World Spider Catalog

Salticidae Blackwall, 1841

Family | ACCEPTED

N.B.: Considered a senior synonym of Lyssomanidae by Galiano, 1976b: 67; for illustrations of scale morphology of about 60 species, see Hill, 1979; for a database with diagnostic drawings maintained by J. Prószyński, see: http://www.peckhamia.com/salticidae; for the phylogeny within the family see Maddison & Hedin, 2003b, Maddison, 2015, Maddison et al., 2017 and Prószyński, 2017b.

Gen. Icius Simon, 1876 urn:lsid:nmbe.ch:spidergen:03000

N.B.: type species is not the I. notabilis of Simon, 1871, which is a misidentification of I. subinermis Simon, 1937 (Prószyński, 1983a: 46); considered a senior synonym of Pseudicius Simon, 1885 by Andreeva, Heciak & Prószyński, 1984: 350, but evidence presented by Maddison, 1987: 103, makes this synonymy dubious, and Prószyński, 1987: 151, 164, considers that they "should be considered separate genera."

Transferred to other genera:

Icius abnormis (Bösenberg & Strand, 1906) -- see Phintella

Icius afghanicus Andreeva, Heciak & Prószyński, 1984 -- see Pseudicius

Icius annectans Chamberlin, 1929 -- see Metaphidippus

Icius castriesianus (Grube, 1861) -- see Phintella

Icius cavaleriei (Schenkel, 1963) -- see Phintella

Icius cinctipes Banks, 1900 -- see Attidops

Icius cornutus G. W. Peckham & E. G. Peckham, 1885 -- see Padilla

Icius daisetsuzanus Saito, 1934 -- see Sittisax

Icius davidi (Schenkel, 1963) -- see Phintelloides

Icius decempunctatus Caporiacco, 1941 -- see Helafricanus

Icius difficilis (Bösenberg & Strand, 1906) -- see Phintella

Icius elongatus Karsch, 1879 -- see Mendoza

Icius exornatus G. W. Peckham & E. G. Peckham, 1909 -- see Messua

Icius flavipes Caporiacco, 1935 -- see Afraflacilla

Icius floridanus Banks, 1895 -- see Beata

Icius fontanus Levi, 1951 -- see Paradamoetas

Icius formicarius Emerton, 1891 -- see Tutelina

Icius fulgens (O. Pickard-Cambridge, 1872) -- see Heliophanillus

Icius gyirongensis Hu, 2001 -- see Nandicius

Icius harti Emerton, 1891 -- see Tutelina

Icius himeshimensis (Dönitz & Strand, 1906) -- see Hakka

Icius hongkong Song, Xie, Zhu & Wu, 1997 -- see Psenuc

Icius icioides (Simon, 1889) -- see Pseudicius

Icius kaszabi Żabka, 1985 -- see Pseudicius

Icius koreanus (Wesołowska, 1981) -- see Hakka

Icius linea (Karsch, 1879) -- see Phintella

Icius lucipetus (Simon, 1890) -- see Heliophanillus

Icius magister Karsch, 1879 -- see Mendoza

Icius minimus Caporiacco, 1933 -- see Pellenes

Icius munitus (Bösenberg & Strand, 1906) -- see Phintella

Icius nepalicus Andreeva, Heciak & Prószyński, 1984 -- see Pseudicius

Icius originalis Żabka, 1985 -- see Pseudicius

Icius parvus Wesołowska, 1981 -- see Phintella

Icius pleuralis Banks, 1896 -- see Beata

Icius popovi Prószyński, 1979 -- see Phintella

Icius pseudoicioides Caporiacco, 1935 -- see Pseudicius

Icius pupus (Karsch, 1879) -- see Phintella

Icius rugosus Suguro & Nagano, 2015 -- see Hakka

Icius sexmaculatus Banks, 1895 -- see Ghelna

Icius similis Banks, 1895 -- see Tutelina

Icius spasskyi Andreeva, Heciak & Prószyński, 1984 -- see Pseudicius

Icius suedicola (Simon, 1901) -- see Heliophanillus

Icius texanus Banks, 1904 -- see Metaphidippus

Icius tokarensis Bohdanowicz & Prószyński, 1987 -- see Pseudicius

Icius tschekiangensis (Schenkel, 1963) -- see Phintelloides

Icius versicolor G. W. Peckham & E. G. Peckham, 1909 -- see Terralonus

Icius vulpes (Grube, 1861) -- see Pseudicius

Icius wickhami G. W. Peckham & E. G. Peckham, 1894 -- see Beata

In synonymy:

