Pholcidae C. L. Koch, 1850

Family |   ACCEPTED
N.B.: For comments on the monophyly and relationships of many genera see Huber, 1998d and Huber, 2000. The division of Pholcidae into five subfamilies, Ninetinae Simon, 1890, Arteminae Simon, 1893, Modisiminae Simon, 1893, Smeringopinae Simon, 1893 and Pholcinae, is confirmed by Huber, Eberle & Dimitrov, 2018.

In synonymy:

Oophora Hentz, 1850 = Spermophora Hentz, 1841 (Thorell, 1869: 102)

Rachus Walckenaer, 1847 = Spermophora Hentz, 1841 (Thorell, 1869: 102)

Simonius Kishida, 1913 = Spermophora Hentz, 1841 (Yaginuma, 1974a: 170)

Gen. Spermophora Hentz, 1841 urn:lsid:nmbe.ch:spidergen:00420
N.B.: considered a senior synonym of Rachus Walckenaer, 1847a: 459 [urn:lsid:nmbe.ch:spidergen:05911] (type Pholcus quadripunctatus Lucas, 1846 = Spermophora senoculata (Dugès, 1836)) and of Oophora Hentz, 1850: 285 [urn:lsid:nmbe.ch:spidergen:05940] (type Spermophora meridionalis Hentz, 1841 = Spermophora senoculata (Dugès, 1836)) by Thorell, 1869: 102; considered a senior synonym of Simonius Kishida, 1913c: 1020 [urn:lsid:nmbe.ch:spidergen:04945] (type S. typicus Kishida, 1913 = Spermophora senoculata (Dugès, 1836)) by Yaginuma, 1974a: 170.

Transferred to other genera:

Spermophora akebona Komatsu, 1961 -- see Belisana

Spermophora amabilis Paik, 1978 -- see Belisana

Spermophora andrei Millot, 1946 -- see Zatavua

Spermophora anhuiensis Xu & Wang, 1984 -- see Belisana

Spermophora ankaranae Millot, 1946 -- see Zatavua

Spermophora baso Roewer, 1963 -- see Savarna

Spermophora caesaris Wunderlich, 1987 -- see Spermophorides

Spermophora combesi Millot, 1946 -- see Paramicromerys

Spermophora cuneata Wunderlich, 1987 -- see Spermophorides

Spermophora dubia Kulczyński, 1911 -- see Panjange

Spermophora elevata Simon, 1874 -- see Spermophorides

Spermophora elongata Yin & Wang, 1981 -- see Cantikus

Spermophora ensifera Tullgren, 1910 -- see Buitinga

Spermophora esperanza Wunderlich, 1987 -- see Spermophorides

Spermophora fagei Millot, 1946 -- see Zatavua

Spermophora faveauxi Lawrence, 1967 -- see Pholcus

Spermophora forcipata Tu, 1994 -- see Belisana

Spermophora fuerteventurensis Wunderlich, 1987 -- see Spermophorides

Spermophora gibbifer Wunderlich, 1987 -- see Spermophorides

Spermophora globosa Tullgren, 1910 -- see Buitinga

Spermophora gomerensis Wunderlich, 1987 -- see Spermophorides

Spermophora hermiguensis Wunderlich, 1987 -- see Spermophorides

Spermophora heterogibbifer Wunderlich, 1987 -- see Spermophorides

Spermophora huberti Senglet, 1973 -- see Spermophorides

Spermophora imerinensis Millot, 1946 -- see Zatavua

Spermophora impudica Millot, 1946 -- see Zatavua

Spermophora junkoae Irie, 1997 -- see Belisana

Spermophora limpida Simon, 1909 -- see Belisana

Spermophora lingua Schmidt, 1956 -- see Metagonia

Spermophora longiventris Simon, 1893 -- see Uthina

Spermophora madagascariensis Fage, 1945 -- see Zatavua

Spermophora mamma Wunderlich, 1987 -- see Spermophorides

Spermophora mammata Senglet, 1973 -- see Spermophorides

Spermophora mediterranea Senglet, 1973 -- see Spermophorides

Spermophora megaceros Millot, 1946 -- see Paramicromerys

Spermophora mercedes Wunderlich, 1987 -- see Spermophorides

Spermophora miser Bristowe, 1952 -- see Savarna

Spermophora nigrescens Berland, 1920 -- see Buitinga

Spermophora petraea Senglet, 1973 -- see Spermophorides

Spermophora pseudomamma Wunderlich, 1987 -- see Spermophorides

Spermophora punctata Millot, 1946 -- see Zatavua

Spermophora rollofoliolata Wang, 1983 -- see Belisana

Spermophora sciakyi Pesarini, 1984 -- see Spermophorides

Spermophora simoni Senglet, 1973 -- see Spermophorides

Spermophora speophila Chamberlin & Ivie, 1938 -- see Anopsicus

Spermophora strinatii Brignoli, 1972 -- see Metagonia

Spermophora tenerifensis Wunderlich, 1987 -- see Spermophorides

Spermophora tesselata Simon, 1901 -- see Savarna

Spermophora tilos Wunderlich, 1987 -- see Spermophorides

Spermophora unicolor Keyserling, 1891 -- see Metagonia

Spermophora valentiana Senglet, 1973 -- see Spermophorides

Spermophora yadongensis Hu, 1985 -- see Belisana

Spermophora yanbaruensis Irie, 2002 -- see Belisana

In synonymy:

Spermophora domestica Yin & Wang, 1981 = Spermophora senoculata Dugès, 1836 (Yin et al., 2012: 175)

Spermophora meridionalis Hentz, 1841 = Spermophora senoculata Dugès, 1836 (Yaginuma, 1974a: 170)

Spermophora quadripunctata (Lucas, 1846) = Spermophora senoculata Dugès, 1836 (Simon, 1866a: 119, sub Pholcus)

Spermophora ressenensis (Drensky, 1929, T from Comaroma) = Spermophora senoculata Dugès, 1836 (Deltshev & Blagoev, 2001: 110)

Spermophora topolia Roewer, 1928 = Spermophora senoculata Dugès, 1836 (Wunderlich, 1992a: 324)

Spermophora typica (Kishida, 1913, T from Simonius) = Spermophora senoculata Dugès, 1836 (Yaginuma, 1974a: 170)

