch:spidersp:015958

Eustala mourei Mello-Leitão, 1947b: 241, f. 9-10 (Dmf) Original description

Eustala mourei Poeta, 2014: 455, f. 75-84 (mf)

Eustala mucronatella (Roewer, 1942) | | Brazil urn:lsid:nmbe.ch:spidersp:015041

Epeira mucronata L. Koch, 1871: 74, pl. 6, f. 2-3 (Dmf; preoccupied by Blackwall, 1862, see Araneus mucronatus) [urn:lsid:nmbe.ch:spidersp:061416]

Aranea mucronatella Roewer, 1942a: 830 (replacement name) Original description

Eustala mucronatella Framenau, Scharff & Levi, 2009: 26, f. 4A-H (mf, T from Araneus)

Eustala nasuta Mello-Leitão, 1939 | | Panama, Guyana, Brazil urn:lsid:nmbe.ch:spidersp:015959

Eustala nasuta Mello-Leitão, 1939g: 107, f. 5-6 (Df) Original description

Eustala tumida Chickering, 1955: 508, f. 131-132 (Df) [urn:lsid:nmbe.ch:spidersp:015990]

Eustala nasuta Poeta, Marques & Buckup, 2010b: 273, f. 32-33 (Dm)

Eustala nasuta Levi & Santos, 2013: 222 (S of Eustala tumida)

Eustala novemmamillata Mello-Leitão, 1941 | | Argentina urn:lsid:nmbe.ch:spidersp:015961

Eustala 9-mamillata Mello-Leitão, 1941f: 212, f. 16-17 (Df) Original description

Eustala novemmamillata Brignoli, 1983c: 270

Eustala oblonga Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015962

Eustala oblonga Chickering, 1955: 404, f. 12-18 (Dmf) Original description

Eustala occidentalis Baert, 2014 | | Ecuador (Galapagos Is.) urn:lsid:nmbe.ch:spidersp:046984

Eustala occidentalis Baert, 2014b: 3, f. 1A-E, 2A-F, 5A, 6A (Dmf) Original description

Eustala orientalis Baert, 2014 | | Ecuador (Galapagos Is.) urn:lsid:nmbe.ch:spidersp:046985

Eustala orientalis Baert, 2014b: 9, f. 3A-E, 5C, 6C (Dmf) Original description

Eustala pallida Mello-Leitão, 1940 | | Brazil urn:lsid:nmbe.ch:spidersp:015964

Eustala pallida Mello-Leitão, 1940h: 25, f. 2-3 (Df) Original description

Eustala palmares Poeta, Marques & Buckup, 2010 | | Brazil, Uruguay, Argentina urn:lsid:nmbe.ch:spidersp:043574

Eustala palmares Poeta, Marques & Buckup, 2010a: 152, f. 14-19 (Dmf) Original description

Eustala perdita Bryant, 1945 | | Hispaniola urn:lsid:nmbe.ch:spidersp:015966

Eustala perdita Bryant, 1945a: 373, f. 4, 9 (Dmf) Original description

Eustala perfida Mello-Leitão, 1947 | | Brazil, Uruguay urn:lsid:nmbe.ch:spidersp:015967

Eustala perfida Mello-Leitão, 1947b: 243, f. 11 (Df) Original description

Eustala perfida Poeta, Marques & Buckup, 2010b: 271, f. 22-26 (f, Dm)

Eustala photographica Mello-Leitão, 1944 | | Brazil, Uruguay, Argentina urn:lsid:nmbe.ch:spidersp:015968

Eustala photographica Mello-Leitão, 1944b: 329, f. 13-14 (Df) Original description

Eustala photographica Poeta, Marques & Buckup, 2010a: 158, f. 30-34 (f, Dm)

Eustala redundans Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015970

Eustala redundans Chickering, 1955: 484, f. 106-107 (Df) Original description

Eustala rosae Chamberlin & Ivie, 1935 | | USA, Mexico urn:lsid:nmbe.ch:spidersp:015972

