Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Heser Tuneva, 2004 urn:lsid:nmbe.ch:spidergen:03731

Transferred to other genera:

Heser infumatus (O. Pickard-Cambridge, 1872) -- see Berinda

In synonymy:

Heser incisupalpis (Levy, 1998) = Heser hierosolymitanus Levy, 1998 (Levy, 2009: 38, sub Zelotes)

Heser aradensis (Levy, 1998) | | Israel urn:lsid:nmbe.ch:spidersp:028116
Zelotes aradensis Levy, 1998c: 147, f. 37b, 116-119 (Dmf) Original description
Heser aradensis Tuneva, 2004: 323 (Tmf from Zelotes)
Heser aradensis Deltshev, 2016: 2, f. 1b,d, 2b,d,f, 3b,d, 4b,d, 5b,d,f, 6b,d (mf)
Heser bernardi (Marinaro, 1968) | | Spain, Algeria urn:lsid:nmbe.ch:spidersp:028134
Zelotes bernardi Marinaro, 1968: 689, f. 4-6 (Dmf) Original description
Zelotes bernardi Cárdenas & Barrientos, 2011: 108, f. 8-10 (m)
Heser bernardi Senglet, 2012: 522 (T from Zelotes)
Zelotes bernardi Frétey, 2023: 126 (year of description corrected)
Heser bonneti (Marinaro, 1968) | | Algeria urn:lsid:nmbe.ch:spidersp:028142
Zelotes bonneti Marinaro, 1968: 695, f. 13-14 (Dm) Original description
Heser bonneti Senglet, 2012: 522 (T from Zelotes)
Zelotes bonneti Frétey, 2023: 126 (year of description corrected)
Heser hierosolymitanus (Levy, 1998) | | Israel urn:lsid:nmbe.ch:spidersp:028240
Zelotes hierosolymitanus Levy, 1998c: 136, f. 90-91 (Df) Original description
Zelotes incisupalpis Levy, 1998c: 141, f. 102-103 (Dm) [urn:lsid:nmbe.ch:spidersp:028251]
Zelotes hierosolymitanus Levy, 2009: 38 (S of Zelotes incisupalpis)
Heser incisupalpis Senglet, 2012: 522 (T from Zelotes)
Heser hispanus Senglet, 2012 | | Spain urn:lsid:nmbe.ch:spidersp:046163
Heser hispanus Senglet, 2012: 525, f. 84-89 (Dmf) Original description
Heser malefactor Tuneva, 2004 | | Kazakhstan urn:lsid:nmbe.ch:spidersp:040615
Heser malefactor Tuneva, 2004: 323, f. 10-16 (Dmf) Original description
Heser nilicola (O. Pickard-Cambridge, 1874) | | Canary Is., Mediterranean. Introduced to Burkina Faso, USA, Mexico urn:lsid:nmbe.ch:spidersp:028322
Prosthesima nilicola O. Pickard-Cambridge, 1874a: 380, pl. 5, f. 8 (Dm) Original description
Prosthesima tantula Simon, 1878a: 88, pl. 14, f. 21 (Dm)
Prosthesima nilotica Simon, 1878a: 99
Melanophora nilotica Simon, 1893a: 366
Zelotes tantulus Simon, 1914: 160, 217, f. 312 (m)
Zelotes nilicolus Reimoser, 1919: 169
Zelotes nilicola Platnick & Shadab, 1983a: 186, f. 263-268 (m, Df)
Zelotes nilicola Melic, 1995c: 179, f. 1-2 (f)
Zelotes nilicola Levy, 1998c: 142, f. 104-107 (mf)
Zelotes nilicola Chatzaki, Thaler & Mylonas, 2003: 73, f. 82-86 (mf)
Zelotes nilicola Cornic, Ledoux & Villepoux, 2004: 17, f. 3A-B (mf)
Zelotes nilicola FitzPatrick, 2007: 123, f. 93-96 (mf)
Zelotes nilicola Wunderlich, 2011: 407, f. 20-22 (mf)
Heser nilicola Senglet, 2012: 522, f. 69-75 (Tmf from Zelotes)
Heser nilicola Jiménez et al., 2015: 324, f. 5b-c (m)
Heser nilicola Lecigne, 2016a: 111, f. 7I-K (m)
Heser nilicola Danışman, Ülker & Coşar, 2024: 1030, f. 2A-C, 3A-B (f)
Heser schmitzi (Kulczyński, 1899) | | Spain, Madeira, Canary Is. Introduced to USA urn:lsid:nmbe.ch:spidersp:028378
Prosthesima schmitzii Kulczyński, 1899: 359, pl. 6, f. 32 (Df) Original description
Zelotes schmitzi Platnick & Murphy, 1998: 118, f. 1-4 (f, Dm)
Heser schmitzi Senglet, 2012: 523, f. 76-83 (Tmf from Zelotes)
Heser stoevi Deltshev, 2016 | | Turkmenistan urn:lsid:nmbe.ch:spidersp:048611
Heser stoevi Deltshev, 2016: 3, f. 1a,c, 2a,c,e, 3a,c, 4a,c, 5a,c,e, 6a,c (Dmf) Original description
Heser vijayanagara Bosselaers, 2010 | | India urn:lsid:nmbe.ch:spidersp:043727
Heser vijayanagara Bosselaers, 2010b: 3, f. 1-8, 10-19 (Dmf) Original description

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