World Spider Catalog

Lycosidae Sundevall, 1833

Family | ACCEPTED

N.B.: according to Brignoli, 1983c: 432, "Guy (1966, 1969) has proposed many synonymies and has downgraded many genera to subgenus rank; as this author based his conclusions more on nomenclature than on the study of material (or even of the original descriptions)", Platnick (WSC 15.0) did not accept Guy's conclusions; the changes by Guy, 1966 are noted (except where more recent work has superseded them); Guy, 1969 is merely a literature compilation that reflects those changes and is not cited separately; several genera of Lycosidae have been found not to be monophyletic (Murphy et al., 2006). Piacentini & Ramírez, 2019: 233-237 confirm ten valid subfamilies of Lycosidae: Venoniinae Lehtinen & Hippa, 1979, Zoicinae Lehtinen & Hippa, 1979 (= Piratinae Zyuzin, 1993), Evippinae Zyuzin, 1985, Sosippinae Dondale, 1986 (= Hygrolycosinae He & Song, 1996), Artoriinae Framenau, 2007, Tricassinae Alderweireldt & Jocqué, 1993 (= Arctosinae He & Song, 1996), Hippasinae Simon, 1898, Allocosinae Dondale, 1986, Pardosinae Simon, 1898 (= Wadicosinae Zyuzin, 1985) and Lycosinae Sundevall, 1833; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 518, f. 17-18.

In synonymy:

Potamia C. L. Koch, 1847 = Pirata Sundevall, 1833 (Simon, 1876a: 294)

Sosilaus Simon, 1898 = Pirata Sundevall, 1833 (Wallace & Exline, 1978: 79)

Gen. Pirata Sundevall, 1833 urn:lsid:nmbe.ch:spidergen:01650

N.B.: considered a senior synonym of Potamia C. L. Koch, 1847: v. 14: 98 [urn:lsid:nmbe.ch:spidergen:06159] (type Araneus piraticus Clerck, 1757) by Simon, 1876a: 294, Potamia was described as subgenus of Lycosa Latreille, 1804 and elevated to genus by Thorell, 1856: 63, Potamia is preoccupied several times (see Bonnet, 1958: 3767); considered a senior synonym of Sosilaus Simon, 1898a: 350 [urn:lsid:nmbe.ch:spidergen:05878] (type S. spiniger Simon, 1898) by Wallace & Exline, 1978: 79 (through transfer of type species); not a senior synonym of Piratula Roewer, 1960 (Omelko, Marusik & Koponen, 2011: 213, contra Dondale & Redner, 1981b: 107, Breitling, 2019b: 21 and Breitling, 2021: 26 (the two latter mainly based on COI, see FAQ 7)); see notes under Caporiaccosa and Foxicosa.

Transferred to other genera:

Pirata abalosi (Mello-Leitão, 1942) -- see Prolycosides

Pirata boreus Tanaka, 1974 -- see Piratula

Pirata brisbanae (L. Koch, 1878) -- see Venatrix

Pirata canadensis Dondale & Redner, 1981 -- see Piratula

Pirata cantralli Wallace & Exline, 1978 -- see Piratula

Pirata cereipes (L. Koch, 1878) -- see Halocosa

Pirata clercki (Bösenberg & Strand, 1906) -- see Piratula

Pirata denticulatus Liu, 1987 -- see Piratula

Pirata emertoni Dahl, 1908 -- see Trebacosa

Pirata eugeni (Roewer, 1951) -- see Piratula

Pirata exiguus Tanaka, 1974 -- see Piratula

Pirata fabella (Karsch, 1879) -- see Trochosa

Pirata figuratus (Simon, 1876) -- see Arctosa

Pirata giganteus Gertsch, 1934 -- see Piratula

Pirata hiroshii Tanaka, 1986 -- see Piratula

Pirata hokkaidensis Tanaka, 2003 -- see Piratula

Pirata humicola Montgomery, 1902 -- see Piratula

Pirata hurkai Buchar, 1966 -- see Piratula

Pirata hygrophilus Thorell, 1872 -- see Piratula

Pirata insularis Emerton, 1885 -- see Piratula

Pirata iriomotensis Tanaka, 1989 -- see Piratula

Pirata japonicus Tanaka, 1974 -- see Piratula

Pirata kempi (Gravely, 1924) -- see Lycosa

Pirata knorri (Scopoli, 1763) -- see Piratula

Pirata latitans (Blackwall, 1841) -- see Piratula

Pirata longjiangensis Yin, Peng, Xie, Bao & Wang, 1997 -- see Piratula

Pirata marxi Stone, 1890 -- see Trebacosa

Pirata mascatensis (Simon, 1898) -- see Allocosa

Pirata meridionalis Tanaka, 1974 -- see Piratula

Pirata minutus Emerton, 1885 -- see Piratula

Pirata molensis (Strand, 1908) -- see Trochosa

Pirata montigena Liu, 1987 -- see Piratula

Pirata mossambicus (Roewer, 1960) -- see Trochosa

Pirata nanshanensis Yin & Wang, 1984 -- see Piratula

Pirata oneili (Purcell, 1903) -- see Arctosa

Pirata piccolo Dahl, 1908 -- see Piratula

Pirata piratoides (Bösenberg & Strand, 1906) -- see Piratula

Pirata praedatorius Schenkel, 1953 -- see Piratula

Pirata procurvus (Bösenberg & Strand, 1906 -- see Piratula

Pirata procurvus sinensis Song, Shang, Wang, Cheng & Cheng, 1978 -- see Piratula

Pirata qinlingensis Zheng & Qiu, 1980 -- see Piratula

Pirata sagaphilus (Strand, 1918) -- see Piratula

Pirata sanya Barrion, Barrion-Dupo & Heong, 2013 -- see Lysania

Pirata serrulatus Song & Wang, 1984 -- see Piratula

Pirata simplex (L. Koch, 1882) -- see Hogna

Pirata soukupi (Mello-Leitão, 1942) -- see Prolycosides

Pirata sternomaculatus (Mello-Leitão, 1943) -- see Hogna

Pirata tanakai Brignoli, 1983 -- see Piratula

Pirata taurirtensis (Schenkel, 1937) -- see Hogna

Pirata tenuisetaceus Chai, 1987 -- see Piratula

Pirata uliginosus (Thorell, 1856) -- see Piratula

Pirata velox Keyserling, 1891 -- see Agalenocosa

Pirata wuchangensis Schenkel, 1963 -- see Piratula

Pirata yaginumai Tanaka, 1974 -- see Piratula

Pirata yesoensis Tanaka, 1985 -- see Piratula

In synonymy:

