World Spider Catalog

Zodariidae Thorell, 1881

Family | ACCEPTED

N.B.: the family name Zodariidae is protected by usage over the earlier name Cryptothelidae L. Koch, 1872a: 238 to keep nomenclatorial stability; separation of Cryptothelidae (Murphy & Roberts, 2015: viii) not sufficiently justified and is not accepted here; Jocqué, 1991a stated the subfamilies Cryptothelinae, Cydrelinae Simon, 1893, Cyriocteinae Jocqué, 1991, Lachesaninae Jocqué, 1991, Storeninae Simon, 1893, Storenomorphinae Simon, 1890 and Zodariinae; synonymy of the Cydrelinae with the Cryptothelinae considered by Ramírez et al., 2014: 392; a new subfamily Procydrelinae Henrard & Jocqué, 2025 is described in Ortiz et al., 2025: 998.

In synonymy:

Laches Thorell, 1869 = Lachesana Strand, 1932 (Strand, 1932: 140, preoccupied, replacement name)

Lachesis Audouin, 1826 = Lachesana Strand, 1932 (Gistel, 1848: IX, sub Lestes, preoccupied, replacement name)

Lestes Gistel, 1848 = Lachesana Strand, 1932 (Thorell, 1869: 37, sub Laches, preoccupied, replacement name)

Gen. Lachesana Strand, 1932 urn:lsid:nmbe.ch:spidergen:02296

type Lachesis perversa Audouin, 1826; gender feminine

N.B.: Lachesana is a replacement name for Laches Thorell, 1869: 37 [urn:lsid:nmbe.ch:spidergen:05825] (preoccupied in Hymenoptera by Gistel, 1848), Laches is a replacement name for Lestes Gistel, 1848: IX [urn:lsid:nmbe.ch:spidergen:05824] (preoccupied in Odonata by Leach, 1815) and Lestes is a replacement name for Lachesis Audouin, 1826: 110 [urn:lsid:nmbe.ch:spidergen:05823] (preoccupied in Reptilia by Daudin, 1802).

In synonymy:

Lachesana sundaica (Reimoser, 1927) = Lachesana blackwalli (O. Pickard-Cambridge, 1872) (Jocqué, 1991a: 36)

Nomen dubium:

Lachesana vittata (Strand, 1906b: 669, m, Tunisia, originally in Laches) -- Nentwig et al., 2020: 27

Lachesana bayramgocmeni Özkütük, Yağmur, Gücel, Shafaie, Özden & Kunt, 2020 | | Cyprus urn:lsid:nmbe.ch:spidersp:052330

Lachesana bayramgocmeni Özkütük et al., 2020: 335, f. 1-2, 4-7, 9-12 (Dmf) Original description

Lachesana blackwalli (O. Pickard-Cambridge, 1872) | | Greece, Cyprus, Turkey, Israel, Lebanon urn:lsid:nmbe.ch:spidersp:025790

Lachesis blackwalli O. Pickard-Cambridge, 1872: 266, pl. 13, f. 5 (Dmf) Original description

Lachese blackwalli Simon, 1873a: 67

Laches blackwalli Simon, 1893a: 395, 399

Laches blackwalli Kulczyński, 1908c: 56, pl. 2, f. 11-12 (m)

Laches sundaica Reimoser, 1927c: 3, f. 1 (Dm) [urn:lsid:nmbe.ch:spidersp:056312]

Lachesana blackwalli Levy, 1990b: 333, f. 1-6, 8-14 (mf)

Lachesana blackwalli Jocqué, 1991a: 36, f. 16, 52-58 (mf, S of Lachesana sundaica)

Lachesana blackwalli Thaler & Knoflach, 2004a: 41, f. 4, 12, 14, 17, 19, 22, 25 (m)

Lachesana blackwalli Özkütük et al., 2020: 337, f. 3, 8, 13-14 (mf)

Lachesana blackwalli Zamani & Marusik, 2021b: 152, f. 4H (m)

Lachesana blackwalli Bosmans & Gavalas, 2023: 33, f. 18A, 19A-B, 20A-C (m)

Lachesana dyachkovi Fomichev & Marusik, 2019 | | Kazakhstan urn:lsid:nmbe.ch:spidersp:051563

Lachesana dyachkovi Fomichev & Marusik, 2019b: 559, f. 1–5, 8-19, 22 (Dm) Original description

Lachesana dyachkovi Zamani & Marusik, 2021b: 148, f. 4A-C, F (m)

