World Spider Catalog

Agelenidae C. L. Koch, 1837

Family | ACCEPTED

N.B.: Miller et al., 2010: 801 rejected the placement of Coelotes and allied genera by Wunderlich, 1986: 24 as being more closely related to Amaurobius than to Agelena; Coelotinae F. O. Pickard-Cambridge, 1893: 158 were treated as a separate family Coelotidae by Ono, 2008; transfers of Cryphoeca and Tuberta to the Hahniidae and genera Blabomma, Chorizomma, Cicurina and Mastigusa to the Dictynidae by Lehtinen, 1967.

In synonymy:

Asiacoelotes Wang, 2000 = Iwogumoa Kishida, 1955 (Nishikawa & Ono, 2004: 4, contra Yaginuma, in Brignoli, 1983c: 468)

Gen. Iwogumoa Kishida, 1955 urn:lsid:nmbe.ch:spidergen:03777

N.B.: considered a nomen nudum by Yaginuma, in Brignoli, 1983c: 468; treated as valid and a senior synonym of Asiacoelotes Wang, 2002: 31 [urn:lsid:nmbe.ch:spidergen:04775] (type Coelotes xinhuiensis Chen, 1984) by Nishikawa & Ono, 2004: 4.

Transferred to other genera:

Iwogumoa sparus (Dankittipakul, Chami-Kranon & Wang, 2005) -- see Notiocoelotes

In synonymy:

Iwogumoa atrata (Wang, Yin, Peng & Xie, 1990, T from Coelotes) = Iwogumoa xinhuiensis Chen, 1984 (Wang & Ono, 1998: 151, sub Coelotes)

Iwogumoa cincta (Zhu, Song, Gao & Guan, 1993, T from Coelotes) = Iwogumoa dicranata Wang, Yin, Peng & Xie, 1990 (Wang, 2002: 31, sub Asiacoelotes)

Iwogumoa curta (Bösenberg & Strand, 1906, T from Coelotes) = Iwogumoa insidiosa L. Koch, 1878 (Wang, 2002: 32, sub Asiacoelotes)

Iwogumoa jucunda (Chen & Zhao, 1997, T from Coelotes) = Iwogumoa plancyi Simon, 1880 (Zhu, Wang & Zhang, 2017: 441)

Iwogumoa paradicranata (Guan, Song, Zhu & Gao, 1993, T from Coelotes) = Iwogumoa dicranata Wang, Yin, Peng & Xie, 1990 (Wang, 2002: 32, sub Asiacoelotes)

Iwogumoa portus Nishikawa, 2009 = Iwogumoa plancyi Simon, 1880 (Okumura, Koike & Nakano, 2017: 77)

Iwogumoa tropidosata (Wang & Zhu, 1991) = Iwogumoa songminjae Paik & Yaginuma, 1969 (Kim & Lee, 2006a: 50, sub Asiacoelotes)

Iwogumoa yueluensis (Peng, 1999, T from Coelotes) = Iwogumoa pengi Ovtchinnikov, 1999 (Yin et al., 2012: 981, sub Asiacoelotes)

Homonym replaced:

Iwogumoa ovata (Peng & Wang, 1997) -- see Iwogumoa pengi Ovtchinnikov, 1999 Ovtchinnikov, 1999

Iwogumoa acco (Nishikawa, 1987) | | Japan urn:lsid:nmbe.ch:spidersp:022710

Coelotes acco Nishikawa, 1987b: 24, f. 1-8 (Dmf) Original description

Asiacoelotes acco Wang, 2002: 31 (T from Coelotes)

Iwogumoa acco Okumura et al., 2009: 197, f. 366-372 (mf)

Iwogumoa dalianensis Zhang, Zhu & Wang, 2017 | | China urn:lsid:nmbe.ch:spidersp:050001

Iwogumoa dalianensis Zhang, Zhu & Wang in Zhu, Wang & Zhang, 2017: 430, f. 285A-B (Df) Original description

Iwogumoa dalianensis Hu & Zhang, 2019: 52, f. 1A-G, 2 (Dm, f)

