World Spider Catalog

Linyphiidae Blackwall, 1859

Family | ACCEPTED

N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; Tian et al., 2022 show five subfamilies: Linyphiinae (incl. Mynogleninae Lehtinen, 1967: 250 and Dubiaraneinae Millidge, 1993c: 145), Erigoninae Emerton, 1882: 7, Micronetinae Hull, 1920a: 8, Ipainae Saaristo, 2007b: 34 and Stemonyphantinae Wunderlich, 1986: 120; da Silva-Moreira, Kulkarni & Hormiga, 2025: 60-62 list resp. newly circumscribe six subfamilies: Erigoninae, Lepthyphantinae Simon, 1929: 562, Linyphiinae (syn. nov.: Dubiaraneinae), Mynogleninae, Pocobletinae Silva-Moreira & Hormiga, 2025: 61 (new subfamily) and Stemonyphantinae, state Ipainae and Micronetinae to be polyphyletic, leave several linyphiid clades unranked and also several genera as incertae sedis.

In synonymy:

Myagrus Templeton, 1864 = Saaristoa Millidge, 1978 (Millidge, 1978b: 123, nomen oblitum)

Gen. Saaristoa Millidge, 1978 urn:lsid:nmbe.ch:spidergen:01236

type Neriene abnormis Blackwall, 1841; gender feminine

N.B.: Myagrus Templeton, in Blackwall, 1864a: 283 [urn:lsid:nmbe.ch:spidergen:06400] (type Neriene pilosa Templeton, 1864 = Saaristoa abnormis (Blackwall, 1841)) was described as subgenus of Neriene Blackwall, 1833, considered a senior synyonm (nomen oblitum) of Oreonetides Strand, 1901 by Bonnet, 1957: 3201 and considered a synonym of Saaristoa by Millidge, 1978b: 123 (by transfer of the type species); the synonymy of Saaristoa with Aphileta Hull, 1920, as proposed by Breitling, 2019b: 27 and Breitling, 2021: 20, is not accepted here, as it is based mainly on COI, see FAQ 7); not included in a subfamily of Linyphiidae but in an unranked linyphiid clade by da Silva-Moreira, Kulkarni & Hormiga, 2025: 62.

In synonymy:

Saaristoa decipiens (O. Pickard-Cambridge, 1879, T from Linyphia) = Saaristoa abnormis (Blackwall, 1841) (Bristowe, 1941: 514, sub Oreonetides)

Saaristoa douglasi (O. Pickard-Cambridge, 1875, T from Neriene) = Saaristoa abnormis (Blackwall, 1841) (Simon, 1884a: 386, sub Tmeticus)

Saaristoa higoensis (Saito, 1984) = Saaristoa ebinoensis (Oi, 1979) (Ono, Matsuda & Saito, 2009: 323)

Saaristoa icterica (Thorell, 1875, T from Lepthyphantes) = Saaristoa abnormis (Blackwall, 1841) (Holm, 1977c: 25, sub Oreonetides)

Saaristoa linguata (O. Pickard-Cambridge, 1873) = Saaristoa abnormis (Blackwall, 1841) (O. Pickard-Cambridge, 1881a: 578, sub Linyphia)

Saaristoa pilosa (Templeton, 1864, T from Neriene) = Saaristoa abnormis (Blackwall, 1841) (Hull, 1920b: 31, sub Centromerus)

Saaristoa pusilla (Schenkel, 1923) = Saaristoa firma (O. Pickard-Cambridge, 1906) (Schenkel, 1925d: 282, sub Macrargus)

Saaristoa spelaea (Simon, 1872, T from Erigone) = Saaristoa abnormis (Blackwall, 1841) (Simon, 1884a: 386, sub Tmeticus)

Saaristoa abnormis (Blackwall, 1841) | | Europe urn:lsid:nmbe.ch:spidersp:012483 TYPE SPECIES

Neriene abnormis Blackwall, 1841a: 649 (Dmf) Original description

Argus abnormis Walckenaer, 1847a: 512

Neriene abnormis Blackwall, 1864a: 286, pl. 19, f. 200 (mf)

Neriene pilosa Templeton in Blackwall, 1864a: 283, pl. 19, f. 196 (Df) [urn:lsid:nmbe.ch:spidersp:063692]

Erigone spelaea Simon, 1872b: 474, pl. 16, f. 8-9 (Dmf) [urn:lsid:nmbe.ch:spidersp:063693]

Linyphia linguata O. Pickard-Cambridge, 1873c: 537, pl. 46, f. 8 (Df) [urn:lsid:nmbe.ch:spidersp:063694]

Linyphia icterica Thorell, 1875a: 83 (Dmf) [urn:lsid:nmbe.ch:spidersp:063691]

Linyphia icterica Thorell, 1875c: 21 (mf)

Erigone douglasii O. Pickard-Cambridge, 1875e: 247, pl. 8, f. 5 (Df) [urn:lsid:nmbe.ch:spidersp:063695]

Linyphia abnormis O. Pickard-Cambridge, 1879d: 207

Linyphia decipiens O. Pickard-Cambridge, 1879d: 208 (Df) [urn:lsid:nmbe.ch:spidersp:063696]

Neriene douglasii O. Pickard-Cambridge, 1881a: 488

Linyphia abnormis O. Pickard-Cambridge, 1881a: 578 (S of Linyphia linguata)

Helophora abnormis Förster & Bertkau, 1883: 254

Lepthyphantes ictericus Simon, 1884a: 330

Tmeticus abnormis Simon, 1884a: 386, f. 160-162 (mf, S of Erigone spelaea and Neriene douglasi)