Pirata arenicola Emerton, 1909 = Pirata aspirans Chamberlin, 1904 (Wallace & Exline, 1978: 30)

Pirata blabakensis Barrion & Litsinger, 1995 = Pirata subpiraticus Bösenberg & Strand, 1906 (Omelko, Marusik & Koponen, 2011: 213)

Pirata californicus Banks, 1904 = Pirata sedentarius Montgomery, 1904 (Wallace & Exline, 1978: 72)

Pirata catindigae Barrion, Barrion-Dupo & Heong, 2013 = Pirata subpiraticus Bösenberg & Strand, 1906 (Wang et al., 2021a: 53)

Pirata gertschi Chamberlin & Ivie, 1944 = Pirata mayaca Gertsch, 1940 (Wallace & Exline, 1978: 48)

Pirata gratus Gertsch & Davis, 1940, T from Allocosa = Pirata felix O. Pickard-Cambridge, 1898 (Wallace & Exline, 1978: 55)

Pirata haploapophysis Chai, 1987 = Pirata subpiraticus Bösenberg & Strand, 1906 (Omelko, Marusik & Koponen, 2011: 213)

Pirata luzonensis Barrion & Litsinger, 1995 = Pirata subpiraticus Bösenberg & Strand, 1906 (Omelko, Marusik & Koponen, 2011: 213)

Pirata maculatus Emerton, 1909 = Pirata sedentarius Montgomery, 1904 (Wallace & Exline, 1978: 72)

Pirata piraticus moravicus Kratochvíl, 1931 = Pirata tenuitarsis Simon, 1876 (Michelucci & Tongiorgi, 1976: 156)

Pirata piraticus utahensis Chamberlin, 1908 = Pirata piraticus Clerck, 1757 (Wallace & Exline, 1978: 66)

Pirata shibatai Tanaka, 1995 = Pirata praedo Kulczyński, 1885 (Marusik, Omelko & Koponen, 2010: 33)

Pirata wacondana Scheffer, 1904 (removed from S of P. piraticus) = Pirata sedentarius Montgomery, 1904 (Wallace & Exline, 1978: 72)

Pirata zelotes Wallace & Exline, 1978 = Pirata praedo Kulczyński, 1885 (Omelko, Marusik & Koponen, 2011: 209)

Nomina dubia:

Pirata albicomaculatus Franganillo, 1913: 125 (mf, Spain) -- Breitling et al., 2016b: 54

Pirata rubicundicoloratus (Strand, 1906b: 675, f, Algeria, originally in Tarentula) -- Nentwig et al., 2020: 24

Pirata subniger Franganillo, 1913: 124 (f, Spain) -- Breitling et al., 2016b: 54

Pirata affinis Roewer, 1960 | | Congo urn:lsid:nmbe.ch:spidersp:019052

Pirata affinis Roewer, 1960d: 836, f. 463a-d (Dmf) Original description

Pirata africana (Roewer, 1960) | | Namibia urn:lsid:nmbe.ch:spidersp:017891

Hippasa africana Roewer, 1960d: 994, f. 550a-b (Df) Original description

Pirata africana Alderweireldt & Jocqué, 2005: 65 (Tf from Hippasa)

Pirata africana Dippenaar-Schoeman et al., 2021l: 32, p. 31: 2 f., p. 32: 5 f. (f)

Pirata alachuus Gertsch & Wallace, 1935 | | USA urn:lsid:nmbe.ch:spidersp:019053

Pirata alachua Gertsch & Wallace, 1935: 9, f. 34, 36 (Dmf) Original description

Allocosa alachua Roewer, 1955c: 207

Pirata alachuus Wallace & Exline, 1978: 82, f. 8, 169-174 (Tmf from Allocosa)

Pirata allapahae Gertsch, 1940 | | USA urn:lsid:nmbe.ch:spidersp:019055

Pirata allapahae Gertsch, 1940: 23, f. 9 (Dm) Original description

Allocosa allapahae Roewer, 1955c: 207

Pirata allapahae Wallace & Exline, 1978: 16, f. 14-19, 61 (Tmf from Allocosa)

Pirata apalacheus Gertsch, 1940 | | USA urn:lsid:nmbe.ch:spidersp:019056

Pirata apalacheus Gertsch, 1940: 17, f. 3-4 (Dmf) Original description

Lycosa apalachea Roewer, 1955c: 273

Pirata apalacheus Wallace & Exline, 1978: 18, f. 20-25 (Tmf from Lycosa)

Pirata aspirans Chamberlin, 1904 | | USA, Canada urn:lsid:nmbe.ch:spidersp:019057

Pirata aspirans Chamberlin, 1904b: 286 (Df) Original description

Pirata aspirans Chamberlin, 1908: 303, pl. 27, f. 5-6 (f, Dm)

Pirata insularis Chamberlin, 1908: 309, pl. 22, f. 4 (m, misidentified, not f)

Pirata arenicola Emerton, 1909: 208, pl. 6, f. 9 (Dmf) [urn:lsid:nmbe.ch:spidersp:056629]

Pirata aspirans Comstock, 1912: 643, f. 724 (f, not m)

Pirata insularis Comstock, 1912: 645, f. 724e (f, misidentified, not m)

Pirata arenicola Kaston, 1938a: 16, f. 2 (f)

Pirata arenicola Kaston, 1948: 311, f. 989, 1002, 1012 (mf)

Allocosa arenicola Roewer, 1955c: 209

Allocosa aspirans Roewer, 1955c: 209

Pirata aspirans Wallace & Exline, 1978: 30, f. 4, 10, 51-52, 55-60 (mf, T from Allocosa, S of Allocosa arenicola)

