Cybaeidae Banks, 1892

Family |   ACCEPTED
N.B.: traditionally considered a subfamily of Agelenidae, placed as a subfamily of Dictynidae by Lehtinen, 1967 and elevated to family by Forster, 1970b: 13; after Wheeler et al., 2017: 606 Cybaeidae includes Cryphoecinae Lehtinen, 1967 syn. nov. [authorship correctly Simon, 1898a: 250, sub Cryphoeceae; also Cryphoecini Kishida, 1955: 6 is older] (genera Cryphoeca Thorell, 1870, Calymmaria Chamberlin & Ivie, 1937, Ethobuella Chamberlin & Ivie, 1937 and Willisus Roth, 1981 as well as Cryphoecina Deltshev, 1997, Dirksia Chamberlin & Ivie, 1942, Neocryphoeca Roth, 1970 and Tuberta Simon, 1884, which are not explicitely listed by Wheeler et al., 2017; all transferred from Hahniidae) and the genera Blabomma Chamberlin & Ivie, 1937 and Yorima Chamberlin & Ivie, 1942 (transferred from Dictynidae); furthermore Argyroneta Latreille, 1804 (monotypic in Argyronetinae Thorell, 1869) is transferred to the Dictynidae by Wheeler et al., 2017: 607; Mastigusa Menge, 1854 is transferred from the Hahniidae to the Cybaeidae by Castellucci et al., 2023: 6; the transfers of Blabomma, Cryphoeca and Tuberta to Cicurinidae Murphy & Roberts, 2015 [oldest available name is Cicurininae Kishida, 1955: 4] (Murphy & Roberts, 2015: viii) not sufficiently justified and is not accepted here; Cybaeidae were considered a synonym of Argyronetidae Thorell, 1869 by Grothendieck & Kraus, 1994: 272; Cicurinidae is new circumscribed (Bromella, Cicurina and Chorizomma) and elevated to family by Gorneau et al., 2023: 11; included genera of Cybaeidae are listed by Gorneau et al., 2023: 11; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 515, f. 15-16.
Gen. Cryphoeca Thorell, 1870 urn:lsid:nmbe.ch:spidergen:01936
N.B.: transferred from the Agelenidae to the Hahniidae by Lehtinen, 1967: 225 and to the Cybaeidae by Wheeler et al., 2017: 606; considered a possible member of Dictynidae by Wunderlich, 1995z: 719; transfer to the Cicurinidae (Murphy & Roberts, 2015: viii) not sufficiently justified and is not accepted here.

Transferred to other genera:

Cryphoeca peckhami Simon, 1898 -- see Cicurina

Cryphoeca pyrenaea Simon, 1898 -- see Dirksia

Cryphoeca recisa O. Pickard-Cambridge, 1908 -- see Mastigusa

Cryphoeca tibetana Hu & Li, 1987 -- see Huangyuania

In synonymy:

Cryphoeca saxatilis (Ausserer, 1867, T from Apostenus) = Cryphoeca silvicola C. L. Koch, 1834 (Thaler, 1988: 118)

Nomina dubia:

Cryphoeca cavicola Caporiacco, 1938d: 40 (j, Slovenia) -- Thaler, 1978b: 335

Cryphoeca latitans Menge, 1871: 275, pl. 51, f. 161A-N (j, Poland) -- Thaler, 1978b: 335

