Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.

In synonymy:

Upognampa Tucker, 1923 = Trephopoda Tucker, 1923 (Murphy, 2007: 34)

Gen. Trephopoda Tucker, 1923 urn:lsid:nmbe.ch:spidergen:02491
N.B.: considered a senior synonym of Upognampa Tucker, 1923: 271  [urn:lsid:nmbe.ch:spidergen:02494] (type U. aplanita Tucker, 1923) by Murphy, 2007: x, 34.

Transferred to other genera:

Trephopoda lineatipes (Purcell, 1908) -- see Smionia

Trephopoda aplanita (Tucker, 1923) | | South Africa urn:lsid:nmbe.ch:spidersp:028044
Upognampa aplanita Tucker, 1923: 273, pl. 8, f. 9 (Df) Original description
Trephopoda aplanita Murphy, 2007: 34, f. 134-135 (f)
Trephopoda aplanita Dippenaar-Schoeman et al., 2021D: 21, 2 f. (f)
Trephopoda biamenta (Tucker, 1923) | | South Africa urn:lsid:nmbe.ch:spidersp:028045
Upognampa biamenta Tucker, 1923: 274, pl. 8, f. 10 (Df) Original description
Trephopoda biamenta Dippenaar-Schoeman et al., 2021D: 22, 3 f. (f)
Trephopoda ctenipalpis (Lawrence, 1927) | | Namibia urn:lsid:nmbe.ch:spidersp:028046
Upognampa ctenipalpis Lawrence, 1927: 11, pl. 1, f. 5 (Df) Original description
Trephopoda hanoveria Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:028039
Trephopoda hanoveria Tucker, 1923: 271, pl. 8, f. 8 (Df) Original description
Trephopoda hanoveria Murphy, 2007: 32, f. 130-131 (f)
Trephopoda hanoveria Dippenaar-Schoeman et al., 2021D: 23, 3 f. (f)
Trephopoda kannemeyeri (Tucker, 1923) | | South Africa urn:lsid:nmbe.ch:spidersp:028047
Upognampa kannemeyeri Tucker, 1923: 274, pl. 8, f. 11 (Df) Original description
Trephopoda kannemeyeri Murphy, 2007: 34, f. 136-137 (f, f. m)
Trephopoda kannemeyeri Dippenaar-Schoeman et al., 2021D: 24, 12 f. (mf)
Trephopoda parvipalpa (Tucker, 1923) | | Botswana, South Africa urn:lsid:nmbe.ch:spidersp:028049
Upognampa parvipalpa Tucker, 1923: 277, pl. 10, f. 13 (Dm) Original description
Trephopoda parvipalpa Dippenaar-Schoeman et al., 2021D: 25, 3 f. (m)

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