Iwogumoa dicranata (Wang, Yin, Peng & Xie, 1990) | | China urn:lsid:nmbe.ch:spidersp:022753

Coelotes dicranatus Wang et al., 1990: 186, f. 25-28 (Dmf) Original description

Coelotes cinctus Zhu, Song, Gao & Guan in Song et al., 1993: 94, f. 2a-d (Dmf) [urn:lsid:nmbe.ch:spidersp:022741]

Coelotes paradicranatus Guan, Zhu, Song & Gao in Song et al., 1993: 96, f. 4a-d (Dmf) [urn:lsid:nmbe.ch:spidersp:022836]

Coelotes cinctus Song, Zhu & Chen, 1999: 374, f. 217G-H, 219B, I (mf)

Coelotes dicranatus Song, Zhu & Chen, 1999: 374, f. 217I-J, 219E, L (mf)

Coelotes paradicranatus Song, Zhu & Chen, 1999: 377, f. 221S-T, 222P, 224D (mf)

Coelotes dicranatus Song, Zhu & Chen, 2001: 290, f. 183A-D (mf)

Asiacoelotes dicranatus Wang, 2002: 31 (T from Coelotes, S of Coelotes cinctus and C. paradicranatus)

Iwogumoa dicranata Zhu, Wang & Zhang, 2017: 431, f. 286A-E (mf)

Iwogumoa ensifer (Wang & Ono, 1998) | | Taiwan urn:lsid:nmbe.ch:spidersp:022758

Coelotes ensifer Wang & Ono, 1998: 143, f. 3-7 (Dmf) Original description

Asiacoelotes ensifer Wang, 2002: 32 (T from Coelotes)

Iwogumoa ensifer Wang & Jäger, 2010: 1176, f. 2B (f)

Iwogumoa ensifer Zhu, Wang & Zhang, 2017: 433, f. 287A-E (mf)

Iwogumoa filamentacea (Tang, Yin & Zhang, 2002) | | China urn:lsid:nmbe.ch:spidersp:038746

Coelotes filamentaceus Tang, Yin & Zhang, 2002: 79, f. 1-3 (Dm) Original description

Coelotes filamentaceus Yin et al., 2012: 987, f. 503a-c (m)

Iwogumoa filamentacea Zhu, Wang & Zhang, 2017: 434, f. 288A-D (m, Df, T from Coelotes)

Iwogumoa illustrata (Wang, Yin, Peng & Xie, 1990) | | China urn:lsid:nmbe.ch:spidersp:022779

Coelotes illustratus Wang et al., 1990: 218, f. 93-97 (Dmf) Original description

Coelotes illustratus Song, Zhu & Chen, 1999: 375, f. 220B-C, 22GH, 223J (mf)

Asiacoelotes illustratus Wang, 2002: 32 (T from Coelotes)

Iwogumoa illustrata Zhu & Zhang, 2011: 323, f. 234A-E (mf)

Asiacoelotes illustratus Yin et al., 2012: 979, f. 497a-e (mf)

Iwogumoa illustrata Zhu, Wang & Zhang, 2017: 436, f. 289A-E (mf)

Iwogumoa insidiosa (L. Koch, 1878) | | Russia (Far East), Korea, China, Japan urn:lsid:nmbe.ch:spidersp:022784

Coelotes insidiosus L. Koch, 1878c: 751, pl. 15, f. 12-13 (Dm) Original description

Coras insidiosus Saito, 1959: 38, f. 15a-c (m)

Coras insidiosus Simon, 1898a: 258

Coras insidiosus Yaginuma, 1960: 93, f. 82.6 (f)

Coras insidiosus Bösenberg & Strand, 1906: 300, 305, pl. 16, f. 458 (mf, S of Coelotes japonicus, rejected by Lehtinen, 1967: 224, see Pireneitega luctuosa)

Coelotes curtus Lehtinen, 1967: 224 (T from Tegenaria)

Tegenaria curta Bösenberg & Strand, 1906: 303, pl. 16, f. 478 (Df) [urn:lsid:nmbe.ch:spidersp:022747]

