Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Poecilochroa Westring, 1874 urn:lsid:nmbe.ch:spidergen:02464
type Pythonissa variana C. L. Koch, 1839; gender feminine
N.B.: considered a senior synonym of Sergiolus Simon, 1891 by Ubick & Roth, 1973: 5, but not by Platnick & Shadab, 1981e: 4; not a senior synonym of Kishidaia Yaginuma, 1960 (Kamura, 2001b: 194, contra Paik, 1992a: 118).

Transferred to other genera:

Poecilochroa abjecta Chamberlin, 1936 -- see Sergiolus

Poecilochroa albimaculata (Saito, 1934) -- see Kishidaia

Poecilochroa amphiloga (Chamberlin, 1936) -- see Gertschosa

Poecilochroa angusta (Banks, 1904) -- see Sergiolus

Poecilochroa antineae Fage, 1929 -- see Trichothyse

Poecilochroa atomistica (Chamberlin, 1924) -- see Sergiolus

Poecilochroa bebia (Chamberlin, 1936) -- see Sergiolus

Poecilochroa behni Thorell, 1891 -- see Hitobia

Poecilochroa bellior (Chamberlin, 1936) -- see Sergiolus

Poecilochroa bicolor (Banks, 1900) -- see Sergiolus

Poecilochroa campestrata (O. Pickard-Cambridge, 1874) -- see Trichothyse

Poecilochroa capulata (Walckenaer, 1837) -- see Sergiolus

Poecilochroa cingulata (Roewer, 1933) -- see Gertschosa

Poecilochroa clara (Chamberlin, 1936) -- see Sergiolus

Poecilochroa clericus (Chamberlin, 1922) -- see Sergiolus

Poecilochroa columbiana Emerton, 1917 -- see Sergiolus

Poecilochroa comata (Ohlert, 1867) -- see Kishidaia

Poecilochroa concinna Simon, 1895 -- see Gertschosa

Poecilochroa conspicua (L. Koch, 1866) -- see Kishidaia

Poecilochroa conspicua concolor (Caporiacco, 1951) -- see Kishidaia

Poecilochroa convictrix Simon, 1909 -- see Sergiolus

Poecilochroa coreana Paik, 1992) -- see Kishidaia

Poecilochroa cyaneiventris (Simon, 1893) -- see Sergiolus

Poecilochroa decorata (Kaston, 1945) -- see Sergiolus

Poecilochroa devendrai Gajbe & Rane, 1985 -- see Hitobia

Poecilochroa dimidiata (Simon, 1878) -- see Trichothyse

Poecilochroa famula (Chamberlin, 1922) -- see Sergiolus

Poecilochroa flavomaculata (L. Koch, 1878) -- see Trichothyse

Poecilochroa fruitana (Chamberlin, 1928) -- see Sergiolus

Poecilochroa furcata Simon, 1914 -- see Trichothyse

Poecilochroa golan Levy, 1999 -- see Trichothyse

Poecilochroa hamipalpis (Kroneberg, 1875) -- see Trichothyse

Poecilochroa hosiziro Yaginuma, 1960 -- see Sergiolus

Poecilochroa hungarica Kolosváry, 1934 -- see Aphantaulax

Poecilochroa ilkerakkusi Coşar, Danışman & Marusik, 2024 -- see Trichothyse

Poecilochroa incompta (Pavesi, 1880) -- see Trichothyse

Poecilochroa inconspicua Bryant, 1948 -- see Gnaphosa