Cryphoeca angularis Saito, 1934 | | Japan urn:lsid:nmbe.ch:spidersp:021869
Cryphoeca angularis Saito, 1934b: 348, pl. 13, f. 29, pl. 15, f. 77 (Df) Original description
Cryphoeca brignolii Thaler, 1980 | | Switzerland, Italy urn:lsid:nmbe.ch:spidersp:021870
Cryphoeca brignolii Thaler, 1980b: 402, f. 1-4, 7-8 (Dmf; f. 5-6 see C. thaleri, per Wunderlich, 1995z: 719) Original description
Cryphoeca brignolii Wunderlich, 1995z: 720, f. 3-4 (m)
Cryphoeca carpathica Herman, 1879 | | Poland, Slovakia, Romania, Ukraine urn:lsid:nmbe.ch:spidersp:021871
Cryphoeca carpathica Herman, 1879: 129, 353 (Dmf) Original description
Cryphoeca silvicola carpathica Chyzer & Kulczyński, 1897: 157
Cryphoeca carpathica Thaler, 1980b: 403, f. 9-12 (mf, elevated from subspecies)
Cryphoeca carpathica Heimer & Nentwig, 1991: 360, f. 933 (mf)
Cryphoeca exlineae Roth, 1988 | | USA urn:lsid:nmbe.ch:spidersp:021872
Cryphoeca peckhami Exline, 1938: 18, pl. 1, f. 5, pl. 3, f. 21-22 (f, Dm, misidentified)
Cryphoeca peckhami Roth & Brame, 1972: 29, f. 39-40 (mf, misidentified)
Cryphoeca peckhami Thaler, 1980b: 401, f. 17-18 (mf, misidentified)
Cryphoeca exlineae Roth, 1988: 31, f. 3-6 (Dmf) Original description
Cryphoeca exlineae Bennett, 1992: 7, f. 35 (f)
Cryphoeca exlineae Roth, 1994: 53, f. 39-40 (mf)
Cryphoeca lichenum L. Koch, 1876 | | Germany, Austria urn:lsid:nmbe.ch:spidersp:021873
Cryphoeca lichenum L. Koch, 1876b: 297 (Dmf) Original description
Cryphoeca lichenum Thaler, 1978b: 342, f. 14 (f)
Cryphoeca lichenum Heimer & Nentwig, 1991: 358, f. 930 (mf)
Cryphoeca lichenum nigerrima Thaler, 1978 | | Germany, Austria urn:lsid:nmbe.ch:spidersp:021874
Cryphoeca lichenum nigerrima Thaler, 1978b: 343, f. 3, 6, 9, 12-13 (Dmf) Original description
Cryphoeca montana Emerton, 1909 | | USA, Canada urn:lsid:nmbe.ch:spidersp:021875
Cryphoeca montana Emerton, 1909: 222, pl. 8, f. 4 (Dmf) Original description
Cryphoeca montana Levi & Field, 1954: 454, f. 57, 59-60 (mf)
Cryphoeca montana Thaler, 1980b: 401, f. 13-16 (mf)
Cryphoeca montana Paquin & Dupérré, 2003: 66, f. 545-547 (mf)
Cryphoeca nivalis Schenkel, 1919 | | Switzerland, Austria, Italy urn:lsid:nmbe.ch:spidersp:021876
Cryphoeca silvicola nivalis Schenkel, 1919: 84 (Dmf, omitted by Roewer) Original description
Cryphoeca nivalis Schenkel, 1923: 118 (elevated to species)
Cryphoeca nivalis Thaler, 1978b: 339, f. 2, 5, 10-11, 20-21 (mf)
Cryphoeca nivalis Heimer & Nentwig, 1991: 360, f. 932 (mf)
Cryphoeca nivalis Hansen & Vanin, 2004: 89, f. 1 (f)
Cryphoeca pirini (Drensky, 1921) | | Bulgaria urn:lsid:nmbe.ch:spidersp:021877
Hahnia pirini Drensky, 1921: 42, 77, pl. 1, f. 9-11 (Df) Original description
Cybaeus pirini Drensky, 1942: 56, f. 19 (f)
Cryphoeca pirini Brignoli, 1978g: 508, f. 72 (T from Cybaeus; mentioned females belong to Cryphoeca brignolii per Thaler, 1980b: 403, but can probably be attached to Cryphoeca thaleri, cf. Wunderlich, 1995z: 720)
Cryphoeca shingoi Ono, 2007 | | Japan urn:lsid:nmbe.ch:spidersp:041316