Eustala rosae Chamberlin & Ivie, 1935b: 22, pl. 15, f. 124 (Df) Original description

Eustala rosae Gertsch & Davis, 1936: 14, f. 11 (f; m f. 12 see Eustala clavispina)

Eustala rosae Levi, 1977a: 112, f. 192-204, 297, 313 (Dm, f)

Eustala rosae Orozco-Gil & Desales-Lara, 2021: 11, f. 8-9 (f)

Eustala rubroguttulata (Keyserling, 1879) | | Peru urn:lsid:nmbe.ch:spidersp:015973

Larinia rubroguttulata Keyserling, 1879: 314, pl. 4, f. 15 (Df) Original description

Larinia rubroguttulata Keyserling, 1893: 292, pl. 15, f. 216 (f)

Araneus rubroguttulatus Grasshoff, 1970b: 218 (T from Larinia)

Eustala rubroguttulata Harrod, Levi & Leibensperger, 1991: 246 (T from Araneus)

Eustala rustica Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015974

Eustala rustica Chickering, 1955: 486, f. 108-113 (Dmf) Original description

Eustala saga (Keyserling, 1893) | | Brazil, Uruguay urn:lsid:nmbe.ch:spidersp:015975

Epeira saga Keyserling, 1893: 253, pl. 13, f. 188 (Dmf) Original description

Araneus sagus Petrunkevitch, 1911: 313

Eustala saga Roewer, 1942a: 766

Eustala sagana (Keyserling, 1893) | | Brazil urn:lsid:nmbe.ch:spidersp:015976

Epeira sagana Keyserling, 1893: 254, pl. 13, f. 189 (Dmf) Original description

Araneus saganus Petrunkevitch, 1911: 314

Eustala sagana Roewer, 1942a: 766

Eustala scitula Chickering, 1955 | | Mexico to Panama urn:lsid:nmbe.ch:spidersp:015978

Eustala scitula Chickering, 1955: 492, f. 114-117 (Dm) Original description

Eustala scutigera (O. Pickard-Cambridge, 1898) | | Mexico to Panama urn:lsid:nmbe.ch:spidersp:015979

Epeira scutigera O. Pickard-Cambridge, 1898a: 243, pl. 33, f. 8 (Df) Original description

Epeira nava O. Pickard-Cambridge, 1899a: 300, pl. 37, f. 5 (Df) [urn:lsid:nmbe.ch:spidersp:061415]

Eustala scutigera F. O. Pickard-Cambridge, 1904: 508, pl. 48, f. 11-12 (Dm, f, S of Epeira nava)

Eustala scutigera Chickering, 1955: 496, f. 118-123 (mf)

Eustala secta Mello-Leitão, 1945 | | Brazil, Argentina urn:lsid:nmbe.ch:spidersp:015980

Eustala secta Mello-Leitão, 1945b: 238, f. 11 (Df) Original description

Eustala secta Poeta, Marques & Buckup, 2010b: 272, f. 27-31 (f, Dm)

Eustala sedula Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015981

Eustala sedula Chickering, 1955: 500, f. 124-126 (Df) Original description

Eustala semifoliata (O. Pickard-Cambridge, 1899) | | Central America urn:lsid:nmbe.ch:spidersp:015982

Epeira semifoliata O. Pickard-Cambridge, 1899a: 301, pl. 36, f. 3 (Df) Original description

Eustala semifoliata F. O. Pickard-Cambridge, 1904: 510, pl. 48, f. 21 (f)

Eustala semifoliata Chickering, 1955: 502, f. 127 (f)

Eustala smaragdinea (Taczanowski, 1878) | | Peru urn:lsid:nmbe.ch:spidersp:015983

Epeira smaragdinea Taczanowski, 1878b: 161, pl. 2, f. 12 (Df) Original description

Eustala smaragdinea Simon, 1895a: 789

Eustala tantula Chickering, 1955 | | Panama urn:lsid:nmbe.ch:spidersp:015984

Eustala tantula Chickering, 1955: 505, f. 128-130 (Dm) Original description

Eustala minima Chickering, 1955: 471, f. 94-96 (Df) [urn:lsid:nmbe.ch:spidersp:061408]