Poecilochroa jodhpurensis (Gajbe, 1993) -- see Trichothyse

Poecilochroa josephus Chamberlin & Gertsch, 1940 -- see Cesonia

Poecilochroa lesserti Denis, 1947 -- see Trichothyse

Poecilochroa lesserti (Schenkel, 1950) -- see Herpyllus

Poecilochroa loricata Kritscher, 1996 -- see Trichothyse

Poecilochroa lowelli (Chamberlin & Woodbury, 1929) -- see Sergiolus

Poecilochroa meretrix (Chamberlin, 1922) -- see Sergiolus

Poecilochroa minuta Banks, 1898 -- see Sergiolus

Poecilochroa monodi Fage, 1929 -- see Odontodrassus

Poecilochroa montana Emerton, 1890 -- see Sergiolus

Poecilochroa montanoides Schenkel, 1950 -- see Sergiolus

Poecilochroa ocellata (Walckenaer, 1837) -- see Sergiolus

Poecilochroa ochridana Drensky, 1929 -- see Callilepis

Poecilochroa otonensis Barrion & Litsinger, 1995 -- see Castianeira

Poecilochroa pacifica Banks, 1896 -- see Sergiolus

Poecilochroa perversa Simon, 1914 -- see Trichothyse

Poecilochroa poonaensis (Tikader, 1982) -- see Trichothyse

Poecilochroa pugnax (O. Pickard-Cambridge, 1874) -- see Trichothyse

Poecilochroa quadrimaculata (Yaginuma, 1960) -- see Kishidaia

Poecilochroa rufipes (Thorell, 1875) -- see Trichothyse

Poecilochroa sabulum (Wunderlich, 2011) -- see Trichothyse

Poecilochroa sedula (Simon, 1897) -- see Xizangiana

Poecilochroa senilis (O. Pickard-Cambridge, 1872) -- see Trichothyse

Poecilochroa senilis auspex (Simon, 1878) -- see Trichothyse

Poecilochroa sockniensis (Karsch, 1881) -- see Trichothyse

Poecilochroa stella (Chamberlin, 1922) -- see Sergiolus

Poecilochroa subincompta Strand, 1908 -- see Trichothyse

Poecilochroa taborensis Levy, 1999 -- see Trichothyse

Poecilochroa tennesseensis (Chamberlin, 1922) -- see Sergiolus

Poecilochroa tescorum Simon, 1914 -- see Sernokorba

Poecilochroa tikaderi Patel, 1989 -- see Hitobia

Poecilochroa tribolus (Chamberlin, 1922) -- see Sergiolus

Poecilochroa unifascigera (Bösenberg & Strand, 1906) -- see Hitobia

Poecilochroa unimaculata (Emerton, 1915) -- see Sergiolus

In synonymy:

Poecilochroa nigrofemorata (O. Pickard-Cambridge, 1874, T from Drassus) = Poecilochroa variana (C. L. Koch, 1839) (Simon, 1914: 221)

Poecilochroa picta Simon, 1878 = Poecilochroa albomaculata (Lucas, 1846) (Simon, 1914: 221)

Poecilochroa pulchra (Thorell, 1875, T from Prosthesima) = Poecilochroa albomaculata (Lucas, 1846) (Simon, 1914: 221)

Homonym replaced:

Poecilochroa bicolor (Costa, 1834) -- see Poecilochroa phyllobia Thorell, 1871 Thorell, 1871a

Nomen dubium:

Poecilochroa malagassa Strand, 1907d: 727, Strand, 1907o: 19 (m, Madagascar) -- Nentwig et al., 2020: 24

