Pycnothelidae Chamberlin, 1917

Family |   ACCEPTED
N.B.: downgraded to a subfamily of Nemesiidae by Raven, 1985a: 79 and revalidated by Opatova et al., 2020: 701; redefined and transfer of 12 genera from Nemesiidae by Montes de Oca et al., 2022: 11; furthermore Montes de Oca et al., 2022: 14 define five subfamilies: Pycnothelinae, Diplothelopsinae Schiapelli & Gerschman, 1967, Pionothelinae Indicatti, Montes de Oca, Opatova, Almeida, Pérez-Miles & Bond, 2022 (new subfamily), Prorachiinae Indicatti, Montes de Oca, Opatova, Almeida, Pérez-Miles, Bond, 2022 (new subfamily) and Pselligminae Mello-Leitão, 1923 (elevated from tribe).

In synonymy:

Chubutia Mello-Leitão, 1940 = Acanthogonatus Karsch, 1880 (Gerschman & Schiapelli, 1970a: 152, sub Tryssothele)

Tryssothele Simon, 1902 = Acanthogonatus Karsch, 1880 (Raven, 1985a: 83)

Gen. Acanthogonatus Karsch, 1880 urn:lsid:nmbe.ch:spidergen:00104
N.B.: transferred from the Barychelidae to the Nemesiidae by Raven, 1985a: 83 and to the Pycnothelidae by Opatova et al., 2020: 701; considered a senior synonym of Tryssothele Simon, 1902h: 7 [urn:lsid:nmbe.ch:spidergen:05015] (type T. fuegiana Simon, 1902h: 7) by Raven, 1985a: 83; Tryssothele considered a senior synonym of Chubutia Mello-Leitão, 1940b: 3 [urn:lsid:nmbe.ch:spidergen:05007] (type C. notata Mello-Leitão, 1940) by Gerschman & Schiapelli, 1970a: 152; Acanthogonatus is most likely polyphyletic per Opatova et al., 2020: 701.

Transferred to other genera:

Acanthogonatus antillensis (F. O. Pickard-Cambridge, 1898) -- see Fufius

Acanthogonatus argentina (Simon, 1897) -- see Stenoterommata

In synonymy:

Acanthogonatus australis (Chamberlin, 1917) = Acanthogonatus pissii Simon, 1889 (Gerschman & Schiapelli, 1968: 27, sub Tryssothele)

Acanthogonatus guttulatus (Simon, 1886, T from Stenoterommata) = Acanthogonatus subcalpeianus Nicolet, 1849 (Goloboff, 1995: 78)

Acanthogonatus latastei (Simon, 1902) = Acanthogonatus pissii Simon, 1889 (Gerschman & Schiapelli, 1968: 27, sub Tryssothele)

Acanthogonatus segnis (Simon, 1886) = Acanthogonatus francki Karsch, 1880 (Goloboff, 1995: 104)

Acanthogonatus venosus (Mello-Leitão, 1936, T from Aphantopelma=Thalerommata) = Acanthogonatus pissii Simon, 1889 (Goloboff, 1995: 91)

Nomen dubium:

Acanthogonatus gounellei (Simon, 1886c: 573, j, Brazil, originally in Stenoterommata) -- Goloboff, 1995: 76