Pirata aspirans Dondale & Redner, 1990: 257, f. 387-389 (mf)

Pirata aspirans Paquin & Dupérré, 2003: 166, f. 1864-1868 (mf)

Pirata brevipes (Banks, 1893) | | Congo urn:lsid:nmbe.ch:spidersp:019060

Lycosa brevipes Banks, 1893b: 587, f. 12 (Df) Original description

Pirata brevipes Roewer, 1955c: 285

Pirata brevipes Roewer, 1960d: 835, f. 462 (f)

Pirata browni Gertsch & Davis, 1940 | | Mexico urn:lsid:nmbe.ch:spidersp:019062

Pirata browni Gertsch & Davis, 1940a: 9, f. 18 (Df) Original description

Allocosa browni Roewer, 1955c: 207

Pirata browni Wallace & Exline, 1978: 57, f. 113-114 (Tf from Allocosa)

Pirata bryantae Kurata, 1944 | | Canada, USA (Alaska) urn:lsid:nmbe.ch:spidersp:019063

Pirata bryantae Kurata, 1944: 3, f. 1-4 (Df) Original description

Pirata bryantae Wallace & Exline, 1978: 85, f. 175-180 (f, Dm)

Pirata bryantae Dondale & Redner, 1990: 260, f. 396-399 (mf)

Pirata bryantae Paquin & Dupérré, 2003: 167, f. 1867-1869 (mf)

Pirata chamberlini (Lessert, 1927) | | Congo, East Africa urn:lsid:nmbe.ch:spidersp:019067

Lycosa (P.) chamberlini Lessert, 1927: 409, f. 2 (Dmf) Original description

Piratula chamberlini Roewer, 1955c: 288

Piratula chamberlini Roewer, 1960d: 678, f. 384a-d (mf)

Pirata coreanus Paik, 1991 | | Korea urn:lsid:nmbe.ch:spidersp:019069

Pirata coreanus Paik, 1991g: 61, f. 1-8 (Df) Original description

Pirata davisi Wallace & Exline, 1978 | | USA, Mexico urn:lsid:nmbe.ch:spidersp:019070

Pirata davisi Wallace & Exline, 1978: 87, f. 181-186 (Dmf) Original description

Pirata davisi Breene et al., 1993: 95, f. 131A-B (mf)

Pirata digitatus Tso & Chen, 2004 | | Taiwan urn:lsid:nmbe.ch:spidersp:040103

Pirata digitatus Tso & Chen, 2004: 403, f. 17-24 (Dmf) Original description

Pirata felix O. Pickard-Cambridge, 1898 | | Mexico urn:lsid:nmbe.ch:spidersp:019073

Pirata felix O. Pickard-Cambridge, 1898a: 243, pl. 33, f. 6 (Df) Original description

Pirata felix F. O. Pickard-Cambridge, 1902a: 330, pl. 31, f. 23 (f)

Pirata gratus Gertsch & Davis, 1940a: 10, f. 19, 21 (Dmf) [urn:lsid:nmbe.ch:spidersp:054453]

Allocosa felix Roewer, 1955c: 207

Allocosa grata Roewer, 1955c: 208

Pirata felix Wallace & Exline, 1978: 55, f. 105-112 (mf, T from Allocosa, S of Allocosa grata)

Pirata hiteorum Wallace & Exline, 1978 | | USA urn:lsid:nmbe.ch:spidersp:019077

Pirata hiteorum Wallace & Exline, 1978: 89, f. 6, 192-198, 204 (Dmf) Original description

Pirata indigena Wallace & Exline, 1978 | | USA urn:lsid:nmbe.ch:spidersp:019080

Pirata indigenus Wallace & Exline, 1978: 34, f. 62-65, 72 (Df) Original description

Pirata iviei Wallace & Exline, 1978 | | USA urn:lsid:nmbe.ch:spidersp:019083

Pirata iviei Wallace & Exline, 1978: 36, f. 4, 67-71 (Dmf) Original description

Pirata mayaca Gertsch, 1940 | | USA, Bahama Is., Cuba urn:lsid:nmbe.ch:spidersp:019088

Pirata mayaca Gertsch, 1940: 21, f. 5-6 (Dmf) Original description

Pirata gertschi Chamberlin & Ivie, 1944: 149, pl. 13, f. 190-191 (Dmf) [urn:lsid:nmbe.ch:spidersp:056631]

Allocosa mayaca Roewer, 1955c: 208

Piratula gertschi Roewer, 1955c: 289

Pirata mayaca Wallace & Exline, 1978: 48, f. 5, 87, 90-96 (mf, T from Allocosa, S of Pirata gertschi)

Pirata montanoides Banks, 1892 | | USA urn:lsid:nmbe.ch:spidersp:019092

Pirata montanoides Banks, 1892a: 71, pl. 1, f. 46 (Df) Original description

Pirata montanoides Banks, 1916: 81, pl. 11, f. 20 (f)

Hogna montanoides Roewer, 1955c: 259

Pirata montanoides Wallace & Exline, 1978: 102, f. 11, 217-222 (Tf from Hogna, Dm)

Pirata montanoides Dondale & Redner, 1990: 267, f. 414-416 (mf)

Pirata montanus Emerton, 1885 | | USA, Canada, Russia (Kurile Is.) urn:lsid:nmbe.ch:spidersp:019093

Pirata montanus Emerton, 1885: 493, pl. 48, f. 9 (Df) Original description

Pirata elegans Stone, 1890: 429, pl. 15, f. 5 (Df)

Pirata agilis Banks, 1892a: 72, pl. 1, f. 47 (Df)

Pirata nigromaculatus Montgomery, 1902: 579, pl. 30, f. 44-45 (Dmf)

Pirata elegans Montgomery, 1902: 581, pl. 30, f. 46 (f)

Pirata montanus Chamberlin, 1908: 306, pl. 10, f. 5-9, pl. 2, f. 9 (f)

Pirata montanus Emerton, 1911: 401, pl. 5, f. 3 (m)