Cryphoeca shingoi Ono, 2007: 157, f. 16-28 (Dmf) Original description
Cryphoeca shingoi Ono, 2009a: 207, f. 1-11 (mf)
Cryphoeca shinkaii Ono, 2007 | | Japan urn:lsid:nmbe.ch:spidersp:041315
Cryphoeca shinkaii Ono, 2007: 154, f. 1-15 (Dmf) Original description
Cryphoeca shinkaii Ono, 2009a: 207, f. 12-22 (mf)
Cryphoeca silvicola (C. L. Koch, 1834) | | Europe, Turkey, Russia (Europe to Far East), Japan urn:lsid:nmbe.ch:spidersp:021878
Tegenaria silvicola C. L. Koch, 1834: 125, pl. 16 (Df) Original description
Hahnia silvicola C. L. Koch, 1845: 158, f. 1076-1077 (Dmf)
Amaurobius sylvicolus Menge, 1850: 63 (lapsus)
Agelena hyndmannii Blackwall, 1861a: 154, pl. 10, f. 98 (Df)
Tegenaria silvicola Blackwall, 1861a: 168, pl. 12, f. 108 (mf)
Hahnia sylvicola Ohlert, 1867: 87 (lapsus)
Apostenus saxatilis Ausserer, 1867: 163 (Df) [urn:lsid:nmbe.ch:spidersp:057728]
Hahnia silvicola Menge, 1869: 254, pl. 48, f. 151 (mf)
Cryphoeca silvicola Simon, 1875a: 52, pl. 5, f. 17 (mf)
Cryphoeca silvicola Dahl, 1883: 61
Cryphoeca silvicola Simon, 1898a: 268, f. 262-263 (m)
Cryphoeca silvicola Bösenberg, 1902: 232, pl. 21, f. 330 (mf)
Cryphoeca silvicola Dahl, 1931: 5, f. 1-5 (mf)
Cryphoeca silvicola Simon, 1937: 1021, 1044, f. 1588-1589 (mf)
Cryphoeca silvicola Drensky, 1942: 56, f. 18a-b (mf)
Cryphoeca silvicola Locket & Millidge, 1953: 22, f. 16C-E (mf)
Cryphoeca silvicola Lehtinen, 1967: 454, f. 351-352 (mf)
Cryphoeca silvicola Locket & Millidge, 1967: 182, f. 5C (m)
Cryphoeca silvicola Yaginuma & Nishikawa, 1969: 74, f. 6-9 (m)
Cryphoeca silvicola Loksa, 1969: 114, f. 77A-B (mf)
Cryphoeca silvicola Miller, 1971: 178, pl. XXIX, f. 20-22 (mf)
Cryphoeca silvicola Palmgren, 1977a: 10, f. 1.16-19 (mf)
Cryphoeca silvicola Brignoli, 1977f: 27, f. 12 (f)
Cryphoeca silvicola Thaler, 1978b: 337, f. 1, 4, 7-8, 15-18 (mf)
Cryphoeca silvicola Roberts, 1985: 166, f. 73d (mf)
Cryphoeca sylvicola Barrientos, 1985a: 85, f. 5-8 (mf)
Cryphoeca silvicola Wunderlich, 1986: 179, f. 97 (m)
Cryphoeca silvicola Yaginuma, 1986a: 141, f. 76.1 (m)
Cryphoeca silvicola Thaler, 1988: 118, f. 5 (f, S of Apostenus saxatilis)
Cryphoeca silvicola Heimer & Nentwig, 1991: 358, f. 931 (mf)
Cryphoeca silvicola Roberts, 1995: 252, f. (mf)
Cryphoeca silvicola Roberts, 1998: 269, f. (mf)
Cryphoeca silvicola Wunderlich, 2004j: 1389, f. 5 (m)
Cryphoeca silvicola Almquist, 2006: 308, f. 270a-i (mf)
Cryphoeca silvicola Marusik & Kovblyuk, 2011: 161, f. 18/2, 8, 10-11 (m)
Cryphoeca silvicola Ledoux, 2014: 29, f. 1D (m)
Cryphoeca silvicola Lecigne, 2016d: 13, f. 3A-B (m)
Cryphoeca silvicola Geci et al., 2023: 539, f. 11 (f)
Cryphoeca silvicola Castellucci, Luchetti & Scharff, 2024: 24, f. 18a-b (m)
Cryphoeca thaleri Wunderlich, 1995 | | Turkey urn:lsid:nmbe.ch:spidersp:021879
Cryphoeca brignolii Thaler, 1980b: 402, f. 5-6 (m; misidentified per Wunderlich, 1995z: 719)
Cryphoeca thaleri Wunderlich, 1995z: 719, f. 1-2 (Dm) Original description

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