Icius lamellatus Wunderlich, 2011 = Icius subinermis Simon, 1937 (Wunderlich, 2023: 75)

Icius lucasi (Simon, 1868, T from Attus) = Icius erraticus Lucas, 1846 (Simon, 1876a: 57, sub Icius notabilis)

Nomina dubia:

Icius angustatus (Lucas, 1846: 167, pl. 8, f. 6, f, Algeria, originally in Salticus) -- Alicata & Cantarella, 1994: 115

Icius boryi (Lucas, 1846: 157, pl. 7, f. 3, f, Algeria, originally in Salticus) -- Alicata & Cantarella, 1994: 116

Icius discatus Karsch, 1892: 301 (j, Sri Lanka) -- Roewer, 1955c: 1532

Icius erraticus (Lucas, 1846: 149, pl. 6, f. 5, j, Algeria, originally in Salticus) -- Alicata & Cantarella, 1994: 115

Icius eximius Rainbow, 1899b: 317, pl. 25, f. 8-8b (m, Solomon Is.) -- Roewer, 1955c: 1532

Icius guyoni (Lucas, 1846: 156, pl. 7, f. 6, j, Algeria, originally in Salticus) -- Alicata & Cantarella, 1994: 115

Icius miniamus Franganillo, 1910: 19 (f, Portugal) -- Alicata & Cantarella, 1994: 114

Icius pseudocellatus Strand, 1907a: 547, Strand, 1907n: 718 (f, South Africa) -- Nentwig et al., 2020: 25

Nomina nuda:

Icius afolius Franganillo, 1925: 35 (Spain) [urn:lsid:nmbe.ch:spidersp:054508] -- Alicata & Cantarella, 1994: 114, no description

Icius foliosus Franganillo, 1920: 139, 144 (Portugal) [urn:lsid:nmbe.ch:spidersp:054735] -- Alicata & Cantarella, 1994: 114, no description

Icius abnormis Denis, 1958 | | Afghanistan urn:lsid:nmbe.ch:spidersp:034084

Icius abnormis Denis, 1958b: 108, f. 40 (Df) Original description

Icius alboterminus (Caleb, 2014) | | Pakistan, India, Nepal urn:lsid:nmbe.ch:spidersp:047081

Phintella alboterminus Caleb, 2014: 605, f. 1-12 (Dmf) Original description

Icius alboterminus Caleb, 2017: 327 (T from Phintella)

Icius alboterminus Prajapati & Kamboj, 2020a: 15476, f. 1A-B, Images 1A-H, 2A-H, 3A-D (m)

Icius alboterminus Ali, 2021: 82, f. 1A-B, 2C-D (m)

Icius bamboo Cao & Li, 2016 | | China urn:lsid:nmbe.ch:spidersp:048889

Icius bamboo Cao & Li in Cao, Li & Żabka, 2016: 80, f. 26A-D, 27A-B (Dm) Original description

Icius bandama Wesołowska & Russell-Smith, 2022 | | Ivory Coast urn:lsid:nmbe.ch:spidersp:056028

Icius bandama Wesołowska & Russell-Smith, 2022: 45, f. 27A-D, 28A-D (Dmf) Original description

Icius bilobus Yang & Tang, 1996 | | China urn:lsid:nmbe.ch:spidersp:034085

Icius bilobus Yang & Tang, 1996: 105, f. 2.1-5 (Dm) Original description

Icius bilobus Peng, 2020: 183, f. 121a-c (m)

Icius brunellii Caporiacco, 1940 | | Ethiopia urn:lsid:nmbe.ch:spidersp:034086

Icius brunellii Caporiacco, 1940c: 857, f. 50 (Dm) Original description

Icius cervinus Simon, 1878 | | Russia (Europe) urn:lsid:nmbe.ch:spidersp:034087

Icius cervinus Simon, 1878b: 204 (Df) Original description

Icius cervinus Prószyński, 1984a: 69 (m)

Icius congener (Simon, 1871) | | Portugal, Spain, Andorra, France (Corsica), Italy, Malta, Algeria, Libya urn:lsid:nmbe.ch:spidersp:034088

