Pycnothelidae Chamberlin, 1917

Family |   ACCEPTED
N.B.: downgraded to a subfamily of Nemesiidae by Raven, 1985a: 79 and revalidated by Opatova et al., 2020: 701; redefined and transfer of 12 genera from Nemesiidae by Montes de Oca et al., 2022: 11; furthermore Montes de Oca et al., 2022: 14 define five subfamilies: Pycnothelinae, Diplothelopsinae Schiapelli & Gerschman, 1967, Pionothelinae Indicatti, Montes de Oca, Opatova, Almeida, Pérez-Miles & Bond, 2022 (new subfamily), Prorachiinae Indicatti, Montes de Oca, Opatova, Almeida, Pérez-Miles, Bond, 2022 (new subfamily) and Pselligminae Mello-Leitão, 1923 (elevated from tribe).

In synonymy:

Diplothelopsis Tullgren, 1905 = Lycinus Thorell, 1894 (Montes de Oca et al., 2022: 12)

Levina Zapfe, 1963 = Lycinus Thorell, 1894 (Schiapelli & Gerschman, 1973c: 291)

Parachubutia Mello-Leitão, 1940 = Lycinus Thorell, 1894 (Schiapelli & Gerschman, 1967b: 57)

Gen. Lycinus Thorell, 1894 urn:lsid:nmbe.ch:spidergen:00121
N.B.: transferred from the Nemesiidae to the Pycnothelidae by Montes de Oca et al., 2022: 11; removed from the synonymy of Mygaloides Nicolet, 1849 (regarded as a nomen dubium) by Goloboff, 1995: 148, contra Raven, 1985a: 98; considered a senior synonym of Parachubutia Mello-Leitão, 1940f: 233 [urn:lsid:nmbe.ch:spidergen:05011] (type P. gajardoi Mello-Leitão, 1940) by Schiapelli & Gerschman, 1967b: 55; considered a senior synonym of Levina Zapfe, 1963: 125 [urn:lsid:nmbe.ch:spidergen:05010] (type L. epipiptusa Zapfe, 1963) by Schiapelli & Gerschman, 1973c: 291; considered a senior synonym of Diplothelopsis Tullgren, 1905: 4 [urn:lsid:nmbe.ch:spidergen:00113] (type D. ornata Tullgren, 1905) by Montes de Oca et al., 2022: 12.

In synonymy:

Lycinus canescens (Mello-Leitão, 1939) = Lycinus bonariensis Mello-Leitão, 1938 (Gerschman & Schiapelli, 1966a: 387, sub Diplothelopsis)

Lycinus decarloi (Gerschman & Schiapelli, 1948) = Lycinus ornatus Tullgren, 1905 (Gerschman & Schiapelli, 1966a: 385, sub Diplothelopsis)

Nomina dubia:

Lycinus hastatus (Mello-Leitão, 1923a: 112, f. 25-27, f, Brazil, originally in Diplothelopsis) -- Goloboff, 1995: 164, N.B.: m of Bücherl, 1957 is probably a misidentified Neodiplothele

Lycinus secundus Mello-Leitão, 1941d: 105, pl. 2, f. 5 (j, Argentina) -- Goloboff, 1995: 149

