Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.

In synonymy:

Allodrassus Strand, 1906 = Echemella Strand, 1906 (Murphy & Russell-Smith, 2007: 16)

Maniana Strand, 1906 = Echemella Strand, 1906 (Murphy & Russell-Smith, 2007: 16)

Gen. Echemella Strand, 1906 urn:lsid:nmbe.ch:spidergen:02419
N.B.: considered a senior synonym of Allodrassus Strand, 1906 and Maniana Strand, 1906 by Murphy & Russell-Smith, 2007: 16.

Transferred to other genera:

Echemella aldabrae Strand, 1907 -- see Camillina

In synonymy:

Echemella defecta (Strand, 1906) = Echemella quinquedentata Strand, 1906 (Murphy & Russell-Smith, 2007: 17)

Echemella tridentata (Strand, 1906) = Echemella quinquedentata Strand, 1906 (Murphy & Russell-Smith, 2007: 17)

Echemella occulta (Benoit, 1965) | | Congo urn:lsid:nmbe.ch:spidersp:026573
Allodrassus occultus Benoit, 1965f: 307, f. 12-15 (Df, probably misplaced) Original description
Echemella pavesii (Simon, 1909) | | Ethiopia urn:lsid:nmbe.ch:spidersp:026574
Allodrassus pavesii Simon, 1909c: 34 (Df, probably misplaced) Original description
Echemella quinquedentata Strand, 1906 | | Ethiopia urn:lsid:nmbe.ch:spidersp:027110
Echemella quinquedentata Strand, 1906b: 610 (Df) Original description
Allodrassus tridentatus Strand, 1906b: 610 (Df)
Maniana defecta Strand, 1906b: 614 (Df)
Echemella quinquedentata Murphy & Russell-Smith, 2007: 17, f. 1-10 (Dm, f, S of Allodrassus tridentatus and Maniana defecta)
Echemella quinquedentata Murphy, 2007: 32, f. 106-107 (mf)
Echemella sinuosa Murphy & Russell-Smith, 2007 | | Ethiopia urn:lsid:nmbe.ch:spidersp:040604
Echemella sinuosa Murphy & Russell-Smith, 2007: 19, f. 11-13 (Dmf) Original description
Echemella strandi (Caporiacco, 1940) | | Ethiopia urn:lsid:nmbe.ch:spidersp:026575
Allodrassus strandi Caporiacco, 1940c: 833, f. 35 (Df, probably misplaced) Original description
Echemella tenuis Murphy & Russell-Smith, 2007 | | Ethiopia urn:lsid:nmbe.ch:spidersp:040605
Echemella tenuis Murphy & Russell-Smith, 2007: 21, f. 14-16 (Df) Original description

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