Attus nebulosus Simon, 1868b: 572 (mf, misidentified, not Dendryphantes nebulosus C. L. Koch, 1846 = Macaroeris nidicolens Walckenaer, 1802; not originally described as new species, contra Roewer, 1955: 1220]

Attus congener Simon, 1871a: 184 (Dm) Original description

Icius congener Simon, 1876a: 58 (Df)

Icius congener Simon, 1937: 1216, 1217, 1264, f. 1948 (mf)

Icius congener Prószyński, 1987: 46 (m, not I. nebulosus as per Roewer)

Icius congener Alicata & Cantarella, 1994: 120, f. 2, 8, 14, 37-45 (mf; N.B.: f may be misidentified, per Logunov, 2015: 67)

Icius congener Wunderlich, 2023: 76, f. 191-192 (m)

Icius crassipes (Simon, 1868) | | Spain, Algeria, Tunisia urn:lsid:nmbe.ch:spidersp:034089

Attus crassipes Simon, 1868b: 574 (Dm) Original description

Icius crassipes Simon, 1876a: 56

Icius crassipes Prószyński, 1987: 47 (m)

Icius crassipes Alicata & Cantarella, 1994: 123, f. 4, 10, 46, 59-61 (m)

Icius crassipes Logunov, 2015a: 45, f. 47-52 (m, Df)

Icius crassipes Wunderlich, 2023: 76, f. 190 (m)

Icius dendryphantoides Strand, 1909 | | South Africa urn:lsid:nmbe.ch:spidersp:034091

Icius dendryphantoides Strand, 1909g: 592 (Df) Original description

Icius dendryphantoides Prószyński, 1987: 46 (f)

Icius desertorum Simon, 1901 | | South Africa urn:lsid:nmbe.ch:spidersp:034092

Icius desertorum Simon, 1901c: 161 (Df) Original description

Icius fagei (Lessert, 1925) | | Tanzania urn:lsid:nmbe.ch:spidersp:032945

Cosmophasis fagei Lessert, 1925a: 453, f. 33-39 (Dmf) Original description

Icius fagei Wesołowska, 2012a: 208, f. 19-26 (Tmf from Cosmophasis)

Icius faker Yang & Zhang, 2024 | | China urn:lsid:nmbe.ch:spidersp:060619

Icius faker Yang & Zhang, 2024: 156, f. 28-53 (Dmf) Original description

Icius glaucochirus (Thorell, 1890) | | Indonesia (Sumatra) urn:lsid:nmbe.ch:spidersp:034093

Maevia glaucochira Thorell, 1890c: 163 (Dm) Original description

Chrysilla glaucochira Simon, 1901a: 544

Icius glaucochira Prószyński, 1984a: 69 (Tm from Chrysilla)

Icius grassei (Berland & Millot, 1941) | | Malawi, Nigeria, DR Congo, Kenya urn:lsid:nmbe.ch:spidersp:036597

Mithion grassei Berland & Millot, 1941: 356, f. 56E-H (Df) Original description

Pseudicius grassei Wesołowska & Edwards, 2012: 758, f. 85-87 (Tf from Mithion=Telamonia)

Icius grassei Wesołowska, 2017: 253, f. 1A-F, 2A-D, 3A-G, 4A-E (Dm, f, T from Pseudicius)

Icius hamatus (C. L. Koch, 1846) | | Atlantic Is., North Africa, Southern Europe, Turkey, Georgia. Introduced to Western and Central Europe urn:lsid:nmbe.ch:spidersp:034094

Marpissa hamata C. L. Koch, 1846: 67, f. 1132 (Df) Original description

Icelus notabilis C. L. Koch, 1846: 174, f. 1225 (Dm)

Dendryphantes hamatus Simon, 1864: 314

Attus striatus Simon, 1868b: 566, pl. 6, f. 3 (mf, misidentified)

Attus vicinus Simon, 1868b: 569 (Dmf)

Attus hamatus Simon, 1871a: 183

Icius striatus Simon, 1876a: 59

Icius hamatus Peckham & Peckham, 1886: 306

Icius striatus Simon, 1901a: 614, f. 738 (m)

Icius hamatus Simon, 1937: 1216, 1217, 1264, f. 1946 (mf)

Icius hamatus Prószyński, 1976: 154, f. 233, 403-406 (m)

Icius hamatus Hansen, 1982: 55, f. 1-2 (mf)

Icius hamatus Prószyński, 1983a: 46, f. 9-11 (mf)