Lycinus bonariensis (Mello-Leitão, 1938) | | Argentina urn:lsid:nmbe.ch:spidersp:001117
Diplothelopsis bonariensis Mello-Leitão, 1938a: 89, f. 1 (Dmf) Original description
Diplothelopsis canescens Mello-Leitão, 1939e: 47, f. 4 (Dm) [urn:lsid:nmbe.ch:spidersp:041390]
Diplothelopsis bonariensis Gerschman & Schiapelli, 1966a: 387, f. 1-4, 9-11, 14-17 (mf, S of Diplothelopsis canescens)
Diplothelopsis bonariensis Goloboff, 1995: 166, f. 13, 45, 132A-B (mf)
Diplothelopsis bonariensis Ferretti et al., 2010: 21, f. 28 (m)
Lycinus bonariensis Montes de Oca et al., 2022: 13 (T from Diplothelopsis)
Lycinus caldera Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001163
Lycinus caldera Goloboff, 1995: 154, f. 125A-D (Dmf) Original description
Lycinus choros Lucas & Indicatti, 2010 | | Chile urn:lsid:nmbe.ch:spidersp:043514
Lycinus choros Lucas & Indicatti, 2010: 428, f. 14-19 (Dm) Original description
Lycinus domeyko Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001164
Lycinus domeyko Goloboff, 1995: 162, f. 130 (Dm) Original description
Lycinus epipiptus (Zapfe, 1963) | | Chile, Argentina urn:lsid:nmbe.ch:spidersp:001165
Levina epipiptusa Zapfe, 1963: 126, f. 1-4 (Dm) Original description
Lycinus epipiptus Goloboff, 1995: 157, f. 127A-E (removed m from S of L. gajardoi contra Schiapelli & Gerschman, 1973c: 291, Df)
Lycinus epipiptus Ferretti, 2015a: 10, f. 7A-G (m)
Lycinus frayjorge Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001166
Lycinus frayjorge Goloboff, 1995: 156, f. 126 (Df) Original description
Lycinus gajardoi (Mello-Leitão, 1940) | | Chile urn:lsid:nmbe.ch:spidersp:001167
Parachubutia gajardoi Mello-Leitão, 1940f: 233 (Dm) Original description
Lycinus gajardoi Schiapelli & Gerschman, 1967b: 57, f. 16-19 (m)
Lycinus gajardoi Schiapelli & Gerschman, 1973c: 291 (S, rejected)
Lycinus gajardoi Legendre & Calderón, 1984: 1045, pl. XIII, f. 1-14 (m, Df)
Lycinus gajardoi Goloboff, 1995: 152, f. 7, 16, 21, 124A-E (m, Df)
Lycinus lagigliai Ferretti, 2015 | | Argentina urn:lsid:nmbe.ch:spidersp:047743
Lycinus lagigliai Ferretti, 2015a: 4, f. 2A-G, 3A-D, 5B (Dmf) Original description
Lycinus longipes Thorell, 1894 | | Argentina urn:lsid:nmbe.ch:spidersp:001168
Lycinus longipes Thorell, 1894: 37 (Dm) Original description
Lycinus longipes Carbonell, 1923b: 108 (Df)
Lycinus longipes Schiapelli & Gerschman, 1942: 321, f. 7-8 (m)
Lycinus longipes Bücherl, 1957: 405, f. 94 (m)
Lycinus longipes Schiapelli & Gerschman, 1967b: 56, f. 14-15, 20-22 (m)
Lycinus longipes Legendre & Calderón, 1984: 1048, pl. XIV, f. 1-9 (m)
Mygaloides longipes Raven, 1985a: 98, f. 97-102 (m)
Lycinus longipes Goloboff, 1995: 150, f. 31-32, 123A-F, 134 (m, Df)
Lycinus longipes Ferretti, 2015a: 11, f. 9A-G, 10B (m)
Lycinus nevadoensis Ferretti, 2015 | | Argentina urn:lsid:nmbe.ch:spidersp:047744
Lycinus nevadoensis Ferretti, 2015a: 7, f. 6A-C, 8A-G (Dm) Original description
Lycinus ornatus (Tullgren, 1905) | | Argentina urn:lsid:nmbe.ch:spidersp:001118
Diplothelopsis ornatus Tullgren, 1905: 5, pl. 1, f. 1 (Df) Original description
Diplothelopsis decarloi Gerschman & Schiapelli, 1948: 4, f. 4-7 (Dm) [urn:lsid:nmbe.ch:spidersp:041391]
Diplothelopsis decarloi Bücherl, 1957: 387, f. 22 (m)
Diplothelopsis ornata Gerschman & Schiapelli, 1966a: 385, f. 5-8, 12-13 (mf, S of Diplothelopsis decarloi)
Diplothelopsis ornatus Schiapelli & Gerschman, 1967b: 62, f. 23-25, 27-33 (mf)
Diplothelopsis ornatus Raven, 1985a: 97, f. 92-96
Diplothelopsis ornata Goloboff, 1995: 164, f. 131A-C (mf)
Diplothelopsis ornata Grismado & Goloboff, 2014c: 113, f. 9 (m)
Lycinus ornatus Montes de Oca et al., 2022: 13 (T from Diplothelopsis)
Lycinus portoseguro Lucas & Indicatti, 2010 | | Brazil urn:lsid:nmbe.ch:spidersp:043513
Lycinus portoseguro Lucas & Indicatti, 2010: 425, f. 1-13 (Dmf) Original description
Lycinus quilicura Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001169
Lycinus quilicura Goloboff, 1995: 159, f. 128 (Dm) Original description
Lycinus tofo Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001170
Lycinus tofo Goloboff, 1995: 160, f. 129A-D (Dmf) Original description

Sponsors