Tetragnathidae Menge, 1866

Family |   ACCEPTED
N.B.: considered a senior synonym of Metidae Simon, 1894 by Heimer & Nentwig, 1982: 292 and Levi, 1986a: 93 (although not by Wunderlich, 1986: 115); Metidae/Metinae is preoccupied in the Copepoda and emened to Metainae by ICZN, 2013 (Opinion 2326) following the proposal of Álvarez-Padilla & Hormiga, 2011b; Diphyini is preoccupied in the Hydrozoa and emened to Diphyaini by ICZN, 2021 (Opinion 3774) following the proposal of Marusik, 2018d; see also the notes at the Araneidae, Nephilidae and Phonognathidae; includes taxa placed by Forster & Forster, 1999: 166 in the "Nanometidae" (see Álvarez-Padilla & Hormiga, 2011a: 802); also Leucauginae Caporiacco, 1955: 318 and Pachygnathidae Menge, 1866: 94 are included; mentioning of "Tetragnathiens" by Simon, 1864: 325, but the French ending -iens was not accepted as taxon at familiar level by Bonnet and Platnick, and this is continued here.

In synonymy:

Menosira Chikuni, 1955 = Metellina Chamberlin & Ivie, 1941 (Kallal & Hormiga, 2018a: 415)

Gen. Metellina Chamberlin & Ivie, 1941 urn:lsid:nmbe.ch:spidergen:01389
N.B.: considered a senior synonym of Menosira Chikuni, 1955: 31 [urn:lsid:nmbe.ch:spidergen:01384] (type M. ornata Chikuni, 1955) by Kallal & Hormiga, 2018: 415; the proposed synonymy with Meta C. L. Koch, 1836 (e.g. by Wunderlich, 1987a: 119, 1992a: 361, 2011: 205, Murphy & Roberts, 2015: ix) was not followed by subsequent workers (e.g. Wunderlich, 2008a: 95) and the current status is confirmed by Kallal & Hormiga, 2018.

In synonymy:

Metellina antrorum (Simon, 1907, T from Meta) = Metellina segmentata Clerck, 1757 (Loksa, 1972: 69, sub Meta)

Metellina canariensis (Schmidt, 1975) = Metellina minima Denis, 1953 (Wunderlich, 1987a: 132, sub Meta)

Metellina merianae celata (Blackwall, 1841) = Metellina merianae Scopoli, 1763 (Wiehle, 1931: 126, sub Meta)

Metellina peckhami (McCook, 1894, T from Araneus) = Metellina curtisi McCook, 1894 (Levi, 1980a: 35)

Metellina ramblae (Schmidt, 1975) = Metellina minima Denis, 1953 (Wunderlich, 1987a: 132, sub Meta)

Metellina reticulata (Linnaeus, 1758, T from Meta) = Metellina segmentata Clerck, 1757 (, Clerckian names validated by ICZN, 1959 (Direction 104))

Nomen dubium:

Metellina nigra (Franganillo, 1920: 140, f, Portugal, originally in Meta) -- Wunderlich, 2011: 209

