Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Xerophaeus Purcell, 1907 urn:lsid:nmbe.ch:spidergen:02498
type Xerophaeus capensis Purcell, 1907; gender masculine

Transferred to other genera:

Xerophaeus coruscus (L. Koch, 1875) -- see Sorkinius

Xerophaeus espoir Platnick, 1981 -- see Sorkinius

Xerophaeus kibonotensis Tullgren, 1910 -- see Sorkinius

Xerophaeus oceanicus Schmidt & JocquƩ, 1983 -- see Sorkinius

Xerophaeus zuluensis Lawrence, 1938 -- see Trichothyse

In synonymy:

Xerophaeus delphinurus Purcell, 1907 = Xerophaeus capensis Purcell, 1907 (Tucker, 1923: 397)

Xerophaeus gordonicus Hewitt, 1915 = Xerophaeus hottentottus Purcell, 1908 (Tucker, 1923: 405)

Xerophaeus interrogator Purcell, 1907 = Xerophaeus capensis Purcell, 1907 (Tucker, 1923: 397)

Xerophaeus poweri Hewitt, 1915 = Xerophaeus aridus Purcell, 1907 (Tucker, 1923: 394)

Xerophaeus ahenus Purcell, 1908 | | South Africa urn:lsid:nmbe.ch:spidersp:028055
Xerophaeus ahenus Purcell, 1908: 237, pl. 11, f. 19 (Df) Original description
Xerophaeus ahenus Dippenaar-Schoeman et al., 2021D: 32, 1 f. (f)
Xerophaeus anthropoides Hewitt, 1916 | | South Africa urn:lsid:nmbe.ch:spidersp:028056
Xerophaeus anthropoides Hewitt, 1916b: 211, f. 10 (Df) Original description
Xerophaeus anthropoides Dippenaar-Schoeman et al., 2021D: 33, 4 f. (f)
Xerophaeus appendiculatus Purcell, 1907 | | South Africa urn:lsid:nmbe.ch:spidersp:028057
Xerophaeus appendiculatus Purcell, 1907: 326, pl. 15, f. 41-42 (Dmf) Original description
Xerophaeus appendiculatus Tucker, 1923: 393, pl. 11, f. 86 (mf)
Xerophaeus appendiculatus Rodrigues & Rheims, 2020b: 680, f. 17C, 28C, 29C (f)
Xerophaeus appendiculatus Dippenaar-Schoeman et al., 2021D: 34, 8 f. (mf)
Xerophaeus aridus Purcell, 1907 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:028058
Xerophaeus aridus Purcell, 1907: 319, pl. 15, f. 26 (Df) Original description
Prosthesima aridus Tullgren, 1910: 109, pl. 1, f. 23 (f)
Xerophaeus poweri Hewitt, 1915b: 94, f. 4 (Dmf) [urn:lsid:nmbe.ch:spidersp:057399]
Xerophaeus aridus Tucker, 1923: 394, pl. 11, f. 87 (m, S of Xerophaeus poweri)
Xerophaeus aridus Dippenaar-Schoeman et al., 2021D: 35, 6 f. (mf)
Xerophaeus aurariarum Purcell, 1907 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:028059
Xerophaeus aurariarum Purcell, 1907: 324, pl. 15, f. 35-36 (Dmf) Original description
Xerophaeus aurariarum Dippenaar-Schoeman et al., 2021D: 36, 7 f. (mf)
Xerophaeus bicavus Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028060
Xerophaeus bicavus Tucker, 1923: 397, pl. 10, f. 88 (Dmf) Original description
Xerophaeus bicavus Dippenaar-Schoeman et al., 2021D: 37, 7 f. (mf)
Xerophaeus biplagiatus Tullgren, 1910 | | Tanzania, South Africa urn:lsid:nmbe.ch:spidersp:028061
Xerophaeus biplagiatus Tullgren, 1910: 108, pl. 1, f. 21 (Df) Original description
Xerophaeus biplagiatus Dippenaar-Schoeman et al., 2021D: 38, 1 f. (f)
Xerophaeus capensis Purcell, 1907 | | South Africa urn:lsid:nmbe.ch:spidersp:028062 TYPE SPECIES
Xerophaeus capensis Purcell, 1907: 314, pl. 13, f. 20 (Dmf) Original description
Xerophaeus delphinurus Purcell, 1907: 316, pl. 13, f. 21 (Dm) [urn:lsid:nmbe.ch:spidersp:057397]
Xerophaeus interrogator Purcell, 1907: 316, pl. 14, f. 22 (Dm) [urn:lsid:nmbe.ch:spidersp:057398]
Xerophaeus capensis Tucker, 1923: 397, pl. 11, f. 89 (mf, S of Xerophaeus delphinurus and X. interrogator)
Xerophaeus capensis Murphy, 2007: 50, f. 386-387 (mf)