Spermophora abibae Huber, 2014 | | Congo urn:lsid:nmbe.ch:spidersp:046980
Spermophora abibae Huber in Huber, Le Gall & Mavoungou, 2014: 35, f. 84-93 (Dmf) Original description
Spermophora akwamu Huber & Kwapong, 2013 | | Ghana, Gabon urn:lsid:nmbe.ch:spidersp:046604
Spermophora akwamu Huber & Kwapong, 2013: 31, f. 11-13, 77-91 (Dmf) Original description
Spermophora awalai Huber, 2014 | | Cameroon urn:lsid:nmbe.ch:spidersp:046981
Spermophora awalai Huber in Huber, Le Gall & Mavoungou, 2014: 38, f. 5-6, 97-107 (Dmf) Original description
Spermophora berlandi Fage, 1936 | | Kenya urn:lsid:nmbe.ch:spidersp:003898
Spermophora berlandi Fage in Fage & Simon, 1936: 317, f. 10-11 (Dmf) Original description
Spermophora berlandi Huber, 2003b: 601, f. 23, 46, 262-267, 284-285 (mf)
Spermophora bukusu Huber & Warui, 2012 | | Kenya, Uganda urn:lsid:nmbe.ch:spidersp:045540
Spermophora bukusu Huber & Warui, 2012: 32, f. 45, 47-49, 68, 97-101 (Dmf) Original description
Spermophora deelemanae Huber, 2005 | | Indonesia (Ambon) urn:lsid:nmbe.ch:spidersp:039672
Spermophora deelemanae Huber, 2005b: 68, f. 8, 54-56 (Dm) Original description
Spermophora dieke Huber, 2009 | | Guinea urn:lsid:nmbe.ch:spidersp:042721
Spermophora dieke Huber, 2009b: 2514, f. 12, 36-40, 124-127 (Dmf) Original description
Spermophora dumoga Huber, 2005 | | Indonesia (Sulawesi) urn:lsid:nmbe.ch:spidersp:039673
Spermophora dumoga Huber, 2005b: 66, f. 5-6, 16, 43-46, 49 (Dmf) Original description
Spermophora estebani Simon, 1892 | | Philippines urn:lsid:nmbe.ch:spidersp:003904
Spermophora estebani Simon, 1892c: 42 (Df) Original description
Spermophora estebani Huber, 2005b: 70, f. 11, 19-20, 72-80 (f, Dm)
Spermophora falcata Yao & Li, 2013 | | Laos urn:lsid:nmbe.ch:spidersp:046157
Spermophora falcata Yao & Li, 2013: 43, f. 38A-D, 39A-E, 40A-E (Dmf) Original description
Spermophora gordimerae Huber, 2003 | | South Africa urn:lsid:nmbe.ch:spidersp:038460
Spermophora gordimerae Huber, 2003d: 517, f. 27, 49, 232-237 (Dmf) Original description
Spermophora jocquei Huber, 2003 | | Comoros, Mayotte urn:lsid:nmbe.ch:spidersp:038461
Spermophora jocquei Huber, 2003b: 605, f. 288-291 (Dm) Original description
Spermophora kaindi Huber, 2005 | | New Guinea urn:lsid:nmbe.ch:spidersp:039674
Spermophora kaindi Huber, 2005b: 69, f. 10, 69-71 (Dm) Original description
Spermophora kerinci Huber, 2005 | | Indonesia (Sumatra, Bali). Introduced to Britain, Netherlands, Germany, Austria urn:lsid:nmbe.ch:spidersp:039675
Spermophora kerinci Huber, 2005b: 65, f. 1-2, 14, 22-26, 32-38, 47 (Dmf) Original description
Spermophora kerinci Noordijk, 2020: 106, f. 1 (m)
Spermophora kerinci Vaněk, Hörweg & Milasowszky, 2022: 100, f. 1 (f)
Spermophora kirinyaga Huber & Warui, 2012 | | Kenya urn:lsid:nmbe.ch:spidersp:045541
Spermophora kirinyaga Huber & Warui, 2012: 34, f. 10, 50-52, 102-106 (Dmf) Original description
Spermophora kivu Huber, 2003 | | Congo urn:lsid:nmbe.ch:spidersp:038462
Spermophora kivu Huber, 2003b: 600, f. 22, 258-261 (Dm) Original description