Poecilochroa albomaculata (Lucas, 1846) | | Portugal, Spain, France, Italy, Morocco, Algeria, Tunisia urn:lsid:nmbe.ch:spidersp:027724
Drassus albomaculatus Lucas, 1846: 224, pl. 13, f. 8 (Df) Original description
Macaria albomaculata Simon, 1864: 113
Prosthesima pulchra Thorell, 1875c: 111 (Df) [urn:lsid:nmbe.ch:spidersp:060370]
Poecilochroa picta Simon, 1878a: 160, pl. 14, f. 2 (Dmf) [urn:lsid:nmbe.ch:spidersp:060371]
Poecilochroa albomaculata Simon, 1914: 183, 221 (m, S of Poecilochroa picta and Prosthesima pulchra)
Poecilochroa albomaculata Grimm, 1985: 163, f. 184-189 (f)
Poecilochroa albomaculata Di Franco, 1987: 148, f. 9-10 (m)
Poecilochroa alcala Barrion & Litsinger, 1995 | | Philippines urn:lsid:nmbe.ch:spidersp:027725
Poecilochroa alcala Barrion & Litsinger, 1995: 190, f. 109a-h (Df) Original description
Poecilochroa anomala (Hewitt, 1915) | | South Africa, Lesotho urn:lsid:nmbe.ch:spidersp:027726
Xerophaeus anomalus Hewitt, 1915b: 98, f. 6 (Df) Original description
Poecilochroa anomala Tucker, 1923: 381, pl. 10, f. 81 (f)
Poecilochroa anomala Dippenaar-Schoeman et al., 2021C: 42, 6 f. (f)
Poecilochroa barmani Tikader, 1982 | | India urn:lsid:nmbe.ch:spidersp:027728
Poecilochroa barmani Tikader, 1982b: 451, f. 354-358 (f) Original description
Poecilochroa barmani Gajbe, 2007: 477, f. 157-161 (f)
Poecilochroa barmani Sankaran & Caleb, 2025: 136, f. 8-10 (f)
Poecilochroa bifasciata Banks, 1902 | | Galapagos urn:lsid:nmbe.ch:spidersp:027730
Poecilochroa bifasciata Banks, 1902c: 57, pl. 1, f. 4 (Dm) Original description
Poecilochroa bifaciata Baert et al., 2023: 10, f. 31A-C (mf, lapsus, on p. 67 at f. 31 spelling correct)
Poecilochroa capensis Strand, 1909 | | South Africa urn:lsid:nmbe.ch:spidersp:027731
Poecilochroa capensis Strand, 1909g: 552 (Dj) Original description
Poecilochroa capensis Tucker, 1923: 380 (Df)
Poecilochroa carinata Caporiacco, 1947 | | Uganda urn:lsid:nmbe.ch:spidersp:027732
Poecilochroa carinata Caporiacco, 1947c: 186, pl. 2, f. 38 (Df) Original description
Poecilochroa dayamibrookiana Barrion & Litsinger, 1995 | | Philippines urn:lsid:nmbe.ch:spidersp:027736
Poecilochroa dayamibrookiana Barrion & Litsinger, 1995: 187, f. 107a-j (Dmf) Original description
Poecilochroa exoculata Lissner, 2024 | | Canary Is. urn:lsid:nmbe.ch:spidersp:060059
Poecilochroa exoculata Lissner in Lissner, Suárez & López, 2024: 1019, f. 3, 4A-B, 5A-C, 6A-C, 7A-F, 9A-D (Df) Original description
Poecilochroa faradjensis Lessert, 1929 | | DR Congo urn:lsid:nmbe.ch:spidersp:027738
Poecilochroa faradjensis Lessert, 1929: 103, f. 1 (Df) Original description
Poecilochroa haplostyla Simon, 1907 | | São Tomé and Príncipe urn:lsid:nmbe.ch:spidersp:027741
Poecilochroa haplostylus Simon, 1907d: 235 (Dm) Original description
Poecilochroa insularis Kulczyński, 1911 | | Indonesia (Java) urn:lsid:nmbe.ch:spidersp:027744
Poecilochroa insularis Kulczyński, 1911b: 478, pl. 21, f. 28 (Df) Original description
Poecilochroa involuta Tucker, 1923 | | South Africa urn:lsid:nmbe.ch:spidersp:027745
Poecilochroa involuta Tucker, 1923: 382, pl. 10, f. 82 (Df) Original description
Poecilochroa involuta Dippenaar-Schoeman et al., 2021C: 44, 4 f. (f, undescribed m)
Poecilochroa joreungensis Paik, 1992 | | Korea urn:lsid:nmbe.ch:spidersp:027746
Poecilochroa joreungensis Paik, 1992a: 121, f. 12-21 (Df) Original description
Poecilochroa joreungensis Jung et al., 2005: 191, f. 30, 71, 94 (f)
Poecilochroa joreungensis Kim & Lee, 2013: 126, f. 87A-B (f)
Poecilochroa khodiar (Patel, 1988) | | India urn:lsid:nmbe.ch:spidersp:027922
Sergiolus khodiarae Patel, 1988d: 44, f. 1a-d (Df) Original description
Poecilochroa khodiar Sankaran, Caleb & Sebastian, 2021b: 2854 (T from Sergiolus)
Poecilochroa kuljitae (Tikader, 1982) | | India urn:lsid:nmbe.ch:spidersp:027828
Scopodes kuljitae Tikader, 1982b: 367, f. 170-174 (Df) Original description
Poecilochroa kuljitae Sankaran, Caleb & Sebastian, 2019b: 156, f. 1-4 (f, T from Scopoides)
Poecilochroa latefasciata Simon, 1893 | j | Peru urn:lsid:nmbe.ch:spidersp:027747
