World Spider Catalog

Linyphiidae Blackwall, 1859

Family | ACCEPTED

N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; Tian et al., 2022 show five subfamilies: Linyphiinae (incl. Mynogleninae Lehtinen, 1967: 250 and Dubiaraneinae Millidge, 1993c: 145), Erigoninae Emerton, 1882: 7, Micronetinae Hull, 1920a: 8, Ipainae Saaristo, 2007b: 34 and Stemonyphantinae Wunderlich, 1986: 120; da Silva-Moreira, Kulkarni & Hormiga, 2025: 60-62 list resp. newly circumscribe six subfamilies: Erigoninae, Lepthyphantinae Simon, 1929: 562, Linyphiinae (syn. nov.: Dubiaraneinae), Mynogleninae, Pocobletinae Silva-Moreira & Hormiga, 2025: 61 (new subfamily) and Stemonyphantinae, state Ipainae and Micronetinae to be polyphyletic, leave several linyphiid clades unranked and also several genera as incertae sedis.

In synonymy:

Frontina Simon, 1884 = Floronia Simon, 1887 (Simon, 1887d: 158, preoccupied, replacement name)

Gen. Floronia Simon, 1887 urn:lsid:nmbe.ch:spidergen:00979

type Araneus bucculentus Clerck, 1757; gender feminine

N.B.: replacement name for Frontina Simon, 1884a: 205 [urn:lsid:nmbe.ch:spidergen:05270], preoccupied in Diptera by Meigen, 1838; considered valid by Millidge, 1984b: 246 and Merrett, Locket & Millidge, 1985: 399, contra Millidge, 1977: 45, who considered it a junior synonym of Tapinopa Westring, 1851.

In synonymy:

Floronia albomaculata (Ohlert, 1867) = Floronia bucculenta (Clerck, 1757) (Thorell, 1870a: 54, sub Linyphia frenata)

Floronia elegans (Walckenaer, 1841, T from Linyphia) = Floronia bucculenta (Clerck, 1757) (Simon, 1929: 726)

Floronia frenata (Wider, 1834) = Floronia bucculenta (Clerck, 1757) (ICZN, 1959: 89)

Floronia pallida (C. L. Koch, 1836, T from Theridion) = Floronia bucculenta (Clerck, 1757) (Thorell, 1870a: 54, sub Linyphia frenata)

Floronia sapporensis (Saito, 1934, T from Neriene) = Floronia exornata (L. Koch, 1878) (Ono, 2009a: 589)

Nomen dubium:

Floronia fagei Caporiacco, 1955: 322, f. 18a-e (mf, Venezuela) -- Miller, 2007a: 259

Floronia annulipes Berland, 1913 | | Ecuador urn:lsid:nmbe.ch:spidersp:010611

Floronia annulipes Berland, 1913b: 88, pl. 8, f. 21-23 (Df) Original description

Floronia bucculenta (Clerck, 1757) | | Europe, Russia (Europe to Far East), China urn:lsid:nmbe.ch:spidersp:010612 TYPE SPECIES

Araneus bucculentus Clerck, 1757: 63, pl. 4, f. 1 (Df) Original description

Araneus bucculentus Martyn, 1793: 29, pl. 6, f. 2 (f)

Linyphia frenata Wider, 1834: 262, pl. 18, f. 4a-c (Dmf) [urn:lsid:nmbe.ch:spidersp:054327]

Theridion pallidum C. L. Koch, 1836a: 64, pl. 94, f. 216 (Df) [urn:lsid:nmbe.ch:spidersp:054328]

Linyphia elegans Walckenaer, 1841: 258 (Df) [urn:lsid:nmbe.ch:spidersp:054329]

Linyphia pallida Blackwall, 1843: 126 (Dm)

Linyphia frenata Westring, 1861: 110 (f)

Linyphia frenata Blackwall, 1864a: 228, pl. 16, f. 151 (mf)

Bolyphantes frenatus Menge, 1866: 137, pl. 26, f. 57 (mf)

Linyphia albomaculata Ohlert, 1867: 81 (Df) [urn:lsid:nmbe.ch:spidersp:054330]

Linyphia frenata Thorell, 1870a: 54 (S of Linyphia albomaculata and Theridion pallidum)

Bolyphantes bucculentus Dahl, 1883: 50

Frontina bucculenta Simon, 1884a: 207

Floronia bucculenta Simon, 1887d: 158

Floronia frenata Chyzer & Kulczyński, 1894: 51, pl. 2, f. 17 (mf)

Floronia bucculenta Simon, 1894a: 708, f. 779, 781, 783 (mf)

Frontina bucculenta Becker, 1896: 10, pl. 1, f. 5 (mf)

Bolyphantes frenatus Bösenberg, 1901: 62, pl. 5, f. 60 (mf)

Frontina frenata Lessert, 1910a: 823

Floronia bucculenta Fage, 1919: 78, f. 31 (m)

Floronia bucculenta Simon, 1929: 566, 726, f. 834-835 (mf, S of Linyphia elegans)

Floronia bucculenta Locket & Millidge, 1953: 373, f. 223A-F (mf)

Floronia bucculenta Wiehle, 1956: 146, f. 249-256 (mf)

Bolyphantes frenatus Saito, 1959: 78, f. 80a-e (f)

[Floronia bucculenta] ICZN, 1959: 89 (S of Floronia frenata, Clerckian names validated)

Floronia bucculenta Oi, 1960a: 193 (S of Floronia exornata [revalidated by Saaristo, 1996a: 6], mf f. 225-230 see F. exornata)

Floronia bucculenta Merrett, 1963b: 364, f. 18A-E (m)

Floronia bucculenta Wiehle, 1967b: 187, f. 23-24 (f)