Metellina barreti (Kulczyński, 1899) | | Madeira urn:lsid:nmbe.ch:spidersp:013868
Meta barreti Kulczyński, 1899: 389, pl. 8, f. 79 (Dj) Original description
Meta barreti Schenkel, 1938a: 29, f. 10 (Df)
Meta barretti Wunderlich, 1992a: 362, f. 327-329 (f, Dm)
Metellina barreti Wunderlich, 2008a: 94 (T from Meta)
Metellina barreti Marusik & Larsen, 2018: 209 (T from Meta confirmed)
Metellina curtisi (McCook, 1894) | | North America urn:lsid:nmbe.ch:spidersp:013915
Pachygnatha curtisi McCook, 1894: 271, pl. 26, f. 5 (Dm) Original description
Epeira peckhamii McCook, 1894: 189, pl. 18, f. 5-6 (Dmf) [urn:lsid:nmbe.ch:spidersp:060523]
Araneus peckhami Petrunkevitch, 1911: 308
Meta curtisi Gertsch & Ivie, 1936: 20
Metellina curtisi Chamberlin & Ivie, 1941b: 14, f. 20 (f, T from Meta)
Metellina curtisi Levi, 1980a: 35, f. 72-86, 95-98 (mf, S of Araneus peckhami, removed from S of Metellina mengei, rejecting Levi & Field, 1954: 466)
Metellina curtisi Hormiga, Eberhard & Coddington, 1995: 324, f. 6D-E, 13D-E (m)
Metellina curtisi Dondale et al., 2003: 113, f. 244-254 (mf)
Metellina curtisi Álvarez-Padilla, 2007: 293, f. 7F (m)
Metellina gertschi (Lessert, 1938) | | DR Congo urn:lsid:nmbe.ch:spidersp:013873
Meta gertschi Lessert, 1938: 437, f. 18-19 (Dm) Original description
Metellina gertschi Marusik & Larsen, 2018: 209 (T from Meta)
Metellina haddadi Marusik & Larsen, 2018 | | South Africa urn:lsid:nmbe.ch:spidersp:050449
Metellina haddadi Marusik & Larsen, 2018: 212, f. 1-34 (Dmf) Original description
Metellina haddadi Dippenaar-Schoeman et al., 2020t: 26, p. 25: 1 f., p. 26: 3 f. (mf)
Metellina haddadi Dippenaar-Schoeman et al., 2023a: 29, p. 28: 1 f., p. 29: 1-3 (mf)
Metellina kirgisica (Bakhvalov, 1974) | | Azerbaijan, Central Asia, China urn:lsid:nmbe.ch:spidersp:013916
Meta kirgisicus Bakhvalov, 1974: 101, f. 6-7 (Df) Original description
Meta kirgisica Bakhvalov, 1982: 136, f. 1 (f)
Meta kirgisica Bakhvalov, 1983: 86, f. 1 (f)
Metellina kirgisica Marusik, 1989b: 44 (T from Meta)
Metellina kirgisica Marusik, Fritzén & Song, 2007: 271, f. 31, 52 (f)
Metellina kirgisica Özkütük et al., 2017a: 79, f. 7-8, 11-15, 20-22, 26-28, 37-38, 40-46 (Dm, f)
Metellina longipalpis (Pavesi, 1883) | | Ethiopia urn:lsid:nmbe.ch:spidersp:013875
Meta longipalpis Pavesi, 1883: 19 (Dmf) Original description
Metellina longipalpis Marusik & Larsen, 2018: 209 (T from Meta)
Metellina mengei (Blackwall, 1869) | | Europe to Caucasus, Iran, Russia (Europe to Altai) urn:lsid:nmbe.ch:spidersp:013917
Epeira mengii Blackwall, 1869: 398 (Dmf) Original description
Meta segmentata mengei Chyzer & Kulczyński, 1891: 139
Meta segmentata mengei Kulczyński, 1899: 389, pl. 8, f. 78 (f)
Meta mengei Bösenberg, 1901: 49, pl. 3, f. 40 (mf)