Coelotes insidiosus Lehtinen, 1967: 224 (T from Coras, S of Coelotes plancyi, revalidated by Song & Hubert, 1983: 4)

Coras insidiosus Saito, 1939: 62, f. 8(1) (m)

Coras insidiosus Shinkai, 1969: 34, f. 18 (m)

Coras insidiosus Yaginuma, 1971: 93, f. 82.6 (f)

Coelotes insidiosus Nishikawa, 1974: 177, f. 20-22 (mf)

Coelotes curtus Nishikawa, 1974: 178, f. 32 (f)

Coelotes insidiosus Nishikawa, 1977a: 34, f. 6-7 (m)

Coelotes insidiosus Shinkai, 1978: 95, f. 37 (m)

Coelotes insidiosus Shimojana, 1982: 80, f. 6-8, 12-13 (mf)

Coelotes insidiosus Irie, 1984: 16, f. 1-6 (mf)

Coelotes insidiosus Yaginuma, 1986a: 148, f. 80.4 (mf)

Coelotes insidiosus Chikuni, 1989b: 100, f. 9 (mf)

Coelotes insidiosus Marusik & Logunov, 1991: 93, f. 1.3 (f)

Asiacoelotes insidiosus Wang, 2002: 32 (T from Coelotes, S of Coelotes curtus)

Coelotes insidiosus Guan, 2002: 5, f. 3.1-5 (mf)

Asiacoelotes insidiosus Kim & Lee, 2006a: 56, f. 5A-F, 6A-B, 7A-H, 8A-H, 9B (mf)

Iwogumoa insidiosa Okumura et al., 2009: 197, f. 378-381 (mf)

Iwogumoa insidiosa Kim & Lee, 2018b: 53, f. 31A-B (f)

Iwogumoa insidiosa Okumura, Honki & Ohba, 2019: 248, f. 1b,d,f (mf)

Iwogumoa interuna (Nishikawa, 1977) | | Russia (Far East), Korea, Japan urn:lsid:nmbe.ch:spidersp:022785

Coelotes interunus Nishikawa, 1977a: 34, f. 1-5 (Dmf) Original description

Coelotes interunus Matsuda, 1986: 89, f. 17-18 (m)

Coelotes interunus Yaginuma, 1986a: 151, f. 80.5 (mf)

Coelotes interunus Chikuni, 1989b: 102, f. 18 (mf)

Coelotes interunus Kim, 1998b: 6, f. 1-3 (m)

Asiacoelotes interunus Wang, 2002: 32 (T from Coelotes)

Iwogumoa interuna Okumura et al., 2009: 197, f. 373-377 (mf)

Iwogumoa interuna Kim & Lee, 2018b: 53, f. 32A-B (f)

Iwogumoa longa (Wang, Tso & Wu, 2001) | | Taiwan urn:lsid:nmbe.ch:spidersp:037405

Coelotes longus Wang, Tso & Wu, 2001: 132, f. 22-23 (Dm) Original description

Asiacoelotes longus Wang, 2002: 32 (T from Coelotes)

Iwogumoa longa Zhu, Wang & Zhang, 2017: 437, f. 290A-B (m)

Iwogumoa montivaga (Wang & Ono, 1998) | | Taiwan urn:lsid:nmbe.ch:spidersp:022819

Coelotes montivagus Wang & Ono, 1998: 146, f. 10-14 (Dmf) Original description

Asiacoelotes montivagus Wang, 2002: 32 (T from Coelotes)

Iwogumoa montivaga Zhu, Wang & Zhang, 2017: 438, f. 291A-E (mf)

Iwogumoa nagasakiensis Okumura, 2007 | | Japan urn:lsid:nmbe.ch:spidersp:041557

Iwogumoa nagasakiensis Okumura, 2007: 89, f. 9-12 (Dmf) Original description

Iwogumoa nagasakiensis Okumura et al., 2009: 197, f. 382-385 (mf)

