World Spider Catalog

Linyphiidae Blackwall, 1859

Family | ACCEPTED

N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; other subfamilies are Erigoninae Emerton, 1882: 7 and Mynogleninae Lehtinen, 1967: 250.

In synonymy:

Trachygnatha Kulczyński, 1894 = Gnathonarium Karsch, 1881 (Simon, 1926: 532)

Gen. Gnathonarium Karsch, 1881 urn:lsid:nmbe.ch:spidergen:00989

N.B.: considered a senior synoynm of Trachygnatha Kulczyński, in Chyzer & Kulczyński, 1894: 91 [urn:lsid:nmbe.ch:spidergen:05451] (type Theridion dentatum Wider, 1834) by Simon, 1926: 532.

In synonymy:

Gnathonarium cambridgei Schenkel, 1963 = Gnathonarium taczanowskii O. Pickard-Cambridge, 1873 (Tanasevitch, 2006b: 303)

Gnathonarium columbianum (Emerton, 1923) = Gnathonarium taczanowskii O. Pickard-Cambridge, 1873 (Tanasevitch, 2013b: 177)

Gnathonarium cornigerum Zhu & Wen, 1980 = Gnathonarium taczanowskii O. Pickard-Cambridge, 1873 (Tu & Li, 2004b: 856, sub G. cambridgei)

Gnathonarium famelicum (Keyserling, 1886) = Gnathonarium suppositum Kulczyński, 1885 (Eskov, 1988e: 105)

Gnathonarium flavidum Gao & Zhu, 1993 = Gnathonarium taczanowskii O. Pickard-Cambridge, 1873 (Tu & Li, 2004b: 856, sub G. cambridgei)

Gnathonarium phragmigerum Gao & Zhu, 1988 = Gnathonarium taczanowskii O. Pickard-Cambridge, 1873 (Tu & Li, 2004b: 856, sub G. cambridgei)

Gnathonarium sibirianum (Keyserling, 1886, T from Erigone) = Gnathonarium suppositum Kulczyński, 1885 (Eskov & Marusik, 1994: 67)

Gnathonarium biconcavum Tu & Li, 2004 | | China urn:lsid:nmbe.ch:spidersp:039895

Gnathonarium biconcavum Tu & Li, 2004b: 854, f. 1A-I (Dmf) Original description

Gnathonarium dentatum (Wider, 1834) | | Europe, North Africa, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, Iran, Central Asia, China, Korea, Japan urn:lsid:nmbe.ch:spidersp:010645

Theridion dentatum Wider, 1834: 223, pl. 15, f. 8 (Dmf) Original description

Argus dentatum Walckenaer, 1841: 354

Erigone dentata Westring, 1861: 262

Neriene dentata Blackwall, 1864a: 258, pl. 18, f. 174 (mf)

Tmeticus dentatus Menge, 1868: 187, pl. 35, f. 87 (mf)

Tmeticus cristatus Menge, 1868: 189, pl. 36, f. 88 (Dm)

Gnathonarium rohlfsianum Karsch, 1881a: 10, pl. 1, f. 7 (Dmf)

Gongylidium dentatum Simon, 1884a: 492, f. 276-279 (mf)

Trachygnatha dentata Chyzer & Kulczyński, 1894: 91, pl. 3, f. 41 (mf)

Gongylidium dentatum Becker, 1896: 88, pl. 9, f. 8 (mf)

Gongylidium dentatum Smith, 1901b: 108, p. 9: 1b-e (m)

Tmeticus dentatus Bösenberg, 1902: 166, pl. 15, f. 225 (mf)

Oedothorax dentatus Bösenberg & Strand, 1906: 165, pl. 12, f. 273 (mf)

Gnathonarium dentatum Simon, 1926: 476, 532, f. 812 (mf)

Micryphantes dentatus Hull, 1933: 108

Gnathonarium dentatum Bishop & Crosby, 1935c: 222, pl. 18, f. 14-16 (mf)

Gnathonarium dentatum Locket & Millidge, 1953: 217, f. 134D-G (mf)

Gnathonarium dentatum Tullgren, 1955: 363, f. 65a-c (mf)

Gnathonarium dentatum Oi, 1960a: 147, f. 39-43 (mf)

Gnathonarium dentatum Wiehle, 1960a: 371, f. 676-682 (mf)

Gnathonarium dentatum Merrett, 1963b: 442, f. 124A-B (m)

Gnathonarium dentatum Miller, 1971: 261, pl. LIII, f. 19-20 (mf)