Pirata montanus Comstock, 1912: 644, f. 131a, 724c

Pirata montanus Kaston, 1938a: 16, f. 6 (f)

Pirata montanus Comstock, 1940: 130, 655, f. 131a, 724c (f)

Pirata montanus Kaston, 1948: 308, f. 983, 1004, 1009 (mf)

Hogna montanus Roewer, 1955c: 259

Pirata montanus Wallace & Exline, 1978: 103, f. 6, 223-232 (Tmf from Hogna)

Pirata montanus Dondale & Redner, 1990: 269, f. 417-420 (mf)

Pirata montanus Paquin & Dupérré, 2003: 168, f. 1685-1687 (mf)

Pirata nanatus Gertsch, 1940 | | USA urn:lsid:nmbe.ch:spidersp:019096

Pirata nanatus Gertsch, 1940: 19, f. 7-8 (Dmf) Original description

Lycosa nanata Roewer, 1955c: 274

Pirata nanatus Wallace & Exline, 1978: 20, f. 26-31, 61 (Tmf from Lycosa)

Pirata niokolona Roewer, 1961 | | Senegal urn:lsid:nmbe.ch:spidersp:019097

Pirata niokolona Roewer, 1961b: 42, f. 6 (Df) Original description

Pirata pagicola Chamberlin, 1925 | | Mexico to Panama urn:lsid:nmbe.ch:spidersp:019098

Pirata pagicola Chamberlin, 1925d: 225 (Df) Original description

Pirata pagicola Wallace & Exline, 1978: 52, f. 5, 97-104 (f, Dm)

Pirata pallipes (Blackwall, 1857) | | Algeria urn:lsid:nmbe.ch:spidersp:019099

Lycosa pallipes Blackwall, 1857a: 286 (Df) Original description

Pirata pallipes Roewer, 1955c: 283

Pirata pallipes Roewer, 1960d: 827, f. 457a-b (f)

Pirata piratellus (Strand, 1907) | | Japan urn:lsid:nmbe.ch:spidersp:019100

Tarentula piratella Strand, 1907b: 565 (Df) Original description

Tarentula piratella Strand, 1909f: 64, f. 38 (f)

Pirata piratella Roewer, 1955c: 283

Pirata piraticus (Clerck, 1757) | | North America, Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, Iran, Central Asia, China, Japan urn:lsid:nmbe.ch:spidersp:019101

Araneus piraticus Clerck, 1757: 102, pl. 5, f. 4 (Df) Original description

Pirata piraticus Yaginuma, 1971: 87, f. 78.4 (f)

Aranea piratica Olivier, 1789: 218 (D)

Pirata piraticus Azheganova, 1971: 11, f. 14 (mf)

Araneus piraticus Martyn, 1793: 47, pl. 7, f. 7 (f)

Pirata piraticus Tystshenko, 1971: 182, f. 530, 535 (mf)

Lycosa piratica Walckenaer, 1805: 14

Pirata piraticus Fuhn & Niculescu-Burlacu, 1971: 213, f. 104a-e (mf)

Lycosa piratica Hahn, 1833a: 107, f. 80 (Dmf)

Pirata piraticus Miller, 1971: 169, pl. XXVI, f. 16-17 (mf)

Pirata piraticus Sundevall, 1833b: 24

Pirata piraticus Namkung, Paik & Yoon, 1971: 51, f. 4 (f)

Lycosa argenteomarginata Lucas, 1846: 120, pl. 3, f. 10 (Df)

Pirata piraticus Loksa, 1972: 60, f. 51, 53E-F, 54E-F (mf)

Lycosa piratica C. L. Koch, 1847: 1, f. 1413-1414 (mf)

Pirata piraticus Tanaka, 1974a: 23, f. 1-3 (mf)

Potamia piratica Thorell, 1856: 63

Pirata piraticus Michelucci & Tongiorgi, 1976: 155, f. 3, 5, 7, 9, 11, 13 (mf)

Lycosa piratica Blackwall, 1861a: 34, pl. 2, f. 16 (mf)

Pirata piraticus Jocqué, 1977e: 145, f. 5, 7 (mf)

Potamia argenteomarginata Simon, 1864: 352

Pirata piraticus Wallace & Exline, 1978: 66, f. 3, 53-54, 134-139 (mf, S of Pirata piraticus utahensis)

Pirata piraticus Thorell, 1872a: 341

Pirata piraticus Paik & Namkung, 1979: 68, f. 53.1-2 (mf)

Pirata piraticus Simon, 1876a: 300, pl. 12, f. 7-8 (mf)

Pirata piraticus Kronestedt, 1980b: 65, f. 2A, 3B, D, 4A, 6A-C, 7D-F, 8E-F, 9C-D (mf)

Pirata prodigiosa Keyserling, 1877a: 669, pl. 8, f. 44 (Df)

Pirata piraticus Snazell, 1983b: 97, f. 12-14 (mf)

Pirata piraticus Menge, 1879: 513, pl. 83, f. 290 (mf)

Pirata piraticus Hu, 1984: 243, f. 253.1-2 (f)

Lycosa febriculosa Becker, 1881b: 45, pl. 2, f. 2 (Df)

Pirata piraticus Roberts, 1985: 150, f. 65b, 66b (mf)

Pirata piraticus Hansen, 1882: 73, pl. 5, f. 6 (mf)

Pirata piraticus Zyuzin, 1985a: 44, f. 7

Pirata piraticus Becker, 1882b: 122, pl. 9, f. 9 (mf)

Pirata piraticus Yaginuma, 1986a: 167, f. 92.1 (mf)

Pirata piraticus Bösenberg, 1903: 406, pl. 38, f. 598 (mf)

Pirata piraticus Liu, 1987a: 45, pl. 1, f. 1 (m)

Pirata piraticus Dahl, 1908: 287, f. 34 (mf)

Pirata piraticus Liu, 1987b: 86, f. 24, 26, 28 (mf)

Pirata febriculosa Chamberlin, 1908: 311, pl. 22, f. 1-2 (mf)

Pirata piraticus Tanaka, 1988a: 36, f. 1-4 (mf)