Spermophora kyambura Huber & Warui, 2012 | | Cameroon to Uganda urn:lsid:nmbe.ch:spidersp:045542
Spermophora kyambura Huber & Warui, 2012: 36, f. 57-58, 69, 107-110 (Dmf) Original description
Spermophora lambilloni Huber, 2003 | | Comoros urn:lsid:nmbe.ch:spidersp:038463
Spermophora lambilloni Huber, 2003b: 605, f. 25, 49, 274-279, 286-287 (Dmf) Original description
Spermophora luzonica Huber, 2005 | | Philippines urn:lsid:nmbe.ch:spidersp:039676
Spermophora luzonica Huber, 2005b: 71, f. 81-85 (Dmf) Original description
Spermophora maathaiae Huber & Warui, 2012 | | Kenya urn:lsid:nmbe.ch:spidersp:045539
Spermophora maathaiae Huber & Warui, 2012: 29, f. 46, 66-67, 92-96 (Dmf) Original description
Spermophora maculata Keyserling, 1891 | | Brazil urn:lsid:nmbe.ch:spidersp:003915
Spermophora maculata Keyserling, 1891: 177, pl. 6, f. 122 (Df, may be misplaced in this family, per Huber, 2000: 345) Original description
Spermophora maculata Mello-Leitão, 1918c: 110, f. 25 (f)
Spermophora maculata Mello-Leitão, 1922a: 211, f. 5 (Dm)
Spermophora maros Huber, 2005 | | Indonesia (Sulawesi) urn:lsid:nmbe.ch:spidersp:039677
Spermophora maros Huber, 2005b: 67, f. 7, 17, 50-53 (Dmf) Original description
Spermophora masisiwe Huber, 2003 | | Tanzania urn:lsid:nmbe.ch:spidersp:038464
Spermophora masisiwe Huber, 2003b: 599, f. 21, 45, 244-249, 255-257 (Dmf) Original description
Spermophora mau Huber & Warui, 2012 | | Kenya urn:lsid:nmbe.ch:spidersp:045538
Spermophora mau Huber & Warui, 2012: 27, f. 9, 40-44, 87-91 (Dmf) Original description
Spermophora minotaura Berland, 1920 | | East Africa urn:lsid:nmbe.ch:spidersp:003920
Spermophora minotaura Berland, 1920c: 133, f. 154-161 (Dmf) Original description
Spermophora minotaura Berland, 1932b: 276, f. 416 (m)
Spermophora minotaura Huber, 2003b: 602, f. 24, 47-48, 268-273, 280-283 (mf)
Spermophora morogoro Huber, 2003 | | Tanzania urn:lsid:nmbe.ch:spidersp:038465
Spermophora morogoro Huber, 2003b: 594, f. 18, 42, 221-233 (Dmf) Original description
Spermophora palau Huber, 2005 | | Caroline Is. urn:lsid:nmbe.ch:spidersp:039678
Spermophora palau Huber, 2005b: 68, f. 9, 18, 57-68 (Dmf) Original description
Spermophora palau Beatty, Berry & Huber, 2008: 23, f. 10-11, 36 (mf)
Spermophora paluma Huber, 2001 | | Australia (Queensland) urn:lsid:nmbe.ch:spidersp:036973
Spermophora paluma Huber, 2001: 126, f. 406-415 (Dmf) Original description
Spermophora pembai Huber, 2003 | | South Africa urn:lsid:nmbe.ch:spidersp:038466
Spermophora pembai Huber, 2003d: 519, f. 28, 252-255 (Dm) Original description
Spermophora peninsulae Lawrence, 1964 | | South Africa urn:lsid:nmbe.ch:spidersp:003923
Spermophora peninsulae Lawrence, 1964: 65, f. 9-14 (Dmf) Original description
Spermophora peninsulae Huber, 2003d: 519, f. 29, 50, 238-243, 248-251 (mf)
Spermophora persica Senglet, 2008 | | Iran urn:lsid:nmbe.ch:spidersp:042173
Spermophora persica Senglet, 2008: 372, f. 107-117 (Dmf) Original description