Iwogumoa pengi (Ovtchinnikov, 1999) | | China urn:lsid:nmbe.ch:spidersp:022844

Coelotes ovatus Peng & Wang, 1997: 330, f. 24-26 (Df; preoccupied by Paik, 1976, see Alloclubionoides ovatus) [urn:lsid:nmbe.ch:spidersp:054304]

Coelotes ovatus Song, Zhu & Chen, 1999: 377, f. 221Q-R (f)

Coelotes pengi Ovtchinnikov, 1999: 79 (replacement name) Original description

Coelotes yueluensis Peng, 1999: 285 (superfluous replacement name) [urn:lsid:nmbe.ch:spidersp:061063]

Asiacoelotes pengi Wang, 2002: 32 (T from Coelotes)

Asiacoelotes pengi Yin et al., 2012: 981, f. 498a-c (f, S of Coelotes yueluensis)

Iwogumoa pengi Zhu, Wang & Zhang, 2017: 440, f. 292A-B (f)

Iwogumoa plancyi (Simon, 1880) | | China, Japan urn:lsid:nmbe.ch:spidersp:022849

Coelotes plancyi Simon, 1880b: 115, pl. 3, f. 12-14 (Dm) Original description

Coras plancyi Simon, 1898a: 258

Coelotes insidiosus Lehtinen, 1967: 448, f. 235 (m, type of Coelotes plancyi)

Coelotes plancyi Song & Hubert, 1983: 4, f. 6-9 (m; removed from S of Coelotes insidiosus, rejecting Lehtinen, 1967: 224)

Coelotes plancyi Song, 1987: 194, f. 152 (m)

Coelotes jucundus Chen & Zhao, 1997: 89, f. 7-8 (Df) [urn:lsid:nmbe.ch:spidersp:022789]

Coelotes jucundus Song, Zhu & Chen, 1999: 376, f. 220L-M (f)

Coelotes plancyi Song, Zhu & Chen, 1999: 377, f. 223B-C (m)

Coelotes plancyi Hu, 2001: 143, f. 53.1-3 (m)

Coelotes plancyi Song, Zhu & Chen, 2001: 293, f. 186A-D (m)

Iwogumoa portus Nishikawa, 2009: 70, f. 111-116 (Dmf) [urn:lsid:nmbe.ch:spidersp:042631]

Iwogumoa portus Okumura et al., 2009: 199, f. 393-398 (mf)

Coelotes plancyi Zhu & Zhang, 2011: 318, f. 230A-E (mf)

Iwogumoa plancyi Okumura, Koike & Nakano, 2017: 77, f. 1A-D (mf, T from Coelotes, S of Iwogumoa portus)

Iwogumoa plancyi Zhu, Wang & Zhang, 2017: 441, f. 10F, 293A-E, pl. IIA (mf, S of Coelotes jucundus)

Iwogumoa plancyi Zhang & Wang, 2017: 25, 4 f. (mf)

Iwogumoa plancyi Jiang, Chen & Zhang, 2018: 78, f. 14A-B, 26N (f)

Iwogumoa plancyi Okumura, 2019c: 128, f. 2 (f)

Iwogumoa songminjae (Paik & Yaginuma, 1969) | | Russia (Far East), China, Korea, Japan (Tsushima Is.) urn:lsid:nmbe.ch:spidersp:022875

Coelotes songminjae Paik & Yaginuma in Paik, Yaginuma & Namkung, 1969: 839, f. 65-69 (Df) Original description

Coelotes songminjae Paik, 1971c: 172, f. 1-6 (Dm)

Coelotes songminjae Paik, 1978e: 351, f. 158.1-8 (mf)

Coelotes songminjae Paik & Namkung, 1979: 56, f. 41a-b (mf)

Coelotes tropidosatus Wang & Zhu, 1991: 4, f. 9-10 (Df) [urn:lsid:nmbe.ch:spidersp:022897]

Coelotes tropidosatus Song, Zhu & Chen, 1999: 388, f. 226A-B (f)

Asiacoelotes songminjae Wang, 2002: 32 (T from Coelotes)

Asiacoelotes tropidosatus Wang, 2002: 33 (T from Coelotes)