Pirata piraticus utahensis Chamberlin, 1908: 313 (Dm) [urn:lsid:nmbe.ch:spidersp:056634]

Pirata piraticus Yu & Song, 1988c: 119 (S)

Pirata sylvestris Emerton, 1909: 209, pl. 6, f. 8 (Df, not m)

Pirata piraticus Hu & Wu, 1989: 233, f. 193.1-5 (mf)

Pirata febriculosa Comstock, 1912: 645, f. 724f

Pirata piraticus Chikuni, 1989b: 113, f. 17 (mf)

Pirata piraticus Gerhardt, 1921: 245, pl. 2, f. 1 (m)

Pirata piraticus Dondale & Redner, 1990: 270, f. 421-424, 426-427 (mf)

Pirata piraticus Dahl & Dahl, 1927: 64, f. 166-169 (mf)

Pirata piraticus Chen & Gao, 1990: 130, f. 162a-b (mf)

Pirata piraticus Reimoser, 1928: 107, f. 5 (mf)

Pirata piraticus Heimer & Nentwig, 1991: 346, f. 892 (mf)

Pirata piraticus Kratochvíl, 1931: 70, f. 4 (f)

Pirata piraticus Zhao, 1993: 107, f. 47a-c (mf)

Pirata piraticus Saito, 1934b: 351, pl. 15, f. 81 (f)

Pirata piraticus Barrion & Litsinger, 1994: 315, f. 1646-1648 (mf)

Pirata piraticus Gertsch & Wallace, 1937: 5

Pirata piraticus Roberts, 1995: 232, f. (mf)

Lycosa piraticus Simon, 1937: 1118, 1140, f. 1769-1770 (mf)

Pirata piraticus Zhao, 1995: 825, f. 383a-c (mf)

Pirata piraticus Kaston, 1938a: 16, f. 1 (f)

Pirata piraticus Agnarsson, 1996: 45, f. 27A-B (mf)

Pirata piraticus Palmgren, 1939: 70, f. 104, 109, 115 (mf)

Pirata piraticus Yin et al., 1997c: 37, f. 15a-d (mf)

Pirata febriculosa Comstock, 1940: 653, f. 724f (f)

Pirata piraticus Roberts, 1998: 250, f. (mf)

Pirata piraticus Holm, 1947: 10, f. 4c, pl. 1, f. 7-8, pl. 9, f. 4 (mf)

Pirata piraticus Song, Zhu & Chen, 1999: 344, f. 200K, 201B (mf)

Pirata piraticus Kaston, 1948: 309, f. 1003, 1010 (mf)

Pirata piraticus Paquin & Dupérré, 2003: 168, f. 1888-1891 (mf)

Pirata piraticus Locket & Millidge, 1951: 287, f. 139B, F (mf)

Pirata piraticus Almquist, 2005: 243, f. 238a-f (mf)

Pirata piraticus Brændegaard, 1958: 22, f. 13-14 (mf)

Pirata piraticus Yoo, Park & Kim, 2007: 25, f. 11-13 (m)

Pirata piraticus Saito, 1959: 58, f. 44a-b (f)

Pirata piraticus Tanaka, 2009: 223, f. 12-13 (mf)

Pirata piraticus Wiebes, 1959b: 61, f. 82, 92, 104-105 (mf)

Pirata piraticus Zhu & Zhang, 2011: 284, f. 206A-D (mf)

Pirata piratica Yaginuma, 1960: 87, f. 78.4 (f)

Pirata piraticus Nadolny & Kovblyuk, 2011: 186, f. 4, 8, 13-14, 108, 114-116, 124 (mf)

Pirata piraticus Yaginuma, 1965b: 366, f. 9.9 (f)

Pirata piraticus Omelko, Marusik & Koponen, 2011: 207, f. 1a, 3a, 6-16, 42-43, 167, 175 (mf)

Pirata piraticus Buchar, 1966a: 217, f. 3D, G (mf)

Pirata piraticus Yin et al., 2012: 778, f. 388a-d (mf)

Pirata piraticus Azheganova, 1968: 27, f. 34-35 (mf)

Pirata piraticus Liu, Wang & Zhang, 2015: 421, f. 13-20 (mf)

Pirata piraticus von Helversen & Harms, 1969: 367, f. 3 (m)

Pirata piraticus Suzuki et al., 2021: 255, f. 4a-b (f)

Pirata piraticus Zhang, Peng & Zhang, 2022: 150, f. 111A-H (mf)

Pirata piratimorphus (Strand, 1908) | | USA urn:lsid:nmbe.ch:spidersp:019102

Tarentula piratimorpha Strand, 1908e: 244 (Df) Original description

Lycosa piratimorpha Petrunkevitch, 1911: 564

Piratula piratimorpha Roewer, 1955c: 289 (N.B.: presumably misplaced in Pirata)

Pirata piscatorius (Clerck, 1757) | | Europe, Russia (Europe to Middle Siberia) urn:lsid:nmbe.ch:spidersp:019104

Araneus piscatorius Clerck, 1757: 103, pl. 5, f. 5 (Df) Original description

Aranea piscatoria Olivier, 1789: 218 (D)

Araneus piscatorius Martyn, 1793: 48, pl. 11, f. 9 (f)

Trochosa umbraticola C. L. Koch, 1847: 137, f. 1368 (Df)

Lycosa piscatorius Westring, 1851: 54 (D)

Potamia piscatorius Thorell, 1856: 64

Lycosa de greyi O. Pickard-Cambridge, 1871a: 396, pl. 54, f. 3 (Dm)

Pirata piscatorius Thorell, 1872a: 339

Pirata piscatorius Menge, 1875: 512, pl. 83, f. 289 (Dmf)

Pirata piscatorius Becker, 1882b: 121, pl. 9, f. 8 (mf)

Pirata piscatorius Bösenberg, 1903: 405, pl. 38, f. 597 (mf)

Pirata piscatorius Dahl, 1908: 292, f. 33 (mf)

Pirata piscatorius Dahl & Dahl, 1927: 63, f. 164-165 (mf)