Icius hamatus Andreeva, Hęciak & Prószyński, 1984: 350, f. 1-5 (mf)

Icius hamatus Prószyński, 1984a: 41 (f, S of Euophrys altera, rejected by Prószyński, Lissner & Schäfer, 2018: 43)

Icius hamatus Prószyński, 1987: 47-48 (mf; pp. 151-152 treats Araneus striatus Clerck, 1757, which was "nicht zu deuten! (Salticidae?)" for Roewer, as a synonym; if this synonymy is corroborated, Clerck's name would have priority but insufficient usage; see also Bonnet, 1957: 2284, footnote 7, and Alicata & Cantarella, 1994: 116)

Icius hamatus Hu & Li, 1987a: 379, f. 42.1-2, 43.1-3 (mf)

Icius hamatus Baldacchino et al., 1993: 45, f. 3d (m)

Icius hamatus Alicata & Cantarella, 1994: 116, f. 1, 7, 13, 18-36 (mf)

Icius hamatus Fuhn & Gherasim, 1995: 147, f. 68A-E (mf)

Icius hamatus Metzner, 1999: 96, f. 61a-l (mf)

Icius hamatus Hu, 2001: 388, f. 245.1-2 (f; belongs to another species per Rozwałka & Czaja, 2021b: 1)

Icius hamatus Tomasiewicz & Wesołowska, 2006: 339, f. 1-4 (f)

Icius hamatus Schäfer & Deepen-Wieczorek, 2014: 49, f. 1-2 (m)

Icius hamatus Prószyński, 2017b: 26, f. 11A, 12A (mf)

Icius hamatus Prószyński, Lissner & Schäfer, 2018: 43, f. 17A2 (f)

Icius hamatus Breitling, 2020: 355, f. 30A, 32A (mf)

Icius hamatus Peng, 2020: 185, f. 123a-d (mf)

Icius hamatus Rozwałka & Czaja, 2021b: 1, f. 1a-e (mf)

Icius hamatus Killick, 2023: 6, f. 1-5 (m)

Icius hamatus Wunderlich, 2023: 76, f. 189 (m)

Icius hamatus Seropian et al., 2023: 274, f. 98-99 (m)

Icius han Yang & Zhang, 2024 | | China urn:lsid:nmbe.ch:spidersp:060620

Icius han Yang & Zhang, 2024: 161, f. 54-67 (Dmf) Original description

Icius ildefonsus Chamberlin, 1924 | | Mexico urn:lsid:nmbe.ch:spidersp:034096

Icius ildefonsus Chamberlin, 1924b: 679, f. 119 (Df) Original description

Icius indicus (Simon, 1901) | | India, China urn:lsid:nmbe.ch:spidersp:035541

Heliophanus indica Simon, 1901c: 152 (Dm) Original description

Pseudicius indicus Wesołowska, 1986: 231, f. 876-879 (m, T from Heliophanus)

Phintella indica Prószyński, 1992b: 199, f. 190-192 (m, T from Pseudicius)

Icius koreanus Xiao, 1993: 123, f. 1-6 (m; misidentified per Wang et al., 2023e: 9)

Pseudicius koreanus Peng et al., 1993: 192, f. 671-679 (mf; misidentified per Wang et al., 2023e: 9, f uncertain, maybe the unknown f)

Pseudicius koreanus Song, Zhu & Chen, 1999: 542, f. 312M, 313O, 328P (mf = reproduced figs. of Peng et al., 1993: 192, f. 671-679; misidentified per Wang et al., 2023e: 9)

Pseudicius koreanus Yin et al., 2012: 1451, f. 791a-i (mf = reproduced figs. of Peng et al., 1993: 192, f. 671-679; misidentified per Wang et al., 2023e: 9; m = unknown species, per Suguro & Yahata, 2014: 94)

Pseudicius koreanus Peng, 2020: 367, f. 264a-i (mf = reproduced figs. of Peng et al., 1993: 192, f. 671-679; misidentified per Wang et al., 2023e: 9)

Icius indicus Wang et al., 2023e: 9, f. 4A-G (m, T from Phintella)

Icius inhonestus Keyserling, 1878 | | Uruguay urn:lsid:nmbe.ch:spidersp:034097

Icius inhonestus Keyserling, 1878a: 621, pl. 14, f. 23 (Df) Original description

Icius insolidus (Wesołowska, 1999) | | Nigeria, southern Africa urn:lsid:nmbe.ch:spidersp:034557