Gnathonarium dentatum Yaginuma, 1971: 45, f. 43.5-6 (m)

Gnathonarium dentatum Tystshenko, 1971: 248, f. 897 (f)

Gnathonarium dentatum Namkung, Paik & Yoon, 1972: 92, f. 5 (f)

Gnathonarium dentatum Palmgren, 1976: 69, f. 5.5-8 (mf)

Gnathonarium dentatum Millidge, 1977: 40, f. 164 (m)

Gnathonarium dentatum Song et al., 1977b: 37, f. 5A-F (mf)

Gnathonarium dentatum Paik & Namkung, 1979: 34, f. 16.a-2 (mf)

Gnathonarium dentatum Song, 1980: 155, f. 83a-f (mf)

Gnathonarium dentatum Millidge, 1980a: 101, f. 10 (m)

Gnathonarium dentatum Saito, 1983a: 3, f. 1-3 (f)

Gnathonarium dentatum Hu, 1984: 194, f. 203.1-4 (mf)

Gnathonarium dentatum Guo, 1985: 105, f. 2-47.1-3 (mf)

Gnathonarium dentatum Zhu & Shi, 1985: 115, f. 101a-e (mf)

Gnathonarium dentatum Yaginuma, 1986a: 80, f. 43.2 (f)

Gnathonarium dentatum Song, 1987: 150, f. 111 (mf)

Gnathonarium dentatum Roberts, 1987: 42, f. 12c (mf)

Gnathonarium dentatum Zhang, 1987: 127, f. 105.1-5 (mf)

Gnathonarium dentatum Tanasevitch, 1990: 102, f. 22.22, 28.7, 24.50 (mf)

Gnathonarium dentatum Feng, 1990: 134, f. 109.1-5 (mf)

Gnathonarium dentatum Chen & Gao, 1990: 108, f. 135a-b (mf)

Gnathonarium dentatum Chen & Zhang, 1991: 176, f. 174.1-6 (mf)

Gnathonarium dentatum Heimer & Nentwig, 1991: 162, f. 440 (mf)

Gnathonarium dentatum Song, Zhu & Li, 1993: 860, f. 14A-D (mf)

Gnathonarium dentatum Zhao, 1993: 183, f. 84a-c (mf)

Gnathonarium dentatum Millidge, 1993c: 147, f. 30 (f)

Gnathonarium dentatum Zhao, 1995: 899, f. 421a-c (mf)

Gnathonarium dentatum Song, Zhu & Chen, 1999: 169, f. 96A, L (mf)

Gnathonarium dentatum Hu, 2001: 544, f. 365.1-4 (mf)

Gnathonarium dentatum Song, Zhu & Chen, 2001: 129, f. 68A-F (mf)

Gnathonarium dentatum Namkung, 2002: 187, f. 17.39a-b (mf)

Gnathonarium dentatum Namkung, 2003: 189, f. 17.39a-b (mf)

Gnathonarium dentatum Guryanova, 2003: 5, f. 8 (f)

Gnathonarium dentatum Tu & Li, 2004b: 859, f. 3A-G (mf)

Gnathonarium dentatum Bosmans, 2007: 126, f. 32-37 (mf)

Gnathonarium dentatum Lee, Kang & Kim, 2009: 126, f. 25a-c (mf)

Gnathonarium dentatum Ono, Matsuda & Saito, 2009: 294, f. 500-503 (mf)

Gnathonarium dentatum Zhu & Zhang, 2011: 124, f. 77A-F (mf)

Gnathonarium dentatum Yin et al., 2012: 489, f. 225a-e (mf)

Gnathonarium dentatum Tanasevitch, 2013b: 173, f. 7-12, 22-27, 38-94 (mf)

Gnathonarium dentatum Zhang, Peng & Zhang, 2022: 57, f. 34A-H (mf)

Gnathonarium dentatum Irfan, Zhang & Peng, 2022: 87, f. 100A-F, 101A-F (mf)

Gnathonarium dentatum Palmer, 2023: 11, f. 1A-B, 3-8 (f)

Gnathonarium dentatum orientale (O. Pickard-Cambridge, 1872) | | Israel urn:lsid:nmbe.ch:spidersp:010646

Erigone dentata orientalis O. Pickard-Cambridge, 1872: 290 (Dmf) Original description

Gnathonarium exsiccatum (Bösenberg & Strand, 1906) | | Japan urn:lsid:nmbe.ch:spidersp:010647

Oedothorax exsiccatus Bösenberg & Strand, 1906: 166, pl. 12, f. 265 (Dmf) Original description