Floronia bucculenta Tystshenko, 1971: 227, f. 644 (f)

Floronia bucculenta Miller, 1971: 216, pl. XLI, f. 4-7 (mf)

Floronia bucculenta Palmgren, 1975: 34, f. 7.6-7 (mf)

Floronia bucculenta Millidge, 1984b: 246, f. 100 (f)

Floronia bucculenta Roberts, 1987: 142, f. 72b-c (mf)

Floronia bucculenta Izmailova, 1989: 71, f. 49 (f)

Floronia bucculenta Tanasevitch, 1990: 87, f. 13.8-9, 14.21 (mf)

Floronia bucculenta Heimer & Nentwig, 1991: 160, f. 435 (mf)

Floronia bucculenta Roberts, 1995: 363, f. (mf)

Floronia bucculenta Tao, Li & Zhu, 1995: 246, f. 61-68 (mf)

Floronia bucculenta Saaristo, 1996a: 6, f. 13, 15, 17 (mf)

Floronia bucculenta Roberts, 1998: 374, f. (mf)

Floronia bucculenta Song, Zhu & Chen, 1999: 169, f. 94M-N, 95A-B (mf)

Floronia bucculenta Hu, 2001: 498, f. 330.1-5 (f)

Floronia bucculenta Song, Zhu & Chen, 2001: 127, f. 66A-F (mf)

Floronia bucculenta Saaristo in Marusik & Koponen, 2008a: 9, pl. 2 (m)

Floronia bucculenta Dupérré, 2013: 7, f. 55 (m)

Floronia bucculenta Zhou et al., 2023a: 311, f. 1A-D (mf)

Floronia exornata (L. Koch, 1878) | | Korea, Japan urn:lsid:nmbe.ch:spidersp:010613

Linyphia exornata L. Koch, 1878c: 746, pl. 15, f. 10 (Df) Original description

Oecobius sapporense Saito, 1934b: 271, pl. 12, f. 1, pl. 14, f. 33 (Df) [urn:lsid:nmbe.ch:spidersp:005971]

Linyphia exornata Saito, 1934b: 310, pl. 13, f. 17, pl. 15, f. 60 (Df)

Oecobius sapporense Saito, 1959: 34, f. 7a-d (f)

Floronia bucculenta Yaginuma, 1960: 43, f. 41 (f; misidentified sensu Saaristo, 1996a: 6)

Floronia bucculenta Oi, 1960a: 193, f. 225-230 (mf; misidentified per Saaristo, 1996a: 6)

Floronia bucculenta Paik, 1965a: 59, f. 1-2 (mf; misidentified per Saaristo, 1996a: 6)

Oecobius sapporense Kritscher, 1966a: 293, f. 16 (f)

Floronia bucculenta Yaginuma, 1971: 43, f. 41.1-2 (f; misidentified sensu Saaristo, 1996a: 6)

Floronia bucculenta Paik, 1978e: 247, f. 104.1-9 (mf; misidentified sensu Saaristo, 1996a: 6)

Floronia bucculenta Yaginuma, 1986a: 71, f. 38.1 (mf; misidentified sensu Saaristo, 1996a: 6)

Floronia bucculenta Chikuni, 1989b: 52, f. 27 (mf; misidentified sensu Saaristo, 1996a: 6)

Floronia exornata Saaristo, 1996a: 6, f. 14, 16, 18 (mf, removed from S of Floronia bucculenta, rejecting Oi, 1960a: 193)

Floronia exornata Namkung, 2002: 159, f. 17.12a-b (mf)

Floronia exornata Namkung, 2003: 161, f. 17.12a-b (mf)

Floronia exornata Lee, Kang & Kim, 2009: 126, f. 24a-c (mf)

Neriene sapporense Santos, Gonzaga & Hormiga, 2009: 102 (T from Oecobius)

Floronia exornata Ono, Matsuda & Saito, 2009: 316, f. 861-866 (mf)

Floronia exornata Ono, 2009a: 589 (S of Neriene sapporense)

Floronia huishuiensis Zhou & Xu, 2023 | | China urn:lsid:nmbe.ch:spidersp:059520

Floronia huishuiensis Zhou & Xu in Zhou et al., 2023a: 311, f. 2A-D, 3A-B, 4A-E, 5A-F (Dmf) Original description

Floronia hunanensis Li & Song, 1993 | | China urn:lsid:nmbe.ch:spidersp:010615

Floronia hunanensis Li & Song, 1993: 251, f. 3A-I (Dmf) Original description

Floronia hunanensis Song & Li, 1997: 405, f. 8A-G (mf)

Floronia hunanensis Song, Zhu & Chen, 1999: 169, f. 94O-P, 95C (mf)

Floronia hunanensis Yin et al., 2012: 486, f. 223a-i (mf)

Floronia jiuhuensis Li & Zhu, 1987 | | China urn:lsid:nmbe.ch:spidersp:010616

Floronia jiuhuensis Li & Zhu in Li, Sha & Zhu, 1987: 43, f. 1A-C (Dm) Original description

Floronia jiuhuensis Song, Zhu & Chen, 1999: 169, f. 95D-E (m)

Floronia zhejiangensis Zhu, Chen & Sha, 1987 | | China urn:lsid:nmbe.ch:spidersp:010617

Floronia zhejiangensis Zhu, Chen & Sha, 1987: 139, f. 1-8 (Dmf) Original description

Floronia zhejiangensis Chen & Zhang, 1991: 169, f. 167.1-8 (mf)

Floronia zhejiangensis Song, Zhu & Chen, 1999: 169, f. 94Q-R, 95F-G (mf)

Floronia zhejiangensis Zhou et al., 2023a: 318, f. 6A-D (mf)