Spermophora ranomafana Huber, 2003 | | Madagascar urn:lsid:nmbe.ch:spidersp:039240
Spermophora ranomafana Huber, 2003a: 304, f. 218-220, 224-237 (Dmf) Original description
Spermophora sangarawe Huber, 2003 | | Tanzania urn:lsid:nmbe.ch:spidersp:038467
Spermophora sangarawe Huber, 2003b: 596, f. 19, 43, 234-237 (Dmf) Original description
Spermophora schoemanae Huber, 2003 | | South Africa urn:lsid:nmbe.ch:spidersp:038468
Spermophora schoemanae Huber, 2003d: 516, f. 26, 48, 226-231, 244-247 (Dmf) Original description
Spermophora senoculata (Dugès, 1836) | | Middle East. Introduced to USA, Jamaica, southern Europe, China, Korea, Japan urn:lsid:nmbe.ch:spidersp:003927
Pholcus senoculatus Dugès, 1836a: 160 (D) Original description
Spermophora meridionalis Hentz, 1841a: 116, pl. 17, f. 9 (Df) [urn:lsid:nmbe.ch:spidersp:057483]
Pholcus quadri-punctatus Lucas, 1846: 239, pl. 15, f. 2 (Df) [urn:lsid:nmbe.ch:spidersp:057482]
Rachus quadripunctatus Walckenaer, 1847a: 459
Oophora meridionalis Hentz, 1850: 286, pl. 10, f. 9 (f)
Pholcus sex-oculatus Simon, 1866a: 119, pl. 2, f. 8-9 (lapsus, S of Pholcus quadripunctatus)
Spermophora meridionalis Emerton, 1882: 31, pl. 6, f. 3 (f)
Spermophora senoculata Chyzer & Kulczyński, 1891: 150
Spermophora meridionalis Petrunkevitch, 1910: 208, pl. 21, f. 5 (Dm)
Simonius typicus Kishida, 1913f: 1020 (Df) [urn:lsid:nmbe.ch:spidersp:057485]
Spermophora senoculata Simon, 1914: 238, f. 492 (mf)
Spermophora topolia Roewer, 1928b: 120, pl. 2, f. 35-36 (Dm) [urn:lsid:nmbe.ch:spidersp:057484]
Comaroma ressenensis Drensky, 1929: 36, 70, pl. 3, f. 9-10 (Df) [urn:lsid:nmbe.ch:spidersp:009098]
Comaroma resnensis Drensky, 1935: 105 (lapsus, Dm)
Spermophora meridionalis Gertsch, 1939a: 1, f. 3 (m)
Spermophora topolia Hadjissarantos, 1940: 48, f. 15 (Df)
Simonius typicus Shinkai, 1969: 11, f. 5 (f)
Spermophora senoculata Dresco & Hubert, 1969: 170, f. 1-5 (mf)
Spermophora senoculata Yaginuma, 1974a: 170 (S of Simonius typicus and Spermophora meridionalis)
Spermophora senoculata Yaginuma, 1975b: 23, f. A-D (m)
Spermophora senoculata Miller & Žitňanská, 1976a: 81, pl. I, f. 1-3 (mf)
Spermophora domestica Yin & Wang, 1981c: 378, f. 1A-F (Dmf) [urn:lsid:nmbe.ch:spidersp:003900]
Spermophora senoculata Zhu, 1983: 17 (S of Spermophora domestica)
Spermophora senoculata Yaginuma, 1986a: 29, f. 17.7 (m)
Spermophora domestica Song, 1987: 111, f. 74 (mf; regarded as valid, contra Zhu, 1983: 17)
Spermophora senoculata Chikuni, 1989b: 27, f. 2 (mf)
Spermophora domestica Chen & Zhang, 1991: 76, f. 68.1-4 (mf)
Spermophora senoculata Heimer & Nentwig, 1991: 40, f. 84 (mf)
Spermophora senoculata Wunderlich, 1992a: 324 (S of Spermophora topolia)
Spermophora senoculata Paik, 1996b: 23, f. 1-7 (m)
Spermophora senoculata Song, Zhu & Chen, 1999: 65, f. 26B'-D' (mf)
Spermophora senoculata Szinetár, Kenyeres & Kovács, 1999: 162, f. 2A-C (f)
Spermophora senoculata Huber, 2000: 11, f. 11, 24-25, 70-71, 122, 162, 193 (m)
Spermophora senoculata Senglet, 2001: 50, f. 29-30, 46-49 (mf)