Gnathonarium exsiccatus Yaginuma, 1960: 45, f. 43.5 (removed mf from S of G. dentatum)

Gnathonarium exsiccatum Saito, 1983a: 3, f. 4-6 (f)

Gnathonarium exsiccatum Yaginuma, 1986a: 80, f. 43.1 (mf)

Gnathonarium exiccatum Chikuni, 1989b: 58, f. 55 (mf)

Gnathonarium exsiccatum Gao & Zhu, 1990: 153, f. 9-13 (mf, misidentified per Tu & Li, 2004b: 851)

Gnathonarium exsiccatum Song, Zhu & Chen, 1999: 169, f. 96B-C, M-N (mf, misidentified per Tu & Li, 2004b: 851)

Gnathonarium exsiccatum Ono, Matsuda & Saito, 2009: 294, f. 504-509 (mf)

Gnathonarium gibberum Oi, 1960 | | Russia (South Siberia), Nepal, China, Korea, Japan urn:lsid:nmbe.ch:spidersp:010649

Gnathonarium gibberum Oi, 1960a: 149, f. 44-49 (Dmf) Original description

Gnathonarium gibberum Song et al., 1977b: 37, f. 6A-E (mf)

Gnathonarium gibberum Paik & Namkung, 1979: 35, f. 17.1-3 (mf)

Gnathonarium gibberum Song, 1980: 158, f. 84a-e (mf)

Gnathonarium gibberum Wang, 1981: 109, f. 50A-C (f)

Gnathonarium gibberum Saito, 1983a: 4, f. 7-9 (f)

Gnathonarium gibberum Hu, 1984: 194, f. 204.1-4 (mf)

Gnathonarium gibberum Guo, 1985: 106, f. 2-48.1-2 (f)

Gnathonarium gibberum Yaginuma, 1986a: 80, f. 43.3 (f)

Gnathonarium gibberum Song, 1987: 152, f. 112 (mf)

Gnathonarium gibberum Zhang, 1987: 128, f. 106.1-3 (mf)

Gnathonarium gibberum Feng, 1990: 135, f. 110.1-5 (mf)

Gnathonarium gibberum Chen & Gao, 1990: 109, f. 136a-b (mf)

Gnathonarium gibberum Chen & Zhang, 1991: 177, f. 175.1-5 (mf)

Gnathonarium gibberum Song, Zhu & Li, 1993: 860, f. 15A-E (mf)

Gnathonarium gibberum Zhao, 1993: 185, f. 85a-d (mf)

Gnathonarium gibberum Barrion & Litsinger, 1994: 322, f. 1685-1688 (mf)

Gnathonarium gibberum Zhao, 1995: 901, f. 422a-d (mf)

Gnathonarium gibberum Song, Zhu & Chen, 1999: 170, f. 96F-G, 97A (mf)

Gnathonarium gibberum Song, Zhu & Chen, 2001: 131, f. 69A-E (mf)

Gnathonarium gibberum Namkung, 2002: 188, f. 17.40a-b (mf)

Gnathonarium gibberum Namkung, 2003: 190, f. 17.40a-b (mf)

Gnathonarium gibberum Tu & Li, 2004b: 861, f. 4A-G (mf)

Gnathonarium gibberum Lee, Kang & Kim, 2009: 126, f. 26a-c (mf)

Gnathonarium gibberum Ono, Matsuda & Saito, 2009: 294, f. 510-515 (mf)

Gnathonarium gibberum Zhu & Zhang, 2011: 125, f. 78A-E (mf)

Gnathonarium gibberum Yin et al., 2012: 491, f. 226a-e (mf)

Gnathonarium gibberum Fu, 2018: 37, f. 5-4.A-C, pl. 8 (f)

Gnathonarium gibberum Irfan, Zhang & Peng, 2022: 88, f. 102A-F, 103A-F (mf)

Gnathonarium luzon Tanasevitch, 2017 | | Philippines (Luzon) urn:lsid:nmbe.ch:spidersp:049080

Gnathonarium luzon Tanasevitch, 2017a: 332, f. 4-5, 26-29 (Dmf) Original description

Gnathonarium suppositum (Kulczyński, 1885) | | Russia (Middle Siberia to Far East), USA (Alaska), Canada urn:lsid:nmbe.ch:spidersp:010651

Gongylidium suppositum Kulczyński, 1885: 36, pl. 10, f. 15 (Df) Original description