Lycosa piscatorius Simon, 1937: 1118, 1140, f. 1767-1768 (mf)

Pirata piscatorius Palmgren, 1939: 69, f. 101, 106, 114 (mf)

Pirata piscatorius Holm, 1947: 10, pl. 2, f. 11-12, pl. 9, f. 3 (mf)

Pirata piscatorius Locket & Millidge, 1951: 290, f. 139A, G (mf)

Pirata piscatorius Wiebes, 1959b: 62, f. 90, 97 (mf)

Pirata piscatorius Azheganova, 1968: 28, f. 36-37 (mf)

Pirata piscatorius Azheganova, 1971: 12, f. 15 (mf)

Pirata piscatorius Tystshenko, 1971: 182, f. 531 (m)

Pirata piscatorius Fuhn & Niculescu-Burlacu, 1971: 216, f. 106a-e (mf)

Pirata piscatorius Miller, 1971: 169, pl. XXVI, f. 21-22 (mf)

Pirata piscatorius Loksa, 1972: 61, f. 53C-D, 54C-D (mf)

Pirata piscatorius Kronestedt, 1980b: 65, f. 9G-H (m)

Pirata piscatorius Roberts, 1985: 150, f. 65a (mf)

Pirata piscatorius Heimer & Nentwig, 1991: 346, f. 899 (mf)

Pirata piscatorius Roberts, 1995: 232, f. (mf)

Pirata piscatorius Roberts, 1998: 249, f. (mf)

Pirata piscatorius Almquist, 2005: 244, f. 239a-e (mf)

Pirata piscatorius Dolejš, 2015: 4, f. 1-2 (m)

Pirata piscatorius Lecigne, 2020: 58, pl. 4K-L, ph. 2 (mf)

Pirata praedo Kulczyński, 1885 | | Russia (Urals to Far East), Mongolia, China, Japan, USA, Canada urn:lsid:nmbe.ch:spidersp:019105

Pirata praedo Kulczyński, 1885: 56, pl. 11, f. 34 (Dmf) Original description

Pirata praedo Schenkel, 1930b: 31, f. 12a-b (mf)

Piratula praedo Roewer, 1955c: 288

Pirata zelotes Wallace & Exline, 1978: 70, f. 13, 140-142, 145 (Dmf) [urn:lsid:nmbe.ch:spidersp:019139]

Pirata zelotes Dondale & Redner, 1990: 273, f. 425, 428-432 (mf)

Pirata praedo Logunov, 1992e: 60, f. 6 (mf, removed from S of P. piraticus, contra Yu & Song, 1988c: 119)

Pirata shibatai Tanaka, 1995: 47, f. 1-4 (Dmf) [urn:lsid:nmbe.ch:spidersp:019113]

Pirata zelotes Paquin & Dupérré, 2003: 169, f. 1895-1898 (mf)

Pirata shibatai Tanaka, 2009: 225, f. 16-17 (mf)

Pirata praedo Marusik, Omelko & Koponen, 2010: 33, f. 1-29 (mf, S of Pirata shibatai)

Pirata praedo Omelko, Marusik & Koponen, 2011: 209, f. 1b, 3b-c, 17-26b, 39-41, 169, 177 (mf, S of Pirata zelotes)

Pirata praedo Liu, Wang & Zhang, 2015: 421, f. 1-12 (mf)

Pirata proximus O. Pickard-Cambridge, 1876 | | Egypt urn:lsid:nmbe.ch:spidersp:019107

Pirata proxima O. Pickard-Cambridge, 1876b: 598 (Df) Original description

Pirata proxima Roewer, 1960d: 830, f. 459a-b (f)

Pirata punctipes (Gravely, 1924) | | India urn:lsid:nmbe.ch:spidersp:019341

Lycosa punctipes Gravely, 1924: 603, f. 3I (Dmf) Original description

Crocodilosa punctipes Roewer, 1955c: 238

Trochosa punctipes Tikader & Malhotra, 1980: 436, f. 346-348 (Tf from Crocodilosa)

Pirata punctipes Sankaran & Caleb, 2023b: 545, f. 9A-B, 10A-C (f, T from Trochosa)

Pirata sagitta (Mello-Leitão, 1941) | | Argentina urn:lsid:nmbe.ch:spidersp:019109

Lycosa sagitta Mello-Leitão, 1941d: 133, f. 30, pl. 6, f. 24 (Dm) Original description

Sosilaus sagitta Roewer, 1955c: 296

Pirata sedentarius Montgomery, 1904 | | North America, Greater Antilles urn:lsid:nmbe.ch:spidersp:019110

Pirata sedentarius Montgomery, 1904: 312, pl. 19, f. 28-29 (Dmf) Original description

Pirata californica Banks, 1904b: 356, pl. 39, f. 13 (Dmf) [urn:lsid:nmbe.ch:spidersp:056630]

Lycosa wacondana Scheffer, 1904b: 260, pl. 17, f. 7 (Df) [urn:lsid:nmbe.ch:spidersp:056635]

Pirata maculatus Emerton, 1909: 209, pl. 6, f. 10 (Df) [urn:lsid:nmbe.ch:spidersp:056632]

Pirata wacondana Crosby & Bishop, 1928b: 1069

Pirata maculatus Kaston, 1938a: 16, f. 3 (f)

Pirata sedentarius Comstock, 1940: 653, f. 724e (f)

Pirata maculatus Kaston, 1948: 311, f. 990-991, 1007, 1013 (f, Dm)

Allocosa maculata Roewer, 1955c: 211

Piratula sedentarius Roewer, 1955c: 289

Pirata sedentarius Wallace & Exline, 1978: 72, f. 1, 143-144, 146-162 (mf, S of Allocosa maculata, Pirata californicus and P. wacondana)

Pirata sedentarius Dondale & Redner, 1990: 266, f. 408-413 (mf)

Pirata sedentarius Paquin & Dupérré, 2003: 168, f. 1892-1894 (mf)

Pirata seminolus Gertsch & Wallace, 1935 | | USA urn:lsid:nmbe.ch:spidersp:019111