Xerophaeus capensis Dippenaar-Schoeman et al., 2021D: 39, 10 f. (mf)
Xerophaeus communis Purcell, 1907 | | South Africa urn:lsid:nmbe.ch:spidersp:028063
Xerophaeus communis Purcell, 1907: 322, pl. 14, f. 33-34 (Dmf) Original description
Xerophaeus communis Dippenaar-Schoeman et al., 2021D: 40, 7 f. (mf)
Xerophaeus crusculus Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028066
Xerophaeus crusculus Tucker, 1923: 400, pl. 11, f. 90 (Dmf) Original description
Xerophaeus crusculus Dippenaar-Schoeman et al., 2021D: 42, 3 f. (mf)
Xerophaeus crustosus Purcell, 1907 | | South Africa urn:lsid:nmbe.ch:spidersp:028067
Xerophaeus crustosus Purcell, 1907: 325, pl. 15, f. 40 (Dm) Original description
Xerophaeus crustosus Dippenaar-Schoeman et al., 2021D: 43, 4 f. (m)
Xerophaeus druryi Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028068
Xerophaeus druryi Tucker, 1923: 402, pl. 11, f. 91 (Dm) Original description
Xerophaeus exiguus Purcell, 1907 | | South Africa urn:lsid:nmbe.ch:spidersp:028070
Xerophaeus exiguus Purcell, 1907: 324, pl. 14, f. 37 (Dm) Original description
Xerophaeus exiguus Tucker, 1923: 403 (Df)
Xerophaeus exiguus Dippenaar-Schoeman et al., 2021D: 44, 2 f. (m)
Xerophaeus flammeus Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028071
Xerophaeus flammeus Tucker, 1923: 404, pl. 10, f. 92 (Df) Original description
Xerophaeus flammeus Dippenaar-Schoeman et al., 2021D: 45, 1 f. (f)
Xerophaeus flavescens Purcell, 1907 | | South Africa urn:lsid:nmbe.ch:spidersp:028072
Xerophaeus flavescens Purcell, 1907: 317, pl. 14, f. 23 (Dm) Original description
Xerophaeus flavescens Dippenaar-Schoeman et al., 2021D: 46, 1 f. (m)
Xerophaeus hottentottus Purcell, 1908 | | South Africa urn:lsid:nmbe.ch:spidersp:028073
Xerophaeus hottentottus Purcell, 1908: 236, pl. 11, f. 17 (Df) Original description
Xerophaeus gordonicus Hewitt, 1915b: 97, f. 5 (Df) [urn:lsid:nmbe.ch:spidersp:057400]
Xerophaeus hottentottus Tucker, 1923: 405 (S of Xerophaeus gordonicus)
Xerophaeus hottentottus Dippenaar-Schoeman et al., 2021D: 47, 7 f. (f)
Xerophaeus kiwuensis Strand, 1913 | | Central Africa urn:lsid:nmbe.ch:spidersp:028074
Xerophaeus kiwuensis Strand, 1913a: 334 (Dm) Original description
Xerophaeus lightfooti Purcell, 1907 | | South Africa urn:lsid:nmbe.ch:spidersp:028075
Xerophaeus lightfooti Purcell, 1907: 321, pl. 14, f. 31-32 (Dmf) Original description
Xerophaeus lightfooti Dippenaar-Schoeman et al., 2021D: 48, 2 f. (mf)
Xerophaeus longispina Purcell, 1908 | | South Africa urn:lsid:nmbe.ch:spidersp:028076
Xerophaeus longispina Purcell, 1908: 235, pl. 11, f. 15-16 (Dmf) [N.B.: longispina is a noun and no adjective] Original description
Xerophaeus longispinus Dippenaar-Schoeman et al., 2021D: 49, 2 f. (mf)
Xerophaeus lunulifer Purcell, 1907 | | South Africa urn:lsid:nmbe.ch:spidersp:028077
Xerophaeus lunulifer Purcell, 1907: 319, pl. 14, f. 27-28 (Dmf) Original description
Xerophaeus lunulifer Dippenaar-Schoeman et al., 2021D: 50, 2 f. (mf)
Xerophaeus maritimus Lawrence, 1938 | | South Africa urn:lsid:nmbe.ch:spidersp:028078
Xerophaeus maritimus Lawrence, 1938a: 480, f. 15 (Df) Original description
Xerophaeus maritimus Dippenaar-Schoeman et al., 2021D: 51, 4 f. (f)
Xerophaeus matroosbergensis Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028079
Xerophaeus matroosbergensis Tucker, 1923: 408, pl. 11, f. 93 (Df) Original description