Meta segmentata mengei Fedotov, 1912a: 78, f. 27-28 (mf)
Meta reticulata mengei Wiehle, 1931: 123, f. 198-201 (mf)
Meta reticulata mengei Drensky, 1943: 247, f. 30 (mf)
Meta segmentata mengei Locket & Millidge, 1953: 117, f. 76C, 77B, E (mf)
Meta mengei Chrysanthus, 1953: 160, f. 1b-d, 2b, d-e, 3a-b (mf, elevated from subspecies of Meta reticulata = segmentata)
Meta mengei Levi & Field, 1954: 466 (S of Epeira peckhami; rejected by Levi, 1980a: 36, see Metellina curtisi)
Meta mengei Chrysanthus, 1955c: 59, f. 2A-D (m)
Meta segmentata mengei Zamaraev, 1964: 352, f. 15B, 16B, 17 (mf)
Meta mengei Wiehle, 1967b: 191, f. 33-34 (m)
Meta segmentata mengei Loksa, 1972: 70, f. 60D-F (mf)
Meta mengei Locket, Millidge & Merrett, 1974: 64, f. 37A, C (mf)
Meta mengei Palmgren, 1974a: 39, f. 17a-g (mf)
Meta mengei Punda, 1975b: 42, f. 79, 81 (mf)
Metellina mengei Levi, 1980a: 34, f. 105-107 (mf, T from Meta)
Meta mengei Roberts, 1985: 202, f. 90b, g, 91b (mf)
Metellina mengei Heimer & Nentwig, 1991: 64, f. 140 (mf)
Meta mengei Roberts, 1995: 308, f. (mf)
Metallina mengei Agnarsson, 1996: 59, f. 45A-C (mf)
Metellina mengei Bellmann, 1997: 98, f. (mf)
Meta mengei Roberts, 1998: 322, f. (mf)
Metellina mengei Trotta, 2005: 174, f. 430, 433 (mf)
Metellina mengei Almquist, 2005: 113, f. 132a-h (mf)
Meta mengei Wunderlich, 2011: 206, f. 5, 5a (mf)
Metellina mengei Marusik, Özkütük & Kunt, 2012: 70, f. 1, 3 (m)
Metellina mengei Kallal & Hormiga, 2018a: 408, f. 3B (m)
Metellina mengei Morano, 2020: 105, f. 8, 12-13 (mf)
Metellina merianae (Scopoli, 1763) | | Europe, Caucasus, Turkey, Iran, Russia (Europe to Central Asia) urn:lsid:nmbe.ch:spidersp:013918
Aranea merianae Scopoli, 1763: 395 (D) Original description
Aranea fusca De Geer, 1778: 335, pl. 11, f. 9-12 (Df)
Aranea fusca Olivier, 1789: 204
Aranea antriada Walckenaer, 1802: 201 (Df)
Epeira antriada Walckenaer, 1805: 62
Epeira inclinata Sundevall, 1833a: 250 (Df, var. b)
Meta merianae C. L. Koch, 1835: 134, pl. 14-16 (mf)
Meta merianae C. L. Koch, 1841a: 121, f. 688-690 (Dmf)
Meta muraria C. L. Koch, 1841a: 125, f. 693-694 (Dmf)
Epeira celata Blackwall, 1841a: 668 (Dmf) [urn:lsid:nmbe.ch:spidersp:013919]
Meta fusca Thorell, 1856: 98
Meta fusca Westring, 1861: 76 (Dm)
Zilla antriada Simon, 1864: 237
Epeira antriada Blackwall, 1864a: 351, pl. 26, f. 253 (mf)
Epeira celata Blackwall, 1864a: 353, pl. 26, f. 254 (mf)
Meta muraria Menge, 1866: 88, pl. 14, f. 25 (mf)
Meta merianae Thorell, 1870a: 36
Meta subterranea Lebert, 1877: 137 (Dm)
Meta merianae Chyzer & Kulczyński, 1891: 139, pl. 6, f. 2 (mf)
Meta merianae celata Chyzer & Kulczyński, 1891: 139
Meta merianae Simon, 1894a: 730, f. 811 (m)
Meta merianae Becker, 1896: (2): 58, pl. 11, f. 4 (mf)
Meta merianae Bösenberg, 1901: 47, pl. 3, f. 38 (mf)
