World Spider Catalog

Pholcidae C. L. Koch, 1850

Family | ACCEPTED

N.B.: for comments on the monophyly and relationships of many genera see Huber, 1998d and Huber, 2000; the division of Pholcidae into five subfamilies, Ninetinae Simon, 1890, Arteminae Simon, 1893, Modisiminae Simon, 1893, Smeringopinae Simon, 1893 and Pholcinae, is confirmed by Huber, Eberle & Dimitrov, 2018; two additional subfamilies are described by Meng et al., 2026: 99, 113 (Caipirinae Meng, Carvalho, Podsiadlowski & Huber, 2026 and Physocyclinae Meng, Carvalho, Podsiadlowski & Huber, 2026).

In synonymy:

Coroia González-Sponga, 2005 = Artema Walckenaer, 1837 (Huber, Colmenares & Ramírez, 2014: 416)

Pholciella Roewer, 1960 = Artema Walckenaer, 1837 (Huber, 2009a: 67)

Gen. Artema Walckenaer, 1837 urn:lsid:nmbe.ch:spidergen:00370

type Artema atlanta Walckenaer, 1837; gender feminine

N.B.: considered a senior synonym of Pholciella Roewer, 1960b: 42 [urn:lsid:nmbe.ch:spidergen:00409] (type P. ziaretana Roewer, 1960) by Huber, 2009a: 67 and of Coroia González-Sponga, 2005: 102 [urn:lsid:nmbe.ch:spidergen:04197] (type C. magna González-Sponga, 2005 = Artema atlanta Walckenaer, 1837) by Huber, Colmenares & Ramírez, 2014: 416.

In synonymy:

Artema magna (González-Sponga, 2005, T from Coroia = Artema) = Artema atlanta Walckenaer, 1837 (Huber, Colmenares & Ramírez, 2014: 417)

Artema mauriciana Walckenaer, 1837 = Artema atlanta Walckenaer, 1837 (Caporiacco, 1948a: 626, after F. O. Pickard-Cambridge, 1902a: 366)

Artema sexsignata (Franganillo, 1926, T from Crossopriza) = Artema atlanta Walckenaer, 1837 (Pérez, 1996: 432)

Artema sisyphoides (Doleschall, 1857) = Artema atlanta Walckenaer, 1837 (Millot, 1941: 3, sub A. mauriciana)

Artema atlanta Walckenaer, 1837 | | North Africa, Middle East. Introduced to both Americas, St. Helena, tropical Africa, Britain, Belgium, Greece, Turkey, tropical Asia, China, Japan, Taiwan, Australia, Pacific Is. urn:lsid:nmbe.ch:spidersp:003327 TYPE SPECIES

Artema atlanta Walckenaer, 1837: 656 (Dmf) Original description

Artema mauriciana Walckenaer, 1837: 657, pl. 15, f. 1 (Dmf) [urn:lsid:nmbe.ch:spidersp:056687]

Pholcus sisyphoides Doleschall, 1857: 408 (D) [urn:lsid:nmbe.ch:spidersp:056689]

Artema convexa Blackwall, 1858b: 332 (Dj)

Pholcus borbonicus Vinson, 1863: 132, pl. 3, f. 4 (Df)

Artema mauricia Vinson, 1863: 141, 307 (Dm)

Artema convexa Blackwall, 1866: 459 (Df)

Pholcus rotundatus Karsch, 1879h: 106 (Df)

Artema mauricia Simon, 1885h: 19

Artema sisyphoides Karsch, 1892: 276

Artema mauricia Simon, 1893a: 463, f. 441, 451-454 (mf)

Artema atlanta Pocock, 1900a: 239, f. 81 (Df)

Artema mauriciana Kulczyński, 1901a: 20, pl. 1, f. 15 (f)

Artema atlanta F. O. Pickard-Cambridge, 1902a: 366, pl. 34, f. 15-16 (mf)

Artema mauriciana Tullgren, 1910: 119, pl. 2, f. 35 (f)

Crossopriza sex-signata Franganillo, 1926a: 49 (Dmf) [urn:lsid:nmbe.ch:spidersp:056688]