Eustala tantula Levi & Santos, 2013: 223 (S of Eustala minima, removed from S of Eustala devia, rejecting Levi, 1977a: 101)

Eustala taquara (Keyserling, 1892) | | Brazil, Uruguay, Argentina urn:lsid:nmbe.ch:spidersp:015985

Epeira taquara Keyserling, 1892: 143, pl. 7, f. 105 (Df) Original description

Araneus taquara Petrunkevitch, 1911: 318

Eustala taquara Roewer, 1942a: 766

Eustala taquara Poeta, Marques & Buckup, 2010a: 157, f. 25-29 (f, Dm)

Eustala tribrachiata Badcock, 1932 | | Paraguay urn:lsid:nmbe.ch:spidersp:015986

Eustala tribrachiata Badcock, 1932: 20, f. 15 (Df) Original description

Eustala trinitatis (Hogg, 1918) | | Trinidad urn:lsid:nmbe.ch:spidersp:015988

Araneus trinitatis Hogg, 1918: 166, f. 1 (Df) Original description

Eustala trinitatis Levi, 1991a: 178 (Tf from Araneus)

Eustala ulecebrosa (Keyserling, 1892) | | Brazil, Argentina urn:lsid:nmbe.ch:spidersp:015991

Epeira ulecebrosa Keyserling, 1892: 104, pl. 5, f. 77 (Df) Original description

Araneus ulecebrosus Petrunkevitch, 1911: 321

Eustala ulecebrosa Mello-Leitão, 1919: 470

Eustala ulecebrosa Poeta, 2014: 455, f. 69-74 (f)

Eustala unimaculata Franganillo, 1930 | | Cuba urn:lsid:nmbe.ch:spidersp:015993

Eustala unimaculata Franganillo, 1930: 22 (Df) Original description

Eustala unimaculata Bryant, 1936b: 327, pl. 23, f. 1, 5 (Dm)

Eustala vegeta (Keyserling, 1865) | | Mexico to Brazil, Hispaniola urn:lsid:nmbe.ch:spidersp:015994

Epeira vegeta Keyserling, 1865: 819, pl. 19, f. 31-34 (Dmf) Original description

Epeira vegeta Keyserling, 1892: 144, pl. 7, f. 106 (mf)

Acacesia vegeta Simon, 1895a: 789, f. 858-859 (m)

Eustala vegeta F. O. Pickard-Cambridge, 1904: 509, pl. 48, f. 16-17 (mf)

Eustala vegeta Bryant, 1945a: 374, f. 7 (m)

Eustala vegeta Chickering, 1955: 511, f. 133-138 (mf)

Eustala vellardi Mello-Leitão, 1924 | | Brazil, Paraguay, Argentina urn:lsid:nmbe.ch:spidersp:015995

Eustala vellardi Mello-Leitão, 1924a: 279 (Df) Original description

Eustala vellardi Poeta & Teixeira, 2017: 113, f. 1-9 (Dm, f)

Eustala vellardi Zanone, Larrea & Raimundo, 2019: 151, f. 2-9 (mf)

Eustala viridipedata (Roewer, 1942) | | Peru urn:lsid:nmbe.ch:spidersp:015997

Epeira viridipes Taczanowski, 1878b: 155, pl. 1, f. 8 (Dmf; preoccupied by Doleschall, 1859, see Cyrtophora cylindroides) [urn:lsid:nmbe.ch:spidersp:061417]

Araneus viridipes Petrunkevitch, 1911: 324

Aranea viridipedata Roewer, 1942a: 856 (replacement name) Original description

Eustala viridipedata Levi, 1991a: 177 (T from Araneus)

Eustala wiedenmeyeri Schenkel, 1953 | | Venezuela urn:lsid:nmbe.ch:spidersp:015998

Eustala wiedenmeyeri Schenkel, 1953a: 21, f. 19a-b (Df) Original description