Meta merianae Bacelar, 1929: 260, f. 13-14 (mf)
Meta merianae Simon, 1929: 656, 752, f. 1007-1008 (mf)
Meta merianae Wiehle, 1931: 125, f. 202-204 (mf, S of Meta merianae celata)
Meta merianae Drensky, 1943: 247, f. 31 (mf)
Meta merianae Locket & Millidge, 1953: 117, f. 76D, 77C (mf)
Meta merianae Miller, 1971: 201, pl. XXXVII, f. 7 (m)
Meta merianae Loksa, 1972: 71, f. 61A-B (mf)
Meta merianae Locket, Millidge & Merrett, 1974: 61, f. 36C (m)
Meta merianae Palmgren, 1974a: 41, f. 18a-d (mf)
Metellina merianae Levi, 1980a: 34, f. 108-111 (mf, T from Meta)
Meta merianae Heimer, 1982: 45, f. 14 (m)
Meta merianae Roberts, 1985: 202, f. 90c, 91c (mf)
Metellina merianae Levy, 1987: 255, f. 57-62 (f)
Metellina merianae Heimer & Nentwig, 1991: 64, f. 138 (mf)
Meta merianae Roberts, 1995: 309, f. (mf)
Metellina merianae Roberts, 1998: 322, f. (mf)
Metellina merianae Trotta, 2005: 174, f. 428, 431 (mf)
Metellina merianae Almquist, 2005: 114, f. 133a-g (mf)
Metellina merianae Jäger, 2006d: 3, f. 1-3c-f (mf)
Meta merianae Wunderlich, 2008a: 95, f. 18, 19a-b (m)
Meta merianae Wunderlich, 2011: 209, f. 9 (m, S of Meta merianae celata confirmed)
Metellina merianae Isaia et al., 2011: 148, f. 2.85A-B, 2.86 (mf)
Metellina merianae Marusik et al., 2014a: 262, f. 8a-d (mf)
Metellina merianae Morano, 2020: 105, f. 7, 10-11 (mf)
Metellina merianae Hoyas, 2022: 168, f. 3-4 (f)
Metellina merianopsis (Tullgren, 1910) | | Tanzania urn:lsid:nmbe.ch:spidersp:013880
Meta merianopsis Tullgren, 1910: 151, pl. 3, f. 75 (Df) Original description
Metellina merianopsis Marusik & Larsen, 2018: 210 (T from Meta)
Metellina mimetoides Chamberlin & Ivie, 1941 | | North America urn:lsid:nmbe.ch:spidersp:013920
Metellina mimetoides Chamberlin & Ivie, 1941b: 15, f. 19 (Df) Original description
Metellina mimetoides Levi, 1980a: 36, f. 87-94 (f, Dm)
Metellina mimetoides Dondale et al., 2003: 109, f. 235-239 (mf)
Metellina minima (Denis, 1953) | | Canary Is. urn:lsid:nmbe.ch:spidersp:013883
Meta obscura minima Denis, 1953c: 6 (Df) Original description
Meta obscura Denis, 1953c: 4, f. 4 (f; misidentified per Denis, 1962a: 82)
Meta minima Denis, 1962a: 82 (elevated from subspecies)
Meta canariensis Schmidt, 1975a: 226, f. 6-7 (Dmf) [urn:lsid:nmbe.ch:spidersp:060522]
Meta ramblae Schmidt, 1975a: 227, f. 8-9 (Df) [urn:lsid:nmbe.ch:spidersp:060524]
Meta minima Wunderlich, 1987a: 132 (S of Meta canariensis and M. ramblae)
Meta minima Wunderlich, 1992a: 363, f. 330-332 (mf)
Metellina minima Wunderlich, 2008a: 94, f. 17 (m, T from Meta)
Metellina minima Marusik & Larsen, 2018: 211 (T from Meta confirmed)
Metellina orientalis (Spassky, 1932) | | Turkey, Armenia, Iran, Kazakhstan, Turkmenistan urn:lsid:nmbe.ch:spidersp:013921