Artema atlanta Petrunkevitch, 1929b: 119, f. 102-104 (mf)

Artema atlanta Dyal, 1935: 170, pl. 15, f. 97-103 (mf)

Artema mauriciana Millot, 1941: 3, f. 1F (m, S of Artema kochi and A. sisyphoides; S of A. kochi rejected by Aharon, Huber & Gavish-Regev, 2017: 26; other figs. see A. bunkpurugu)

Artema mauriciana Millot, 1946a: 129, f. 1 (f)

Artema atlanta Caporiacco, 1948a: 626 (S of Artema mauriciana)

Artema mauritiana Bonnet, 1955: 746 (unjustified emendation)

Artema atlanta Chrysanthus, 1967b: 92, f. 15-19 (mf)

Artema atlanta Tikader & Biswas, 1981: 18, f. 12 (f)

Artema atlanta Brignoli, 1981c: 92, f. 1-7 (mf)

Artema atlanta Pérez, 1996: 432 (S of Crossopriza sexsignata)

Artema atlanta Huber, 2000: 12, f. 12-13, 48, 56-57, 99, 121, 145, 169, 195 (m)

Artema atlanta Van Keer & Van Keer, 2001: 83, f. 1.1-7 (mf)

Artema atlanta Saaristo, 2001b: 15, f. 16-22 (mf)

Artema atlanta Lee, 2005: 7, f. a-d (m)

Coroia magna González-Sponga, 2005: 102, pl. 2, f. 1-11 (Dmf) [urn:lsid:nmbe.ch:spidersp:047256]

Tibiosa coreana González-Sponga, 2006: 17, pl. 3, f. 8-9 (f; misidentified per Huber, 2009a: 65)

Artema atlanta Beatty, Berry & Huber, 2008: 3, f. 15-16, 42-43 (mf)

Artema atlanta Colmenares-García, 2008: 87, f. 1A-C (m)

Artema atlanta Irie, 2009: 106, f. 1-2 (f)

Artema atlanta Saaristo, 2010: 159, f. 25.1-5 (mf)

Artema atlanta Gao & Li, 2010b: 11, f. 1A-D, 2A-D, 3A-C (mf)

Artema atlanta Le Peru, 2011: 152, f. 179 (mf)

Artema atlanta Tong, 2013: 55, f. 71A-D, 72A-D, 73A-C (mf)

Artema atlanta Huber, Colmenares & Ramírez, 2014: 417 (S of Coroia magna)

Artema atlanta Aharon, Huber & Gavish-Regev, 2017: 8, f. 3, 15-34, 201, 208 (mf)

Artema atlanta Al-Abbad, Al-Mayah & Al-Abody, 2019: 499, f. 2a-d, 3a-e, 4a-b, 5a-b (mf)

Artema bahla Huber, 2019 | | Oman urn:lsid:nmbe.ch:spidersp:050889

Artema bahla Huber in Huber & Carvalho, 2019: 6, f. 1-2, 7-8, 13-14, 19-20, 25, 30, 33-34 (Dmf) Original description

Artema bunkpurugu Huber & Kwapong, 2013 | | West Africa urn:lsid:nmbe.ch:spidersp:046602

Artema mauriciana Millot, 1941: 3, f. 1A-E, G-I (mf; misidentified per Huber & Kwapong, 2013: 24)

Artema bunkpurugu Huber & Kwapong, 2013: 24, f. 14-15, 48-62 (Dmf) Original description

Artema bunkpurugu Aharon, Huber & Gavish-Regev, 2017: 50, f. 202, 209 (f)

Artema dhofar Huber, 2019 | | Oman urn:lsid:nmbe.ch:spidersp:050891

Artema dhofar Huber in Huber & Carvalho, 2019: 14, f. 5-6, 11-12, 17-18, 23-24, 27-28, 32, 37-38 (Dmf) Original description

Artema doriae (Thorell, 1881) | | Turkey, Azerbaijan, Israel, United Arab Emirates, Iraq, Iran, Afghanistan. Introduced to Japan urn:lsid:nmbe.ch:spidersp:003328