Lachesana gavishae Shafaie & Pekár, 2025 | | Israel urn:lsid:nmbe.ch:spidersp:066042

Lachesana gavishae Shafaie & Pekár in Shafaie, Pekár & Seyyar, 2025: 4, f. 1A-F, 2A-G, 3A-D (Dmf) Original description

Lachesana graeca Thaler & Knoflach, 2004 | | Greece urn:lsid:nmbe.ch:spidersp:040398

Lachesana graeca Thaler & Knoflach, 2004a: 41, f. 1-3, 6, 9-11, 15, 20, 23, 26 (Dm) Original description

Lachesana graeca Bosmans & Gavalas, 2023: 33, f. 18B, 19C-D, 20D-F (m)

Lachesana insensibilis Jocqué, 1991 | | Israel, Saudi Arabia, United Arab Emirates, Iran urn:lsid:nmbe.ch:spidersp:025791

Lachesana insensibilis Jocqué, 1991a: 37, f. 59 (Dm) Original description

Lachesana kavirensis Zamani & Marusik, 2021 | | Iran urn:lsid:nmbe.ch:spidersp:053108

Lachesana kavirensis Zamani & Marusik, 2021b: 148, f. 1A-C, 3A-E, 4I (Dm) Original description

Lachesana lubinae Shafaie & Pekár, 2025 | | Israel urn:lsid:nmbe.ch:spidersp:066043

Lachesana lubinae Shafaie & Pekár in Shafaie, Pekár & Seyyar, 2025: 9, f. 4A-F, 5A-G, 6A-D (Dmf) Original description

Lachesana naxos Wunderlich, 2022 | | Greece urn:lsid:nmbe.ch:spidersp:053746

Lachesana naxos Wunderlich, 2022: 47, f. 99-104 (Dm) Original description

Lachesana naxos Bosmans & Gavalas, 2023: 31, f. 18C, 19E-F, 20G-I (m)

Lachesana perseus Zamani & Marusik, 2021 | | Iran urn:lsid:nmbe.ch:spidersp:053109

Lachesana perseus Zamani & Marusik, 2021b: 149, f. 1D-F, 2, 3F-J, 4J (Dm) Original description

Lachesana perversa (Audouin, 1826) | | Egypt, Syria urn:lsid:nmbe.ch:spidersp:025792 TYPE SPECIES

Lachesis perversa Audouin, 1826: 311, pl. 1, f. 4 (Dm) Original description

Lestes [perversa] Gistel, 1848: IX (generic replacement name)

Laches perversa Thorell, 1869: 37 (generic replacement name)

Lachesana perversa Strand, 1932: 140 (generic replacement name)

Lachesana rufiventris (Simon, 1873) | | Greece, Syria, Israel urn:lsid:nmbe.ch:spidersp:025793

Lachese rufiventris Simon, 1873a: 68 (Dm) Original description

Laches scutiventris Simon, 1893a: 427, f. 396-398 (m, lapsus)

Lachesana rufiventris Levy, 1990b: 335, f. 7, 15-19 (m, Df)

Lachesana rufiventris Jocqué, 1991a: 36, f. 33

Lachesana rufiventris Bosmans & Gavalas, 2023: 33, f. 18D, 19G-H, 20J-L (m)

Lachesana rufiventris Finkel et al., 2024: 25, f. 22C-F, 23B, 24E-H, 25E-H (m)

Lachesana salam Hammouri, Ganem & Gavish-Regev, 2024 | | Syria, Israel, Palestine urn:lsid:nmbe.ch:spidersp:062021

Lachesana rufiventris Thaler & Knoflach, 2004a: 43, f. 5, 7-8, 13, 16, 18, 21, 24, 27 (m; misidentified per Finkel et al., 2024: 24)

Lachesana salam Hammouri, Ganem & Gavish-Regev in Finkel et al., 2024: 24, f. 22A-C, 23A, C, 24A-D, 25A-D, 26A-B (Dm) Original description

Lachesana tarabaevi Zonstein & Ovtchinnikov, 1999 | | Kyrgyzstan, Uzbekistan, Tajikistan urn:lsid:nmbe.ch:spidersp:037328

Lachesana tarabaevi Zonstein & Ovtchinnikov, 1999: 59, f. 1-5 (Dmf) Original description

Lachesana tarabaevi Fomichev & Marusik, 2019b: 559, f. 6-7, 20-21, 23-25 (mf)

Lachesana tarabaevi Zamani & Marusik, 2021b: 152, f. 4D-E, G (m)