Menemerus insolidus Wesołowska, 1999: 299, f. 158-161 (Df) Original description

Icius insolidus Wesołowska, 2006b: 234, f. 43-52 (Tf from Menemerus, Dm)

Icius insolidus Haddad & Wesołowska, 2011: 75, f. 45-46, 55-56 (m)

Icius insolitus Alicata & Cantarella, 1994 | | Algeria urn:lsid:nmbe.ch:spidersp:034098

Icius insolitus Alicata & Cantarella, 1994: 127, f. 5, 11, 16, 47, 62-68 (Dm) Original description

Icius insolitus Wunderlich, 2011: 330, f. 10-13 (m)

Icius insolitus Logunov, 2015a: 47, f. 53-62 (m, Df)

Icius kui Yang & Zhang, 2024 | | China urn:lsid:nmbe.ch:spidersp:060621

Icius kui Yang & Zhang, 2024: 164, f. 68-91 (Dmf) Original description

Icius kulakangri Yang & Zhang, 2024 | | China urn:lsid:nmbe.ch:spidersp:060622

Icius kulakangri Yang & Zhang, 2024: 167, f. 92-115 (Dmf) Original description

Icius kumariae Caleb, 2017 | | India urn:lsid:nmbe.ch:spidersp:049820

Icius kumariae Caleb, 2017: 323, f. 1-19 (Dmf) Original description

Icius mbitaensis Wesołowska, 2011 | | Kenya urn:lsid:nmbe.ch:spidersp:044488

Icius mbitaensis Wesołowska, 2011a: 483, f. 7-13, 33 (Dmf) Original description

Icius menemeriformis (Strand, 1907) | | Tanzania urn:lsid:nmbe.ch:spidersp:033831

Heliophanus menemeriformis Strand, 1907a: 546 (Dmf) Original description

Heliophanus menemeriformis Strand, 1907n: 712 (Dmf)

Heliophanus menemeriformis Wesołowska, 1986: 230, f. 866-870 (mf)

Icius menemeriformis Wesołowska, 2024: 91 (T from Heliophanus)

Icius minimus Wesołowska & Tomasiewicz, 2008 | | Ethiopia urn:lsid:nmbe.ch:spidersp:041757

Icius minimus Wesołowska & Tomasiewicz, 2008: 19, f. 72-81, 207-208 (Dmf) Original description

Icius niger Peelle & Saito, 1933 | | Russia (Kurile Is.) urn:lsid:nmbe.ch:spidersp:050725

Icius niger Peelle & Saito, 1933: 119, f. 7 (Df) Original description

Icius niger Saito, 1959: 156, f. 214a-d (f)

Icius nigra Marusik et al., 1993b: 82 (dubious species = nomen dubium)

Icius niger Platnick, 1998: 894 (nomen dubium)

Icius niger Ono & Ogata, 2018: 641 (removed from nomen dubium)

Icius nigricaudus Wesołowska & Haddad, 2009 | | South Africa urn:lsid:nmbe.ch:spidersp:042462

Icius nigricaudus Wesołowska & Haddad, 2009: 51, f. 87-92, 215-217 (Dmf) Original description

Icius ocellatus Pavesi, 1883 | | East Africa urn:lsid:nmbe.ch:spidersp:034100

Icius ocellatus Pavesi, 1883: 85 (Dj) Original description

Icius ocellatus Caporiacco, 1949a: 483, f. 94a-b (Dmf)

Icius ocellatus Prószyński, 1987: 56 (f, "apparently misclassified")

Icius olokomei Wesołowska & Russell-Smith, 2011 | | Nigeria urn:lsid:nmbe.ch:spidersp:044526

Icius olokomei Wesołowska & Russell-Smith, 2011: 577, f. 81-82, 227 (Df) Original description

Icius pallidulus Nakatsudi, 1943 | | Micronesia urn:lsid:nmbe.ch:spidersp:034101

Icius pallidulus Nakatsudi, 1943a: 171, f. 19a-b (Df) Original description

Icius peculiaris Wesołowska & Tomasiewicz, 2008 | | Ethiopia urn:lsid:nmbe.ch:spidersp:041758

Icius peculiaris Wesołowska & Tomasiewicz, 2008: 20, f. 82-86 (Dm) Original description