Acanthogonatus alegre Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001021
Acanthogonatus alegre Goloboff, 1995: 103, f. 94A-B (Df) Original description
Acanthogonatus birabeni Goloboff, 1995 | | Argentina urn:lsid:nmbe.ch:spidersp:001022
Acanthogonatus birabeni Goloboff, 1995: 116, f. 102A-B (Dm) Original description
Acanthogonatus brunneus (Nicolet, 1849) | | Chile urn:lsid:nmbe.ch:spidersp:001023
Mygale brunnea Nicolet, 1849: 336, pl. 1, f. 5 (Df) Original description
Brachythele brunnea Simon, 1889a: 221
Acanthogonatus brunneus Goloboff, 1995: 127, f. 113A-D (f)
Acanthogonatus campanae (Legendre & Calderón, 1984) | | Chile urn:lsid:nmbe.ch:spidersp:001024
Tryssothele subcalpetana Gerschman & Schiapelli, 1968: 29, f. 9-13 (m, misidentified)
Tryssothele campanae Legendre & Calderón, 1984: 1052, pl. XVII, f. 1-14 (Dmf) Original description
Acanthogonatus subcalpeiana Raven, 1985a: 83, f. 35-39 (m, misidentified)
Acanthogonatus campanae Goloboff, 1995: 100, f. 92A-E, 93A-B (mf)
Acanthogonatus centralis Goloboff, 1995 | | Argentina urn:lsid:nmbe.ch:spidersp:001025
Acanthogonatus centralis Goloboff, 1995: 96, f. 9, 77-78, 89A-F, 90 (Dmf) Original description
Acanthogonatus centralis Ferretti et al., 2010: 21, f. 29-30 (m)
Acanthogonatus centralis Grismado & Goloboff, 2014c: 113, f. 10-11 (m)
Acanthogonatus chilechico Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001026
Acanthogonatus chilechico Goloboff, 1995: 115, f. 101A-B (Dm) Original description
Acanthogonatus confusus Goloboff, 1995 | | Chile, Argentina urn:lsid:nmbe.ch:spidersp:001027
Acanthogonatus confusus Goloboff, 1995: 80, f. 62, 80A-G (Dmf) Original description
Acanthogonatus ericae Indicatti, Lucas, Ott & Brescovit, 2008 | | Brazil urn:lsid:nmbe.ch:spidersp:042101
Acanthogonatus ericae Indicatti et al., 2008b: 534, f. 15-26, 35-39 (Dmf) Original description
Acanthogonatus francki Karsch, 1880 | | Chile urn:lsid:nmbe.ch:spidersp:001028
Acanthogonatus franckii Karsch, 1880c: 392 (Df) Original description
Stenoterommata segne Simon, 1886h: 574 (Df) [urn:lsid:nmbe.ch:spidersp:039276]
Acanthognatus franckii Petrunkevitch, 1911: 42 (lapsus)
Stenoteromata segne Schiapelli & Gerschman, 1962b: 72, pl. III, f. 2 (f)
Acanthogonatus frankii Raven, 1985a: 83, f. 31-34
Acanthogonatus segne Raven, 1985a: 107 (T from Stenoterommata)
Acanthogonatus franki Goloboff, 1995: 104, f. 95A-E (f, Dm, S of Acanthogonatus segne)
Acanthogonatus fuegianus (Simon, 1902) | | Chile, Argentina urn:lsid:nmbe.ch:spidersp:001029
Tryssothele fuegiana Simon, 1902h: 8 (Dmf) Original description
Tryssothele fuegiana Gerschman & Schiapelli, 1968: 22, f. 6-8, 14-17 (mf, probably misplaced)
Acanthogonatus fuegianus Goloboff, 1995: 111, f. 60, 99A-E (mf)
Acanthogonatus hualpen Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001030
Acanthogonatus hualpen Goloboff, 1995: 121, f. 107A-D (Dmf) Original description
Acanthogonatus huaquen Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001031
Acanthogonatus huaquen Goloboff, 1995: 83, f. 59, 81A-E, 82 (Dmf) Original description
Acanthogonatus incursus (Chamberlin, 1916) | | Peru urn:lsid:nmbe.ch:spidersp:001032
Brachythele incursus Chamberlin, 1916: 205, pl. 7, f. 12 (Df) Original description
Bolostromus incursus Raven, 1985a: 160 (Tf from Brachythele)
Acanthogonatus incursus Goloboff, 1995: 93, f. 55, 88A-D (Tf from Bolostromus)
Acanthogonatus juncal Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001033
Acanthogonatus juncal Goloboff, 1995: 87, f. 84A-C (Df) Original description
Acanthogonatus messii Signorotto & Ferretti, 2023 | | Argentina urn:lsid:nmbe.ch:spidersp:059223
Acanthogonatus messii Signorotto & Ferretti in Signorotto, Mancini & Ferretti, 2023: 503, f. 2A-D, 3A-I, 4A-F, 5A-H, 7 (Dmf) Original description
Acanthogonatus minimus Indicatti, Folly-Ramos, Vargas, Lucas & Brescovit, 2015 | | Brazil urn:lsid:nmbe.ch:spidersp:047745
Acanthogonatus minimus Indicatti et al., 2015: 126, f. 1-30 (Dmf) Original description
Acanthogonatus mulchen Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001034
Acanthogonatus mulchen Goloboff, 1995: 124, f. 110, 112A-B (Df) Original description