Pirata seminola Gertsch & Wallace, 1935: 7, f. 33, 35 (Dmf) Original description

Piratula seminola Roewer, 1955c: 289

Pirata seminola Wallace & Exline, 1978: 22, f. 1, 32-43, 61 (mf)

Pirata seminola Dondale & Redner, 1990: 264, f. 404-407 (mf)

Pirata seminola Breene et al., 1993: 95, f. 130A-B (mf)

Pirata spatulatus Chai, 1985 | | China urn:lsid:nmbe.ch:spidersp:019116

Pirata spatulatus Chai, 1985: 76, f. 1-6 (Dm) Original description

Pirata spiniger (Simon, 1898) | | USA urn:lsid:nmbe.ch:spidersp:019117

Sosilaus spiniger Simon, 1898a: 350, f. 340-341 (Dm) Original description

Sosilaus spiniger Chamberlin, 1908: 298, pl. 12, f. 5-6 (m)

Sosilaus spiniger Banks, 1913: 182, pl. 12, f. 19 (m)

Pirata spiniger Wallace & Exline, 1978: 79, f. 163-168 (m, Df)

Pirata subannulipes (Strand, 1906) | | Ethiopia urn:lsid:nmbe.ch:spidersp:019118

Tarentula subannulipes Strand, 1906b: 686 (Df) Original description

Tarentula subannulipes Strand, 1908l: 53, pl. 2, f. 26 (f)

Pirata subannulipes Roewer, 1955c: 285

Pirata subannulipes Roewer, 1960d: 834, f. 461 (f)

Pirata subpiraticus (Bösenberg & Strand, 1906) | | Russia (Far East), Korea, China, Japan, Indonesia (Java), Philippines urn:lsid:nmbe.ch:spidersp:019120

Tarentula subpiratica Bösenberg & Strand, 1906: 317, pl. 13, f. 339 (Df) Original description

Piratula subpiraticus Roewer, 1955c: 288

Pirata subpiratica Saito, 1959: 59, f. 47a-b (f)

Pirata subpiratica Yaginuma, 1960: 87, f. 78.1 (f)

Pirata subpiraticus Yaginuma, 1965b: 366, f. 6 (f)

Pirata subpiraticus Yaginuma, 1971: 87, f. 78.1 (f)

Pirata subpiraticus Tanaka, 1974a: 25, f. 4-6 (mf)

Pirata subpiraticus Yin, 1978: 11, f. 5A-D (mf)

Pirata subpiraticus Song et al., 1978: 4, f. 7A-D (mf)

Pirata subpiraticus Paik & Namkung, 1979: 68, f. 54.1-2 (mf)

Pirata subpiraticus Song, 1980: 169, f. 91a-d (mf)

Pirata subpiraticus Wang, 1981: 120, f. 60A-C (mf)

Pirata subpiraticus Hu, 1984: 245, f. 257.1-2 (mf)

Pirata subpiraticus Guo, 1985: 124, f. 2-64.1-3 (mf)

Pirata subpiraticus Yaginuma, 1986a: 169, f. 92.6 (mf)

Pirata subpiraticus Song, 1987: 243, f. 198 (mf)

Pirata haploapophysis Chai, 1987: 362, f. 1.1-9 (Dmf) [urn:lsid:nmbe.ch:spidersp:019075]

Pirata subpiraticus Zhang, 1987: 159, f. 135.1-4 (mf)

Pirata subpiraticus Liu, 1987a: 45, pl. 1, f. 2 (m)

Pirata haploapophysis Liu, 1987a: 45, pl. 1, f. 3 (m, nomen nudum)

Pirata subpiraticus Liu, 1987b: 86, f. 25, 27, 29 (mf)

Pirata subpiraticus Tanaka, 1988a: 39, f. 5-8 (mf)

Pirata subpiraticus Chikuni, 1989b: 113, f. 19 (mf)

Pirata subpiraticus Feng, 1990: 156, f. 131.1-2 (mf)

Pirata subpiraticus Chen & Gao, 1990: 131, f. 164a-b (mf)

Pirata subpiraticus Chen & Zhang, 1991: 214, f. 219.1-4 (mf)

Pirata haploapophysis Zhao, 1993: 104, f. 44a-c (mf)

Pirata subpiraticus Zhao, 1993: 107, f. 48a-b (mf)

Pirata subpiraticus Barrion & Litsinger, 1994: 315, f. 1642-1645 (mf)

Pirata luzonensis Barrion & Litsinger, 1995: 366, f. 214a-h, 215a-h (Dmf) [urn:lsid:nmbe.ch:spidersp:019087]

Pirata blabakensis Barrion & Litsinger, 1995: 368, f. 216a-f, 217a-h (Dmf) [urn:lsid:nmbe.ch:spidersp:019058]

Pirata haloapophysis Zhao, 1995: 822, f. 380a-c (mf)

Pirata subpiraticus Zhao, 1995: 826, f. 384a-b (mf)

Pirata haploapophysis Yin et al., 1997c: 31, f. 11a-d (mf)

Pirata subpiraticus Yin et al., 1997c: 44, f. 17a-d (mf)

Pirata haploapophysis Song, Zhu & Chen, 1999: 335, f. 200G, S (mf)

Pirata subpiraticus Song, Zhu & Chen, 1999: 344, f. 200N, 201E (mf)

Pirata subpiraticus Hu, 2001: 215, f. 113.1-3 (f)

Pirata subpiraticus Song, Zhu & Chen, 2001: 260, f. 162A-D (mf)

Pirata subpiraticus Namkung, 2002: 338, f. 20.32a-b (mf)

Pirata subpiraticus Namkung, 2003: 340, f. 20.32a-b (mf)

Pirata subpiraticus Yoo & Framenau, 2006: 680, f. 4D (m)

Pirata subpiraticus Tanaka, 2009: 223, f. 14-15 (mf)

Pirata haploapophysis Zhu & Zhang, 2011: 283, f. 205A-C (mf)

Pirata subpiraticus Zhu & Zhang, 2011: 287, f. 209A-D (mf)