Poecilochroa latefasciata Simon, 1893e: 309 (Dj) Original description
Poecilochroa parangunifasciata Barrion & Litsinger, 1995 | | Philippines urn:lsid:nmbe.ch:spidersp:027752
Poecilochroa parangunifasciata Barrion & Litsinger, 1995: 191, f. 110a-i, 111a-e (Df) Original description
Poecilochroa patricia (Simon, 1878) | | France (Corsica) urn:lsid:nmbe.ch:spidersp:027753
Drassus patricius Simon, 1878a: 156 (Df) Original description
Poecilochroa patricia Simon, 1914: 185, 221, f. 399 (f)
Macarophaeus patricia Wunderlich, 2011: 46 (possible member of this genus)
Poecilochroa pauciaculeis Caporiacco, 1947 | j | Eritrea urn:lsid:nmbe.ch:spidersp:027754
Poecilochroa pauciaculeis Caporiacco, 1947c: 187 (Dj) Original description
Poecilochroa pauciaculeis Sherwood, 2024b: 207 (immature, species inquirenda)
Poecilochroa phyllobia (Thorell, 1871) | ? | Italy urn:lsid:nmbe.ch:spidersp:027756
Drassus bicolor Costa, 1834: 64 (D; preoccupied by Hahn, 1833, see Gnaphosa bicolor) [urn:lsid:nmbe.ch:spidersp:060372]
Gnaphosa phyllobia Thorell, 1871a: 201 (replacement name) Original description
Poecilochroa bicolor Reimoser, 1919: 172
Poecilochroa phyllobia Roewer, 1955c: 430
Poecilochroa rollini Berland, 1933 | | French Polynesia (Marquesas Is., Tuamotu) urn:lsid:nmbe.ch:spidersp:027758
Poecilochroa rollini Berland, 1933b: 46, f. 7-12 (Df) Original description
Poecilochroa rollini Dierkens & Ramage, 2016: 141, f. 17-18 (f)
Poecilochroa taeguensis Paik, 1992 | | Korea urn:lsid:nmbe.ch:spidersp:027763
Poecilochroa taeguensis Paik, 1992a: 123, f. 22-31 (Df) Original description
Poecilochroa taeguensis Jung et al., 2005: 192, f. 31, 72, 95 (f)
Poecilochroa taeguensis Kim & Lee, 2013: 127, f. 88A-B (f)
Poecilochroa tridotus Caleb & Mathai, 2013 | | India urn:lsid:nmbe.ch:spidersp:046540
Poecilochroa tridotus Caleb & Mathai, 2013: 2, f. 1a-f (Df) Original description
Poecilochroa tridotus Sankaran & Caleb, 2025: 137, f. 12-14 (f)
Poecilochroa trifasciata Mello-Leitão, 1918 | | Brazil urn:lsid:nmbe.ch:spidersp:027766
Poecilochroa trifasciata Mello-Leitão, 1918a: 71 (Df) Original description
Poecilochroa variana (C. L. Koch, 1839) | | Europe to Central Asia urn:lsid:nmbe.ch:spidersp:027767 TYPE SPECIES
Pythonissa variana C. L. Koch, 1839a: 65, f. 478 (Df) Original description
Drassus varianus Westring, 1851: 48
Melanophora variana Westring, 1861: 359 (Dm)
Gnaphosa variana Thorell, 1871a: 201
Poecilochroa variana Westring, 1874: 44
Drassus nigrofemoratus O. Pickard-Cambridge, 1874a: 385, pl. 51, f. 12 (Dm) [urn:lsid:nmbe.ch:spidersp:060373]
Poecilochroa variana Bösenberg, 1902: 318, pl. 30, f. 471 (mf)
Poecilochroa variana Simon, 1914: 183, 221 (S of Drassus nigrofemoratus)
Poecilochroa variana Reimoser, 1937a: 38, f. 91-92 (mf)
Poecilochroa variana Palmgren, 1943: 99, f. 106 (mf)
Poecilochroa variana Tullgren, 1946: 129, f. 37, pl. 21, f. 272-274 (mf)
Poecilochroa variana Azheganova, 1968: 99, f. 232, 236-237 (mf)
Poecilochroa variana Tystshenko, 1971: 101, f. 195 (m)
Poecilochroa variana Miller, 1971: 95, pl. VIII, f. 23, pl. IX, f. 25 (mf)
Poecilochroa variana Platnick & Shadab, 1981e: 4, f. 6-9 (mf)
Poecilochroa variana Grimm, 1985: 167, f. 193-198 (mf)
Poecilochroa variana Heimer & Nentwig, 1991: 438, f. 1153 (mf)
Poecilochroa variana Roberts, 1995: 118, f. (mf)
Poecilochroa variana Roberts, 1998: 124, f. (mf)
Poecilochroa variana Tuneva & Esyunin, 2002: 221, f. 20-24 (mf)
Poecilochroa variana Almquist, 2007: 416, f. 359a-g (mf)
Poecilochroa variana Murphy, 2007: 46, f. 290-291 (mf)
Poecilochroa variana Coşar, Danışman & Marusik, 2024: 113, f. 9A-F (m)
Poecilochroa viduata (Pavesi, 1883) | | Ethiopia urn:lsid:nmbe.ch:spidersp:027768
Drassus viduatus Pavesi, 1883: 49 (Df) Original description
Poecilochroa viduata Simon, 1893a: 372
Poecilochroa vittata Kulczyński, 1911 | | Indonesia (Java) urn:lsid:nmbe.ch:spidersp:027769
Poecilochroa vittata Kulczyński, 1911b: 480, pl. 21, f. 29 (Df) Original description

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