Icius pulchellus Haddad & Wesołowska, 2011 | | South Africa urn:lsid:nmbe.ch:spidersp:044047

Icius pulchellus Haddad & Wesołowska, 2011: 76, f. 47, 57-62 (Dm) Original description

Icius separatus Banks, 1903 | | Hispaniola urn:lsid:nmbe.ch:spidersp:034103

Icius separatus Banks, 1903: 344, pl. 15, f. 5, 11 (Df) Original description

Icius simoni Alicata & Cantarella, 1994 | | Algeria urn:lsid:nmbe.ch:spidersp:034104

Icius simoni Alicata & Cantarella, 1994: 127, f. 6, 12, 16, 69-76 (Dmf) Original description

Icius steeleae Logunov, 2004 | | Sudan, Kenya, Uganda urn:lsid:nmbe.ch:spidersp:040825

Icius steeleae Logunov, 2004d: 86, f. 3-6 (Dmf) Original description

Icius steeleae Wesołowska, 2011a: 485, f. 14-20, 34 (mf)

Icius subinermis Simon, 1937 | | Mediterranean, Germany, Hungary, Romania. Introduced to USA urn:lsid:nmbe.ch:spidersp:034105

Icius notabilis Simon, 1876a: 57, pl. 9, f. 7 (mf, misidentified)

Icius notabilis Peckham & Peckham, 1894a: 108, pl. 10, f. 7 (f, misidentified)

Icius subinermis Simon, 1937: 1216, 1217, 1264, f. 1945, 1947 (Dmf) Original description

Icius notabilis Kraus, 1955a: 392, f. 37-39 (mf, misidentified)

Icius subinermis Prószyński, 1976: 154, f. 234, 397-402 (mf)

Icius subinermis Andreeva, Hęciak & Prószyński, 1984: 350, f. 6-11 (mf)

Icius subinermis Alicata & Cantarella, 1994: 122, f. 3, 9, 15, 48-58 (mf)

Icius subinermis Jäger, 1995b: 38, f. 1-2 (m)

Icius subinermis Jäger, 1996b: 577, f. 1-2 (m)

Icius lamellatus Wunderlich, 2011: 329, f. 1-9 (Dm) [urn:lsid:nmbe.ch:spidersp:044115]

Icius subinermis Stankovic, 2012: 39, f. 2 (f)

Icius lamellatus Logunov, 2015a: 47 (may be a junior synonym of I. subinermis)

Icius subinermis Korányi, Mezőfi & Markó, 2017: 38, f. 1, 2a-b (f)

Icius subinermis Mezőfi, 2021: 76, f. 1-3 (m)

Icius subinermis Wunderlich, 2023: 75, f. 186-188, 193-194 (m, S of Icius lamellatus)

Icius testaceolineatus (Lucas, 1846) | | Algeria urn:lsid:nmbe.ch:spidersp:034106

Salticus testaceolineatus Lucas, 1846: 183, pl. 10, f. 4 (Df) Original description

Atta testaceolineatus Simon, 1864: 311

Icius testaceolineatus Reimoser, 1919: 109

Icius tukarami Prajapati, Kumbhar & Kamboj, 2021 | | India urn:lsid:nmbe.ch:spidersp:053268

Icius tukarami Prajapati, Kumbhar & Kamboj in Prajapati et al., 2021: 236, f. 26-36 (Dm) Original description

Icius vikrambatrai Prajapati, Malamel, Sudhikumar & Sebastian, 2018 | | India urn:lsid:nmbe.ch:spidersp:050793

Icius vikrambatrai Prajapati et al., 2018: 330, f. 1-17 (Dmf) Original description

Icius yadongensis Hu, 2001 | | China urn:lsid:nmbe.ch:spidersp:038251

Icius yadongensis Hu, 2001: 389, f. 247.1-7 (Dmf) Original description

Hakka yadongensis Peng, 2020: 155, f. 100a-e (mf; following Prószyński, 2016 who lists it in Hakka, but a T was never published)

Icius zang Wang, Mi & Li, 2024 | | China urn:lsid:nmbe.ch:spidersp:060025

Icius zang Wang, Mi & Li, 2024: 155, f. 9A-C, 10A-G (Dmf) Original description

Icius zhengi Yang & Zhang, 2024 | | China urn:lsid:nmbe.ch:spidersp:060623

Icius zhengi Yang & Zhang, 2024: 170, f. 116-138 (Dmf) Original description