Pirata subpiraticus Omelko, Marusik & Koponen, 2011: 213, f. 3d, 27-38, 46-47, 168, 176 (mf, S of Pirata blabakensis, P. haploapophysis and P. luzonensis)

Pirata haploapophysis Yin et al., 2012: 775, f. 386a-d (mf)

Pirata subpiraticus Yin et al., 2012: 783, f. 391a-d, 3-19a-b (mf)

Pirata catindigae Barrion, Barrion-Dupo & Heong in Barrion et al., 2013: 18, f. 19A-D (Df) [urn:lsid:nmbe.ch:spidersp:048431]

Pirata subpiraticus Wang et al., 2021a: 53, f. 49A-D, 50A-H, 51A-D (mf, S of Pirata catindigae)

Pirata subpiraticus Zhang, Peng & Zhang, 2022: 154, f. 112A-H (mf)

Pirata subpiraticus Hidaka, 2022b: 46, f. 5-6 (f)

Pirata suwaneus Gertsch, 1940 | | USA, Bahama Is. urn:lsid:nmbe.ch:spidersp:019121

Pirata suwaneus Gertsch, 1940: 20, f. 1-2 (Dmf) Original description

Allocosa suwanea Roewer, 1955c: 209

Pirata suwaneus Wallace & Exline, 1978: 62, f. 7, 9, 125-133 (Tmf from Allocosa)

Pirata sylvanus Chamberlin & Ivie, 1944 | | USA urn:lsid:nmbe.ch:spidersp:019122

Pirata sylvanus Chamberlin & Ivie, 1944: 151, pl. 13, f. 192-193 (Dmf; N.B.: sylvanus is a Latin noun in apposition) Original description

Pirata sylvanus Wallace & Exline, 1978: 106, f. 12, 233-237 (mf)

Pirata tenuitarsis Simon, 1876 | | Europe to Mongolia urn:lsid:nmbe.ch:spidersp:019126

Pirata tenuitarsis Simon, 1876a: 302, pl. 12, f. 9 (Dmf) Original description

Pirata piraticus moravicus Kratochvíl, 1931: 70, f. 3 (Df) [urn:lsid:nmbe.ch:spidersp:056633]

Lycosa piraticus tenuitarsis Simon, 1937: 1118, 1140

Pirata moravicus Buchar, 1966a: 213, f. 3A-C, E-F (Dm, f, elevated from subspecies)

Pirata moravicus von Helversen & Harms, 1969: 367, f. 1-2 (m)

Pirata tenuitarsis Michelucci & Tongiorgi, 1976: 156, f. 1-2, 4, 6, 8, 10, 12 (elevated from subspecies of P. piraticus, S of Pirata moravicus)

Pirata tenuitarsis Jocqué, 1977e: 145, f. 4, 6 (mf)

Pirata tenuitarsis Kronestedt, 1980b: 66, f. 2B, 3A, C, 4B-C, 6D-F, 7A-C, 8A-D, 9A-B, 10A (mf)

Pirata tenuitarsis van Helsdingen, 1981b: 34, f. 1-2 (mf)

Pirata tenuitarsis Snazell, 1983b: 96, f. 9-11 (mf)

Pirata tenuitarsis Roberts, 1985: 150, f. 66a, c (mf)

Pirata tenuitarsis Heimer & Nentwig, 1991: 346, f. 893 (mf)

Pirata tenuitarsis Logunov, 1992e: 61, f. 6 (mf)

Pirata tenuitarsis Roberts, 1995: 233, f. (mf)

Pirata tenuitarsis Roberts, 1998: 250, f. (mf)

Pirata tenuitarsis Almquist, 2005: 245, f. 240a-e (mf)

Pirata tenuitarsis Nadolny & Kovblyuk, 2011: 188, f. 5, 9, 15, 109-113, 117-123, 125 (mf)

Pirata tenuitarsis Omelko, Marusik & Koponen, 2011: 213, f. 44-45 (f)

Pirata timidus (Lucas, 1846) | | Algeria urn:lsid:nmbe.ch:spidersp:019127

Lycosa timida Lucas, 1846: 117, pl. 3, f. 8 (Df) Original description

Sosilaus timidus Roewer, 1955c: 296 (incertae sedis)

Lycosa timida Bonnet, 1957: 2667

Pirata trepidus Roewer, 1960 | | Namibia urn:lsid:nmbe.ch:spidersp:019128

Pirata trepida Roewer, 1960d: 837, f. 464a-b (Df) Original description

Pirata trepidus Brignoli, 1983c: 457

Pirata triens Wallace & Exline, 1978 | | USA urn:lsid:nmbe.ch:spidersp:019129

Pirata triens Wallace & Exline, 1978: 38, f. 4, 66, 73-76 (Dmf) Original description

Pirata turrialbicus Wallace & Exline, 1978 | | Costa Rica, Panama, Cuba urn:lsid:nmbe.ch:spidersp:019130

Pirata turrialbicus Wallace & Exline, 1978: 57, f. 115-121 (Dmf) Original description

Pirata veracruzae Gertsch & Davis, 1940 | | Mexico urn:lsid:nmbe.ch:spidersp:019133

Pirata veracruzae Gertsch & Davis, 1940a: 8, f. 23 (Dm) Original description

Pirata veracruzae Wallace & Exline, 1978: 62, f. 1, 122-124 (m)

Pirata welakae Wallace & Exline, 1978 | | USA urn:lsid:nmbe.ch:spidersp:019134

Pirata welakae Wallace & Exline, 1978: 26, f. 44-50 (Dmf) Original description

Pirata werneri (Roewer, 1960) | | Morocco urn:lsid:nmbe.ch:spidersp:019135

Piratula werneri Roewer, 1960d: 679, f. 385a-b (Df) Original description

Pirata werneri Brignoli, 1983c: 699

Pirata zavattarii (Caporiacco, 1941) | | Ethiopia urn:lsid:nmbe.ch:spidersp:019138

Trochosa zavattarii Caporiacco, 1941b: 59, f. 9 (Df) Original description

Pirata zavattarii Roewer, 1955c: 286

Pirata zavattarii Roewer, 1960d: 831, f. 460a-b (f)