Pholcus borbonicus Simon, 1877b: 87 (from Iran; misidentified per Thorell, 1881: 180)

Pholcus doriae Thorell, 1881: 180 (Df, new name for Pholcus borbonicus sensu Simon, 1877; doriai is an unjustified emendation, see ICZN Article 31.1.1) Original description

Artema doriai Brignoli, 1981c: 92 (removed from S of Artema mauriciana [rejecting Roewer, 1942a: 333], possible senior S of A. transcaspica and A. magna)

Artema doriai Tabrizi, Rad & Hedayati, 2014: 35, f. 1M-N, 3L-M (f)

Artema doriae Aharon, Huber & Gavish-Regev, 2017: 30, f. 90-120, 205, 212 (Dm, f)

Artema doriae Al-Khazali & Najim, 2018: 179, pl. 1A-D, 2A-D, 3 (mf)

Artema doriae Nuruyeva, Snegovaya & Aliyeva, 2026: 136, f. 9 (f)

Artema ghubrat Huber, 2019 | | Oman urn:lsid:nmbe.ch:spidersp:050890

Artema sp. a Aharon, Huber & Gavish-Regev, 2017: 44, f. 170 (f)

Artema ghubrat Huber in Huber & Carvalho, 2019: 11, f. 3-4, 9-10, 15-16, 21-22, 26, 29, 31, 35-36 (Dmf) Original description

Artema kochi Kulczyński, 1901 | | Eritrea, Ethiopia, Sudan, Yemen, Egypt? urn:lsid:nmbe.ch:spidersp:049789

Pholcus borbonicus L. Koch, 1875b: 25, pl. 3, f. 1 (f; misidentified per Aharon, Huber & Gavish-Regev, 2017: 26)

Artema kochii Kulczyński, 1901a: 20, pl. 1, f. 14 (Df) Original description

Artema kochi Siyam, Dunlop & Jäger, 2017: 319, f. 1-3 (m)

Artema kochi Aharon, Huber & Gavish-Regev, 2017: 26, f. 76-89, 204, 211 (mf, removed from S of Artema atlanta, rejecting Simon, 1908a: 426 and Millot, 1941: 3, both sub A. mauriciana)

Artema magna Roewer, 1960 | | Afghanistan, Pakistan? urn:lsid:nmbe.ch:spidersp:003329

Artema magna Roewer, 1960b: 43, f. 14a-h (Df) Original description

Artema magna Aharon, Huber & Gavish-Regev, 2017: 41, f. 155-169 (Dm, f) [see also note at A. ziaretana]

Artema martensi Huber, 2021 | | Morocco urn:lsid:nmbe.ch:spidersp:053252

Artema martensi Huber, 2021: 326, f. 1-24 (Dmf) Original description

Artema nephilit Aharon, Huber & Gavish-Regev, 2017 | | Greece, Turkey, Cyprus, Israel, Jordan, Yemen?, United Arab Emirates? urn:lsid:nmbe.ch:spidersp:049788

Artema nephilit Aharon, Huber & Gavish-Regev, 2017: 15, f. 4-6, 9-14, 35-75, 203, 210 (Dmf) Original description

Artema transcaspica Spassky, 1934 | | Iran, Tajikistan, Turkmenistan, Uzbekistan urn:lsid:nmbe.ch:spidersp:003330

Artema transcaspica Spassky, 1934a: 369, f. 8-10 (Dmf) Original description

Artema transcaspica Aharon, Huber & Gavish-Regev, 2017: 35, f. 121-154, 206, 213 (mf)

Artema transcaspica Zamani et al., 2020: 585, f. 10A-B (m)

Artema ziaretana (Roewer, 1960) | j | Afghanistan urn:lsid:nmbe.ch:spidersp:003658

Pholciella ziaretana Roewer, 1960b: 42, f. 13a-d (Df) Original description

Artema ziaretana Huber, 2009a: 67 (T from Pholciella, type is j not f)

Artema ziaretana Aharon, Huber & Gavish-Regev, 2017: 44 (might be a synonym of A. magna)