Tmeticus decipiens Simon, 1884a: 420

Tmeticus abnormis Becker, 1896: 61, pl. 8, f. 3 (mf)

Lepthyphantes abnormis Bösenberg, 1901: 73, pl. 4, f. 75 (mf)

Centromerus abnormis Lessert, 1904: 335

Macrargus abnormis Reimoser, 1919: 81

Centromerus abnormis Hull, 1920b: 31 (S of Neriene pilosa)

Oreonetides abnormis Simon, 1929: 561, 725

Neriene pilosa Bristowe, 1941: 525 (identity uncertain, probably S of Oreonetides abnormis)

Oreonetides abnormis Bristowe, 1941: 514 (S of Linyphia decipiens)

Oreonetides abnormis Locket & Millidge, 1953: 360, f. 216A-B (mf)

Neriene pilosa Roewer, 1955c: 1603 (nicht zu deuten = nomen dubium)

Oreonetides abnormis Wiehle, 1956: 83, f. 134-137 (mf)

Oreonetides abnormis Merrett, 1963b: 366, f. 20A-D (m)

Oreonetides abnormis Miller, 1971: 240, pl. XLVIII, f. 6-7, pl. LXI, f. 26 (mf)

Oreonetides abnormis Kritscher, 1972: 89, f. 1-3 (mf)

Oreonetides abnormis Palmgren, 1975: 21, f. 3.18-19 (mf)

Saaristoa abnormis Millidge, 1977: 48, f. 194 (m; in generic nomen nudum)

Oreonetides abnormis Holm, 1977c: 25 (S of Lepthyphantes ictericus)

Saaristoa abnormis Millidge, 1978b: 123 (T from Oreonetides)

Oreonetides abnormis Pakhorukov, 1981: 79, f. 25-26 (mf)

Oreonetides abnormis Thaler, 1981a: 144, f. 7 (f)

Saaristoa abnormis Millidge, 1984b: 246, f. 72-73, 76 (f)

Saaristoa abnormis Roberts, 1987: 134, f. 68b (mf)

Saaristoa abnormis Heimer & Nentwig, 1991: 242, f. 649 (mf)

Saaristoa abnormis Agnarsson, 1996: 141, f. 155A-B (mf)

Saaristoa ebinoensis (Oi, 1979) | | Japan urn:lsid:nmbe.ch:spidersp:011150

Lepthyphantes ebinoensis Oi, 1979: 334, f. 19-21 (Dmf) Original description

Centromerus higoensis Saito, 1984: 5, f. 5-6 (Dmf) [urn:lsid:nmbe.ch:spidersp:012485]

Centromerus higoensis Irie & Saito, 1987: 22, f. 19-20 (mf)

Saaristoa higoensis Eskov, 1991a: 45 (T from Centromerus)

Saaristoa ebinoensis Ono, Matsuda & Saito, 2009: 322, f. 977-981 (mf, T from Lepthyphantes, S of Saaristoa higoensis)

Saaristoa ebinoensis Hino, 2024: 56, f. 2P-Q (mf)

Saaristoa ebinoensis Hirotsu, 2024b: 42, f. 12-14 (mf)

Saaristoa firma (O. Pickard-Cambridge, 1906) | | Europe urn:lsid:nmbe.ch:spidersp:012484

Tmeticus firmus O. Pickard-Cambridge, 1906a: 59, pl. A, f. 13a-c (Dmf) Original description

Centromerus firmus O. Pickard-Cambridge, 1909a: 174

Macrargus pusillus Schenkel, 1923: 99, pl. 7, f. 12-13 (Dmf) [urn:lsid:nmbe.ch:spidersp:063696]

Macrargus firmus Schenkel, 1925d: 282 (S of Macrargus pusillus)

Oreonetides firmus Simon, 1929: 560, 725, f. 827 (mf)

Oreonetides firmus Locket & Millidge, 1953: 360, f. 216C-D (mf)

Oreonetides firmus Wiehle, 1956: 85, f. 138-140 (mf)

Oreonetides firmus Casemir, 1963b: 131, pl. 1, f. 6, pl. 2, f. 1-4 (mf)

Oreonetides firmus Miller, 1971: 240, pl. 48, f. 1-2 (mf)

Saaristoa firma Millidge, 1978b: 123 (T from Oreonetides)

Oreonetides firmus Weiss, Varvara & Sarbu, 1979: 256, f. 1 (f)

Oreonetides firmus Thaler, 1981a: 144, f. 8, 16-17 (mf)

Saaristoa firma Roberts, 1987: 134, f. 68c (mf)

Saaristoa firma Heimer & Nentwig, 1991: 242, f. 648 (mf)

Saaristoa firma Komnenov, 2020b: 34, f. 1-2 (mf)

Saaristoa nipponica (Saito, 1984) | | Japan urn:lsid:nmbe.ch:spidersp:012486

Centromerus nipponicus Saito, 1984: 6, f. 7-8 (Dmf) Original description

Saaristoa nipponica Eskov, 1991a: 45 (T from Centromerus)

Saaristoa nipponica Ono, Matsuda & Saito, 2009: 323, f. 982-984 (mf)

Saaristoa sammamish (H. W. Levi & L. R. Levi, 1955) | | Alaska, Canada, USA urn:lsid:nmbe.ch:spidersp:012487

Lepthyphantes sammamish Levi & Levi, 1955: 33, f. 11-13 (Dmf) Original description

Saaristoa sammamish Crawford, 1988: 20 (T from Lepthyphantes)