Xerophaeus matroosbergensis Dippenaar-Schoeman et al., 2021D: 52, 1 f. (f)
Xerophaeus occiduus Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028080
Xerophaeus occiduus Tucker, 1923: 409, pl. 11, f. 94 (Dm) Original description
Xerophaeus pallidus Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028082
Xerophaeus pallidus Tucker, 1923: 409, pl. 10, f. 95 (Dm) Original description
Xerophaeus pallidus Dippenaar-Schoeman et al., 2021D: 53, 1 f. (m)
Xerophaeus patricki Purcell, 1907 | | Mozambique, South Africa urn:lsid:nmbe.ch:spidersp:028083
Xerophaeus patricki Purcell, 1907: 327, pl. 15, f. 43 (Dm) Original description
Xerophaeus patricki Dippenaar-Schoeman et al., 2021D: 54, 3 f. (m)
Xerophaeus perversus Purcell, 1923 | | Namibia urn:lsid:nmbe.ch:spidersp:028084
Xerophaeus aridus Purcell, 1908: 236, pl. 11, f. 18 (m; misidentified per Purcell, in Tucker, 1923: 410)
Xerophaeus perversus Purcell in Tucker, 1923: 410 (Dm) Original description
Xerophaeus phaseolus Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028085
Xerophaeus phaseolus Tucker, 1923: 411, pl. 10, f. 96 (Df) Original description
Xerophaeus phaseolus Dippenaar-Schoeman et al., 2021D: 55, 3 f. (f)
Xerophaeus robustus Lawrence, 1936 | | South Africa urn:lsid:nmbe.ch:spidersp:028086
Xerophaeus robustus Lawrence, 1936: 147, f. 1 (Df) Original description
Xerophaeus rostratus Purcell, 1907 | | South Africa urn:lsid:nmbe.ch:spidersp:028087
Xerophaeus rostratus Purcell, 1907: 321, pl. 14, f. 38-39 (Dmf) Original description
Xerophaeus rostratus Dippenaar-Schoeman et al., 2021D: 56, 3 f. (mf)
Xerophaeus ruandanus Strand, 1913 | | Rwanda urn:lsid:nmbe.ch:spidersp:028088
Xerophaeus ruandanus Strand, 1913a: 333 (Dm) Original description
Xerophaeus rubeus Tucker, 1923 | | South Africa, Lesotho urn:lsid:nmbe.ch:spidersp:028089
Xerophaeus rubeus Tucker, 1923: 412, pl. 10, f. 97 (Df) Original description
Xerophaeus rubeus Dippenaar-Schoeman et al., 2021D: 57, 1 f. (f)
Xerophaeus silvaticus Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028090
Xerophaeus silvaticus Tucker, 1923: 413, pl. 11, f. 98 (Df) Original description
Xerophaeus silvaticus Dippenaar-Schoeman et al., 2021D: 58, 1 f. (f)
Xerophaeus spiralifer Purcell, 1907 | | South Africa urn:lsid:nmbe.ch:spidersp:028091
Xerophaeus spiralifer Purcell, 1907: 318, pl. 14, f. 24-25 (Dmf) Original description
Xerophaeus spiralifer Dippenaar-Schoeman et al., 2021D: 59, 5 f. (mf)
Xerophaeus spoliator Purcell, 1907 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:028092
Xerophaeus spoliator Purcell, 1907: 321, pl. 14, f. 29-30 (Dmf) Original description
Xerophaeus spoliator Dippenaar-Schoeman et al., 2021D: 60, 5 f. (mf)
Xerophaeus tenebrosus Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028093
Xerophaeus tenebrosus Tucker, 1923: 415, pl. 10, f. 99 (Df) Original description
Xerophaeus tenebrosus Dippenaar-Schoeman et al., 2021D: 61, 3 f. (f)
Xerophaeus thomasi (Caporiacco, 1949) | | Kenya urn:lsid:nmbe.ch:spidersp:028413
Zelotes thomasi Caporiacco, 1949a: 430, f. 67a-b (Dm) Original description
Xerophaeus thomasi FitzPatrick, 2007: 98 (T from Zelotes)
Xerophaeus vickermani Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028094
Xerophaeus vickermani Tucker, 1923: 416, pl. 10, f. 100 (Dmf) Original description
Xerophaeus vickermani Dippenaar-Schoeman et al., 2021D: 62, 5 f. (mf)

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