Erigone famelica Keyserling, 1886b: 186, pl. 17, f. 246 (Dm) [urn:lsid:nmbe.ch:spidersp:058988]

Erigone sibiriana Keyserling, 1886b: 189, pl. 17, f. 249 (Df) [urn:lsid:nmbe.ch:spidersp:058989]

Erigone famelica Banks, 1900b: 480, pl. 29, f. 7-8 (m, Df)

Gnathonarium famelicum Bishop & Crosby, 1935c: 224, pl. 18, f. 17-18 (m)

Erigone sibiriana Roewer, 1955c: 1502 (nicht zu deuten = nomen dubium)

Gnathonarium famelicum Holm, 1960b: 117, f. 9 (f)

Gnathonarium suppositum Eskov, 1988e: 105 (S of Gnathonarium famelicum and Gongylidium columbianum, the latter rejected by Tanasevitch, 2013b: 177, see Gnathonarium taczanowskii)

Gnathonarium suppositum Eskov & Marusik, 1994: 67 (S of Erigone sibiriana, removed from nomen dubium)

Gnathonarium famelicum Buckle et al., 2001: 120 (regarded as valid)

Gnathonarium suppositum Tanasevitch, 2013b: 173, f. 1-6, 13-21, 28-37 (mf, S of Gnathonarium famelicum confirmed)

Gnathonarium taczanowskii (O. Pickard-Cambridge, 1873) | | Russia (Urals to Far East), Kazakhstan, Mongolia, China, USA (Alaska), Canada urn:lsid:nmbe.ch:spidersp:039402

Erigone taczanowskii O. Pickard-Cambridge, 1873f: 443, pl. 41, f. 10 (Dm) Original description

Erigone taczanowskii L. Koch, 1879c: 50, pl. 2, f. 5 (m, Df)

Gongylidium taczanowskii Simon, 1884a: 499

Gongylidium columbianum Emerton, 1923: 238, f. 2a-c (Dm) [urn:lsid:nmbe.ch:spidersp:058990]

Gnathonarium cambridgei Schenkel, 1963: 114, f. 67 (Df) [urn:lsid:nmbe.ch:spidersp:010643]

Gnathonarium cornigerum Zhu & Wen, 1980: 19, f. 2A-F (Dmf) [urn:lsid:nmbe.ch:spidersp:010644]

Gnathonarium cornigerum Hu, 1984: 193, f. 202.1-6 (mf)

Gnathonarium cornigerum Zhu & Shi, 1985: 114, f. 100a-f (mf)

Gnathonarium cornigerum Zhang, 1987: 126, f. 104.1-4 (mf)

Gnathonarium phragmigerum Gao & Zhu, 1988: 350, f. 1-7 (Dmf) [urn:lsid:nmbe.ch:spidersp:010650]

Gnathonarium phragmigerum Song, Zhu & Li, 1993: 860, f. 16A-E (mf)

Gnathonarium flavidum Gao & Zhu, 1993: 28, f. 7-13 (Dmf) [urn:lsid:nmbe.ch:spidersp:010648]

Gnathonarium cornigerum Song, Zhu & Chen, 1999: 169, f. 95N-O, 96J-K (mf)

Gnathonarium flavidum Song, Zhu & Chen, 1999: 169, f. 96D-E, O-P (mf)

Gnathonarium phragmigerum Song, Zhu & Chen, 1999: 170, f. 96H-I, 97B-C (mf)

Gnathonarium cornigerum Hu, 2001: 543, f. 364.1-5 (mf)

Gnathonarium cornigerum Song, Zhu & Chen, 2001: 128, f. 67A-F (mf)

Gnathonarium cambridgei Tu & Li, 2004b: 856, f. 2A-G (mf, S of Gnathonarium cornigerum, G. flavidum and G. phragmigerum)

Gnathonarium taczanowskii Tanasevitch, 2006b: 303 (S of Gnathonarium cambridgei)

Gnathonarium taczanowskii Zhu & Zhang, 2011: 126, f. 79A-F (mf)

Gnathonarium taczanowskii Yin et al., 2012: 492, f. 227a-e (mf)

Gnathonarium taczanowskii Tanasevitch, 2013b: 177, f. 95-104 (S of Gongylidium = Gnathonarium columbianum, removed from S of Gnathonarium suppositum, rejecting Eskov, 1988e: 105)

Gnathonarium taczanowskii Zhang, Peng & Zhang, 2022: 59, f. 35A-G (mf)

Gnathonarium taczanowskii Irfan, Zhang & Peng, 2022: 89, f. 104A-C (f)