Acanthogonatus nahuelbuta Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001035
Acanthogonatus nahuelbuta Goloboff, 1995: 118, f. 15, 61, 75-76, 103A-I, 104-105, 106A-B (Dmf) Original description
Acanthogonatus notatus (Mello-Leitão, 1940) | | Argentina urn:lsid:nmbe.ch:spidersp:001036
Chubutia notata Mello-Leitão, 1940b: 4, f. 1-5 (Dm) Original description
Acanthogonatus notatus Goloboff, 1995: 113, f. 100A-E (m, Df, removed from S of Acanthogonatus patagonicus, contra Gerschman & Schiapelli, 1970a: 152, sub Tryssothele)
Acanthogonatus notatus Signorotto, Mancini & Ferretti, 2023: 502, f. 1A-E (m)
Acanthogonatus parana Goloboff, 1995 | | Argentina urn:lsid:nmbe.ch:spidersp:001037
Acanthogonatus parana Goloboff, 1995: 99, f. 91A-B (Df) Original description
Acanthogonatus patagallina Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001038
Acanthogonatus patagallina Goloboff, 1995: 122, f. 108A-B (Dm) Original description
Acanthogonatus patagonicus (Simon, 1905) | | Chile, Argentina urn:lsid:nmbe.ch:spidersp:001039
Tryssothele patagonica Simon, 1905e: 2 (Df) Original description
Tryssothele patagonica Schiapelli & Gerschman, 1962b: 72, pl. IV, f. 2 (f)
Tryssothele patagonica Gerschman & Schiapelli, 1968: 25, f. 4-5 (f)
Tryssothele patagonica Gerschman & Schiapelli, 1970a: 152 (Sm)
Acanthogonatus patagonicus Goloboff, 1995: 109, f. 98A-D (f, Dm)
Acanthogonatus peniasco Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001040
Acanthogonatus peniasco Goloboff, 1995: 108, f. 58, 97 (Df) Original description
Acanthogonatus pissii (Simon, 1889) | | Chile urn:lsid:nmbe.ch:spidersp:001041
Brachythele pissii Simon, 1889a: 221 (Df) Original description
Tryssothele pissii Simon, 1902h: 7 (Df)
Tryssothele latastei Simon, 1902h: 7 (Df) [urn:lsid:nmbe.ch:spidersp:039250]
Tryssothele australis Chamberlin, 1917: 72, pl. 5, f. 8 (Df) [urn:lsid:nmbe.ch:spidersp:039249]
Aphantopelma venosum Mello-Leitão, 1936d: 119, pl. 12 (Df) [urn:lsid:nmbe.ch:spidersp:039251]
Tryssothele pissii Gerschman & Schiapelli, 1968: 27, f. 18-23 (f, S of Tryssothele australis and T. latastei)
Tryssothele pissii Calderón, 1978: 149, f. 1-5 (Dm)
Tryssothele pissii Legendre & Calderón, 1984: 1049, pl. XV, f. 1-14 (mf)
Acanthogonatus pissii Schaefer & Rudloff, 1995: 59, f. 1-13 (mf)
Acanthogonatus pissii Goloboff, 1995: 90, f. 86A-B, 87 (mf, S of Aphantopelma venosum)
Acanthogonatus quilocura Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001042
Acanthogonatus quilocura Goloboff, 1995: 86, f. 83A-C (Dmf) Original description
Acanthogonatus recinto Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001043
Acanthogonatus recinto Goloboff, 1995: 106, f. 57, 96A-F (Dmf) Original description
Acanthogonatus subcalpeianus (Nicolet, 1849) | | Chile urn:lsid:nmbe.ch:spidersp:001044
Mygale subcalpeiana Nicolet, 1849: 334 (Df) Original description
Stenoterommata guttulatum Simon, 1886h: 573 (Df) [urn:lsid:nmbe.ch:spidersp:039273]
Brachythele subcalpetana Simon, 1889a: 221
Tryssothele subcalpetana Simon, 1903a: 965
Tryssothele subcalpetana Schiapelli & Gerschman, 1962b: 72, pl. IV, f. 1 (f)
Tryssothele subcalpetana Gerschman & Schiapelli, 1968: 29, f. 1-3 (f, not m, = A. campanae)
Tryssothele subcalpetana Legendre & Calderón, 1984: 1051, pl. XVI, f. 1-4 (f)
Acanthogonatus subcalpeianus Goloboff, 1995: 78, f. 79A-H (f, Dm, S of Stenoterommata guttulatum)
Acanthogonatus tacuariensis (Pérez-Miles & Capocasale, 1982) | | Brazil, Uruguay urn:lsid:nmbe.ch:spidersp:001045
Pycnothelopsis auripilus Capocasale & Pérez-Miles, 1979: 3, f. 1, 4-6 (f, misidentified)
Pycnothelopsis tacuariensis Pérez-Miles & Capocasale, 1982: 2, pl. 1; pl. 2, f. 3-6 (Dmf) Original description
Acanthogonatus tacuariensis Raven, 1985a: 101 (Tmf from Pycnothelopsis=Pycnothele)
Acanthogonatus tacuariensis Goloboff, 1995: 88, f. 85A-C (mf)
Acanthogonatus tolhuaca Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001046
Acanthogonatus tolhuaca Goloboff, 1995: 126, f. 56, 112C-D (Df) Original description
Acanthogonatus vilches Goloboff, 1995 | | Chile urn:lsid:nmbe.ch:spidersp:001047
Acanthogonatus vilches Goloboff, 1995: 123, f. 109A-D (Df) Original description

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