Meta orientalis Spassky, 1932a: 184, f. 5-8 (Dmf) Original description
Metellina orientalis Marusik, 1985b: 139 (T from Meta)
Metellina orientalis Marusik, 1986: 19, f. 1.1-3 (mf)
Metellina orientalis Malek Hosseini, Zamani & Sadeghi, 2015: 92, f. 3a-c (m)
Metellina orientalis Özkütük et al., 2017a: 76, f. 1-6, 9-10, 17-19, 23-25, 29-36, 39 (mf)
Metellina ornata (Chikuni, 1955) | | Russia (Far East), China, Korea, Japan urn:lsid:nmbe.ch:spidersp:013856
Menosira ornata Chikuni, 1955: 31, f. 2A-H, pl. 4 (Dmf, species attributed to Kishida, which means in this case Kishida had confirmed it, Ono in litt.) Original description
Menosira ornata Namkung, Paik & Yoon, 1972: 93, f. 12 (f)
Menosira ornata Yaginuma, 1986a: 129, f. 69.1 (mf)
Menosira ornata Chikuni, 1989b: 92, f. 9 (mf)
Menosira ornata Kim, Kim & Lee, 1999: 51, f. 7, 32, 54 (mf)
Menosira ornata Namkung, 2002: 227, f. 18.14a-c (mf)
Menosira ornata Kim & Cho, 2002: 317, f. 859-864 (f)
Menosira ornata Zhu, Song & Zhang, 2003: 60, f. 25A-I, pl. IIE-H (mf)
Menosira ornata Namkung, 2003: 229, f. 18.14a-c (mf)
Menosira ornata Tanikawa, 2007c: 96, f. 320-321, 774-775 (mf)
Menosira ornata Kim, Kim & Lee, 2008: 36, f. 7, 32, 32.1-2 (mf)
Menosira ornata Tanikawa, 2009: 406, f. 11-12 (mf)
Menosira ornata Marusik & Kovblyuk, 2011: 235, f. 34.21 (m)
Menosira ornata Kim & Lee, 2013: 43, f. 26A-C, pl. 13 (mf)
Menosira ornata Marusik et al., 2015: 211, f. 36-43 (mf)
Menosira ornata Baba & Tanikawa, 2015: 34, 4 f. (mf)
Metellina ornata Kallal & Hormiga, 2018a: 415 (T from Menosira)
Metellina segmentata (Clerck, 1757) | | Europe, Turkey, Israel, Caucasus, Russia (Europe to South Siberia), Kazakhstan, Iran, China, Japan. Introduced to Canada urn:lsid:nmbe.ch:spidersp:013922
Araneus segmentatus Clerck, 1757: 45, pl. 2, f. 6 (Dmf; Clerckian names validated by ICZN, 1959: Direction 104) Original description
Aranea reticulata Linnaeus, 1758: 619 (D) [urn:lsid:nmbe.ch:spidersp:013923]
Aranea angulata Schrank, 1781: 525 (D)
Aranea senoculata Cyrillus, 1787: 1, pl. 8, f. 7
Aranea segmentata Olivier, 1789: 206
Aranea reticulata Olivier, 1789: 231
Araneus segmentatus Martyn, 1793: 20, pl. 1, f. 1-2 (mf)
Aranea inclinata Walckenaer, 1802: 201
Epeira inclinata Walckenaer, 1805: 62
Epeira variegata Risso, 1826: 170 (D)
Zilla reticulata C. L. Koch, 1839a: 142, f. 532-533 (mf)
Epeira inclinata Walckenaer, 1841: 82 (Dmf)
Epeira segmentata Westring, 1851: 35
Meta segmentata Westring, 1861: 80 (Dmf)
Epeira inclinata Blackwall, 1864a: 354, pl. 26, f. 255 (mf)
Meta segmentata Menge, 1866: 86, pl. 14, f. 24 (mf)
Zilla inclinata Sordelli, 1868: 471
Meta segmentata Thorell, 1870a: 39
Meta segmentata Hansen, 1882: 24, pl. 1, f. 40, pl. 2, f. 1 (mf)
Miranda piniophila Bremi-Wolff in Pavesi, 1884b: 127 (nomen nudum)