Coelotes songminjae Namkung, 2002: 388, f. 28.2a-b (mf)

Coelotes songminjae Namkung, 2003: 390, f. 28.2a-b (mf)

Asiacoelotes songminjae Kim & Lee, 2006a: 50, f. 1A-G, 2A-B, 3A-H, 4A-H, 9A (mf, S of Asiacoelotes tropidosatus)

Iwogumoa songminjae Zhu, Wang & Zhang, 2017: 443, f. 294A-D (mf)

Iwogumoa songminjae Kim & Lee, 2018b: 55, f. 33A-D, pl. 7 (mf)

Iwogumoa songminjae Okumura, Honki & Ohba, 2019: 249, f. 1a,c,e (mf)

Iwogumoa taoyuandong (Bao & Yin, 2004) | | China urn:lsid:nmbe.ch:spidersp:040283

Coelotes taoyuandong Bao & Yin, 2004: 456, f. 4-6 (Df) Original description

Iwogumoa taoyuandong Wang & Jäger, 2007: 46 (T from Coelotes)

Asiacoelotes taoyuandong Yin et al., 2012: 982, f. 499a-c (f)

Iwogumoa taoyuandong Zhu, Wang & Zhang, 2017: 445, f. 295A-B (f)

Iwogumoa tengchihensis (Wang & Ono, 1998) | | Taiwan urn:lsid:nmbe.ch:spidersp:022887

Coelotes tengchihensis Wang & Ono, 1998: 150, f. 20-21 (Df) Original description

Asiacoelotes tengchihensis Wang, 2002: 32 (T from Coelotes)

Iwogumoa tengchihensis Zhu, Wang & Zhang, 2017: 446, f. 296A-B (f)

Iwogumoa xieae Liu & Li, 2008 | | China urn:lsid:nmbe.ch:spidersp:041871

Iwogumoa xieae Liu & Li, 2008b: 458, f. 1-6 (Dmf) Original description

Iwogumoa xieae Zhu, Wang & Zhang, 2017: 447, f. 297A-E (mf)

Iwogumoa xinhuiensis (Chen, 1984) | | China, Taiwan urn:lsid:nmbe.ch:spidersp:022912

Coelotes xinhuiensis Chen, 1984b: 3, f. 9-10 (Df) Original description

Coelotes atratus Wang et al., 1990: 227, f. 115-116 (Df) [urn:lsid:nmbe.ch:spidersp:061062]

Coelotes xinhuiensis Wang & Ono, 1998: 151, f. 22-28 (Dm, f, S of Coelotes atratus)

Coelotes xinhuiensis Song, Zhu & Chen, 1999: 388, f. 216N-O, 226M-N (mf)

Asiacoelotes xinhuiensis Wang, 2002: 33, f. 70-85 (mf, T from Coelotes)

Asiacoelotes xinhuiensis Yin et al., 2012: 983, f. 500a-c (m)

Iwogumoa xinhuiensis Zhu, Wang & Zhang, 2017: 448, f. 298A-E (mf)

Iwogumoa yaeyamensis (Shimojana, 1982) | | Japan urn:lsid:nmbe.ch:spidersp:022913

Coelotes yaeyamensis Shimojana, 1982: 76, f. 1-5, 9-11 (Dmf) Original description

Coelotes yaeyamensis Yaginuma, 1986a: 151, f. 80.6 (mf)

Asiacoelotes yaeyamensis Wang, 2002: 33 (T from Coelotes)

Iwogumoa yaeyamensis Okumura et al., 2009: 199, f. 386-392 (mf)

Iwogumoa yaeyamensis Suzuki, 2022c: 202, f. 10 (f)

Iwogumoa yushanensis (Wang & Ono, 1998) | | Taiwan urn:lsid:nmbe.ch:spidersp:022918

Coelotes yushanensis Wang & Ono, 1998: 154, f. 29-30 (Df) Original description

Asiacoelotes yushanensis Wang, 2002: 33 (T from Coelotes)

Iwogumoa yushanensis Zhu, Wang & Zhang, 2017: 450, f. 299A-B (f)