Hahniidae Bertkau, 1878

Family |   ACCEPTED
N.B.: an updated diagnosis is provided by Burgo et al., 2025: 12; the genus Cybaeolus Simon, 1884 was transferred from the Agelenidae (Cybaeinae) to the Hahniidae (Cybaeolinae Lehtinen, 1967) by Lehtinen, 1967: 226 and the Cybaeolinae are considered a junior synonym of Hahniidae by Burgo et al., 2025: 13; the genera of Cryphoecinae Lehtinen, 1967 (authorship correctly Simon, 1898a: 250, sub Cryphoeceae; also Cryphoecini Kishida, 1955: 6 is older) were transferred from the Agelenidae to the Hahniidae by Lehtinen, 1967, considered a subfamily of Dictynidae by Wunderlich, 2004j: 1385 and lately transferred to Cybaeidae by Wheeler et al., 2017: 606; transfers of Cicurina Menge, 1871 and related genera (Cicurininae sensu Lehtinen, 1967) from the Agelenidae to the Dictynidae (Lehtinen, 1967) have been accepted by Cokendolpher, 2004a: 28; Cicurina, as well as Chorizomma Simon, 1872 and Mastigusa Menge, 1854 (which are not explicitely listed by Wheeler et al., 2017) are transferred from the Dictynidae to the Hahniidae by Wheeler et al., 2017: 607; Mastigusa is transferred to the Cybaeidae by Castellucci et al., 2023: 6; the transfers of Chorizomma, Cicurina and Mastigusa to the Cicurinidae by Murphy & Roberts, 2015: viii were not sufficiently justified and are not accepted here; Ono & Ogata, 2018: 619 mention the authorship of these subfamilial taxa: Cicurininae Kishida, 1955: 4, Tetrilini Kishida, 1955: 5 and Chorizommatini Kishida, 1955: 6; but the oldest found mention of Cicurininae is by F. O. Pickard-Cambridge, 1893: 158 (sub Cicurinae, see also Bonnet, 1956: 1091); Cicurinidae new circumscribed (Bromella, Cicurina and Chorizomma) and elevated to family by Gorneau et al., 2023: 11.

In synonymy:

Clitistes Simon, 1902 = Cybaeolus Simon, 1884 (Burgo et al., 2025: 13)

Mevianes Simon, 1904 = Cybaeolus Simon, 1884 (Lehtinen, 1967: 248)

Gen. Cybaeolus Simon, 1884 urn:lsid:nmbe.ch:spidergen:01938
type Cybaeolus pusillus Simon, 1884; gender masculine
N.B.: transferred from the Agelenidae (Cybaeinae) to the Hahniidae (Cybaeolinae) and considered a senior synonym of Mevianes Simon, 1904e: 110 [urn:lsid:nmbe.ch:spidergen:05146] (type M. delfini Simon, 1904, transferred from the Agelenidae) by Lehtinen, 1967: 226; considered a senior synonym of Clitistes Simon, 1902h: 20 [urn:lsid:nmbe.ch:spidergen:00901] (type C. velutinus Simon, 1902, transferred from the Linyphiidae to the Dictynidae by Dupérré & Harms, 2018: 8) by Burgo et al., 2025: 13 (transfer from the Dictynidae).

In synonymy:

Cybaeolus fuegianus (Simon, 1895, T from Dictyna) = Cybaeolus pusillus Simon, 1884 (Lehtinen, 1967: 226)

Cybaeolus velutinus (Simon, 1902, T from Clitistes) = Cybaeolus pusillus Simon, 1884 (Burgo et al., 2025: 15)

Cybaeolus wilsoni (Simon, 1904, T from Mevianes) = Cybaeolus pusillus Simon, 1884 (Lehtinen, 1967: 226)

Cybaeolus delfini (Simon, 1904) | | Chile, Argentina urn:lsid:nmbe.ch:spidersp:021882
Mevianes delfini Simon, 1904e: 111 (Df) Original description
Cybaeolus delfini Lehtinen, 1967: 226, f. 353 (f, T from Mevianes)
Mevianes delfini Roth, 1967c: 307, pl. 50, f. 4-6 (f)
Cybaeolus delfini Burgo et al., 2025: 34, f. 1G, 19G-I, 20E-F, 25A-G, 26A-H, 27A.1-F (Dmf)
Cybaeolus pusillus Simon, 1884 | | Chile, Argentina urn:lsid:nmbe.ch:spidersp:021883 TYPE SPECIES
Cybaeolus pusillus Simon, 1884l: 125 (Df) Original description
Dictyna fuegiana Simon, 1895d: 168 (Df) [urn:lsid:nmbe.ch:spidersp:060541]
Clitistes velutinus Simon, 1902h: 20 (Df) [urn:lsid:nmbe.ch:spidersp:050095]
Dictyna fuegiana Simon, 1904e: 84 (Dm)
Mevianes wilsoni Simon, 1904e: 111 (Df) [urn:lsid:nmbe.ch:spidersp:060542]
Cybaeolus pusillus Lehtinen, 1967: 226, f. 348, 350 (mf, S of Dictyna fuegiana and Mevianes wilsoni)
Mevianes wilsoni Roth, 1967c: 310, pl. 50, f. 9-10 (Dm, f)
Clitistes velutinus Miller, 2007a: 269 (nomen dubium)
Clitistes velutinus Dupérré & Harms, 2018: 8, f. 5A-C (f, removed from nomen dubium)
Cybaeolus pusillus Burgo et al., 2025: 15, f. 1F, 2A-I, K-L, 3A-D, 4A-E, 17A-I, 19A-C, 20A-B, 21A-J, 22A.1-K.3 (mf, S of Clitistes velutinus)
Cybaeolus pusillus Montana et al., 2025: 80 (S of Clitistes velutinus confirmed)
Cybaeolus rastellus (Roth, 1967) | | Chile, Argentina urn:lsid:nmbe.ch:spidersp:021884
Cybaeolus rastellus Lehtinen, 1967: 226 (T from Mevianes)
Mevianes rastellus Roth, 1967c: 308, pl. 50, f. 7-8 (Dmf) Original description
Cybaeolus rastellus Burgo et al., 2025: 29, f. 2J, M, 5A-D, 6A-E, 15A-L, 18A-I, 19D-F, 20C-D, 23A-D, 24A.1-I (mf)

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