Spermophora senoculata Deltshev & Blagoev, 2001: 110 (S of Comaroma ressenensis)
Spermophora senoculata Van Keer & Van Keer, 2001: 83, f. 5.1-5 (mf)
Spermophora senoculata Namkung, 2002: 50, f. 5.9a-c (mf)
Spermophora senoculata Kim & Cho, 2002: 60, f. 13-18 (m)
Spermophora senoculata Huber, 2002b: 105, f. 1-13 (mf)
Spermophora senoculata Namkung, 2003: 52, f. 5.9a-c (mf)
Spermophora senoculata Lee & Kim, 2003: 117, f. 100-106 (mf)
Spermophora senoculata Deltshev, 2003: 137, f. 13-16 (f)
Spermophora senoculata Trotta, 2005: 171, f. 379-380 (mf)
Spermophora senoculata Jocqué & Dippenaar-Schoeman, 2006: 206, f. 80j-l (mf)
Spermophora senoculata Kovács, Szinetár & Eichardt, 2008: 125, f. 1 (m)
Spermophora senoculata Senglet, 2008: 370, f. 106 (f)
Spermophora senoculata Irie, 2009: 111, f. 46-47 (m)
Spermophora senoculata Le Peru, 2011: 157, f. 201-202 (mf)
Spermophora senoculata Zhu & Zhang, 2011: 51, f. 23A-D (mf, probable S of Spermophora domestica)
Spermophora senoculata Yin et al., 2012: 175, f. 37a-g (mf, S of Spermophora domestica; confirming Zhu, 1983: 17)
Spermophora senoculata Schifani et al., 2019: 6, f. 1 (mf)
Spermophora senoculatoides Senglet, 2008 | | Iran urn:lsid:nmbe.ch:spidersp:042172
Spermophora senoculatoides Senglet, 2008: 370, f. 96-105 (Dmf) Original description
Spermophora sumbawa Huber, 2005 | | Indonesia (Sunda Is.) urn:lsid:nmbe.ch:spidersp:039679
Spermophora sumbawa Huber, 2005b: 71, f. 12-13, 21, 86-103 (Dmf) Original description
Spermophora suurbraak Huber, 2003 | | South Africa urn:lsid:nmbe.ch:spidersp:038469
Spermophora suurbraak Huber, 2003d: 521, f. 30, 256-258 (Dm) Original description
Spermophora thorelli Roewer, 1942 | | Myanmar urn:lsid:nmbe.ch:spidersp:003930
Spermophora maculata Thorell, 1898: 276 (Df; preoccupied by Keyserling, 1891)
Spermophora thorelli Roewer, 1942a: 346 (replacement name) Original description
Spermophora thorelli Yao, Tavano & Li, 2013: 307, f. 5A-B (f)
Spermophora tonkoui Huber, 2003 | | Guinea, Ivory Coast urn:lsid:nmbe.ch:spidersp:038470
Spermophora tonkoui Huber, 2003b: 607, f. 50, 292-298 (Dmf) Original description
Spermophora tumbang Huber, 2005 | | Borneo urn:lsid:nmbe.ch:spidersp:039680
Spermophora tumbang Huber, 2005b: 65, f. 3-4, 15, 27-31, 39-42, 48 (Dmf) Original description
Spermophora usambara Huber, 2003 | | Tanzania urn:lsid:nmbe.ch:spidersp:038471
Spermophora usambara Huber, 2003b: 598, f. 20, 44, 238-243, 250-254 (Dmf) Original description
Spermophora vyvato Huber, 2003 | | Madagascar urn:lsid:nmbe.ch:spidersp:038472
Spermophora vyvato Huber, 2003a: 306, f. 221-223, 238-243 (Dmf) Original description
Spermophora yao Huber, 2001 | | Australia (Queensland) urn:lsid:nmbe.ch:spidersp:036974
Spermophora yao Huber, 2001: 126, f. 392-405 (Dmf) Original description
Spermophora ziama Huber & Kwapong, 2013 | | Guinea urn:lsid:nmbe.ch:spidersp:046605
Spermophora ziama Huber & Kwapong, 2013: 34, f. 92-105 (Dmf) Original description

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