Meta segmentata Chyzer & Kulczyński, 1891: 138, pl. 6, f. 1 (mf)
Meta segmentata Becker, 1896: (2): 56, pl. 11, f. 3 (mf) [includes M. mengei]
Meta segmentata Bösenberg, 1901: 48, pl. 3, f. 39 (mf)
Meta antrorum Simon, 1907a: 544, f. 2B (Dmf) [urn:lsid:nmbe.ch:spidersp:060521]
Meta segmentata Engelhardt, 1910: 40, f. 2-3 (f)
Meta segmentata Osterloh, 1922: 359, f. 17-19, 28-30 (mf)
Meta segmentata Simon, 1929: 653, 752, f. 1005-1006 (mf)
Meta reticulata Reimoser, 1929c: 86, f. 4 (mf)
Meta reticulata Wiehle, 1931: 119, f. 192-197 (mf)
Meta reticulata Drensky, 1943: 247, f. 30 (mf)
Meta segmentata Locket & Millidge, 1953: 115, f. 75, 76B, 77A, d (mf)
Meta segmentata Chrysanthus, 1953: 160, f. 1a, 2a, c, 3c-d (mf)
Meta segmentata Saito, 1959: 101, f. 120a-c (f)
Meta segmentata Zamaraev, 1964: 352, f. 14, 15A, 16A (mf)
Meta segmentata Wiehle, 1967b: 191, f. 30, 32 (mf)
Meta segmentata Miller, 1971: 201, pl. XXXVII, f. 5-6 (mf)
Meta segmentata Loksa, 1972: 69, f. 59A, 60A-C (mf, S of Meta antrorum)
Meta segmentata Locket, Millidge & Merrett, 1974: 64, f. 36A, 37B, D (mf)
Meta segmentata Palmgren, 1974a: 40, f. 1a, c, 17h-m (mf)
Meta segmentata Punda, 1975b: 44, f. 75, 78, 80 (mf)
Metellina segmentata Levi, 1980a: 36, f. 99-104 (Tmf from Meta)
Meta segmentata Roberts, 1985: 202, f. 90a, f, 91a (mf)
Metellina segmentata Yaginuma, 1986c: 193, f. 2B, 3B, D (mf)
Metellina segmentata Levy, 1987: 254, f. 55-56 (f)
Metellina segmentata Coddington, 1990: 14, f. 49 (m)
Metellina segmentata Heimer & Nentwig, 1991: 64, f. 139 (mf)
Meta segmentata Roberts, 1995: 307, f. (mf)
Meta segmentata Mcheidze, 1997: 250, f. 540-541 (m)
Metellina segmentata Bellmann, 1997: 98, f. (mf)
Meta segmentata Roberts, 1998: 321, f. (mf)
Metellina segmentata Dondale et al., 2003: 111, f. 240-243 (mf)
Metellina segmentata Trotta, 2005: 174, f. 427, 429, 432 (mf)
Metellina segmentata Almquist, 2005: 115, f. 134a-k (mf)
Metellina segmentata Álvarez-Padilla et al., 2009: 140, f. 11a, 12c, e (mf)
Meta segmentata Wunderlich, 2011: 206, f. 6-7 (m)
Metellina segmentata Álvarez-Padilla & Hormiga, 2011a: 779, f. 66A-F, 67A-H, 68A-F, 69A-F, 70A-D (mf)
Metellina segmentata Marusik, Özkütük & Kunt, 2012: 70, f. 2, 4 (m)
Metellina segmentata Morano, 2020: 106, f. 9, 14-15 (mf)
Metellina tangchao Lin & Yang, 2023 | | China urn:lsid:nmbe.ch:spidersp:059471
Metellina Metellina tangchao sp. nov Lin & Yang, 2023: 131, f. 1A-B, 2A-C, 3A-B, 4A-B (Dmf) Original description
Metellina villiersi (Denis, 1955) | | Guinea urn:lsid:nmbe.ch:spidersp:013904
Meta villiersi Denis, 1955c: 1031, f. 8-1O (Dmf) Original description
Metellina villiersi Marusik & Larsen, 2018: 212 (T from Meta)

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