Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.

In synonymy:

Berlandia Dalmas, 1921 = Berlandina Dalmas, 1922 (Dalmas, 1922: 85, preoccupied, replacement name)

Gen. Berlandina Dalmas, 1922 urn:lsid:nmbe.ch:spidergen:02405
type Gnaphosa plumalis O. Pickard-Cambridge, 1872; gender feminine
N.B.: replacement name for Berlandia Dalmas, 1921: 266 [urn:lsid:nmbe.ch:spidergen:06117], preoccupied in Sparassidae by Lessert, 1921: 381.

Transferred to other genera:

Berlandina asiatica (Bösenberg & Strand, 1906) -- see Hitobia

Berlandina exornata (C. L. Koch, 1839) -- see Nomisia

In synonymy:

Berlandina afghana spinitarsis Denis, 1958 = Berlandina afghana Denis, 1958 (Marusik et al., 2014b: 416)

Berlandina atlantica (Dalmas, 1921) = Berlandina nigromaculata (Blackwall, 1865) (Berland, 1936: 74)

Berlandina chopardi Denis, 1955 = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Levy, 1995: 940)

Berlandina cinereoplumosa (Simon, 1878, T from Gnaphosa) = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Dalmas, 1921: 268, sub Berlandia)

Berlandina collisarenosa Wunderlich, 2023 = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Wunderlich, 2024b: 28)

Berlandina jovia Denis, 1947 = Berlandina venatrix (O. Pickard-Cambridge, 1874) (Levy, 1995: 942)

Berlandina litvinovi Fomichev & Marusik, 2017 = Berlandina charitonovi Ponomarev, 1979 (Esyunin, Ponomarev & Kabdrakhimov, 2024: 319)

Berlandina macrostigma Denis, 1966 = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Levy, 1995: 940)

Berlandina paludani Denis, 1958 = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Levy, 1995: 940)

Berlandina passerina (Simon, 1884, T from Callilepis) = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Dalmas, 1921: 268, sub Berlandia)

Berlandina rhodopis (L. Koch, 1875, T from Gnaphosa) = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Dalmas, 1921: 268, sub Berlandia)

Berlandina silacea (Simon, 1878, T from Pythonissa) = Berlandina cinerea (Menge, 1872) (Simon, 1914: 222, sub Pterotricha)

Berlandina xinjiangensis Hu & Wu, 1989 = Berlandina spasskyi Ponomarev, 1979 (Tuneva, 2004: 327)

Berlandina afghana Denis, 1958 | | Iran, Afghanistan, Pakistan urn:lsid:nmbe.ch:spidersp:047516
Berlandina afghana Denis, 1958b: 92, f. 14 (Df) Original description
Berlandina afghana spinitarsis Denis, 1958b: 92, f. 15 (Df) [urn:lsid:nmbe.ch:spidersp:059675]
Berlandina afghana Marusik et al., 2014b: 416, f. 1, 3a-b, 4, 5a-b, 6 (f, removed from S of Berlandina plumalis [rejecting Levy, 1995: 940], S of B. afghana spinitarsis [removed from S of B. plumalis, rejecting Levy, 1995: 940])
Berlandina apscheronica Dunin, 1984 | | Russia (Europe), Azerbaijan, Kazakhstan urn:lsid:nmbe.ch:spidersp:026680
Berlandina apscheronicus Dunin, 1984a: 49, f. 4 (Df) Original description
Berlandina apscheronica Ponomarev et al., 2011: 128, f. 3a-b (Dm)
Berlandina artaxerxes Zamani, Chatzaki, Esyunin & Marusik, 2021 | | Iran urn:lsid:nmbe.ch:spidersp:053225
Berlandina artaxerxes Zamani et al., 2021a: 74, f. 3A-D (Dm) Original description
Berlandina asbenica Denis, 1955 | | Niger urn:lsid:nmbe.ch:spidersp:026681
Berlandina asbenica Denis, 1955b: 113, f. 12 (Df) Original description
Berlandina avishur Levy, 2009 | | Israel urn:lsid:nmbe.ch:spidersp:042289
Berlandina avishur Levy, 2009: 3, f. 1-3 (Dm) Original description
Berlandina caspica Ponomarev, 1979 | | Russia (Europe to Central Asia), Caucasus, Iran, Central Asia, Mongolia urn:lsid:nmbe.ch:spidersp:026682
Berlandina caspica Ponomarev, 1979a: 921, f. 1-3 (Dmf) Original description
Berlandina apscheronica Tuneva, 2004: 327, f. 26-27 (f; misidentified per Marusik, Fomichev & Omelko, 2014a: 197)
Berlandina caspica Marusik, Fomichev & Omelko, 2014a: 197, f. 49a-d, 50a-c, 51-53, 54a-c, 55a-b, 56a-c (mf)
Berlandina caspica Zamani et al., 2017c: 61, f. 2A (f)
Berlandina charitonovi Ponomarev, 1979 | | Russia (Europe, Caucasus), Azerbaijan, Iran, Kazakhstan, Mongolia urn:lsid:nmbe.ch:spidersp:026683
Berlandina charitonovi Ponomarev, 1979a: 921, f. 4-6 (Dmf) Original description
Berlandina litvinovi Fomichev & Marusik, 2017b: 73, f. 9-19 (Dmf) [urn:lsid:nmbe.ch:spidersp:049312]
Berlandina charitonovi Esyunin, Ponomarev & Kabdrakhimov, 2024: 319 (S of Berlandina litvinovi)
Berlandina charitonovi Zamani et al., 2025b: 16, f. 11C, 12E-G (f)
Berlandina cinerea (Menge, 1872) | | Europe, Russia (Europe to South Siberia), Iran, Kazakhstan urn:lsid:nmbe.ch:spidersp:026684
Gnaphosa cinerea Menge, 1872: 319, pl. 57, f. 183 (Df) Original description
Gnaphosa cinerea Menge, 1875: 414, pl. 70, f. 183 (Dm)
Pythonissa silacea Simon, 1878a: 198 (Dm) [urn:lsid:nmbe.ch:spidersp:059681]
Pterotricha cinerea Chyzer & Kulczyński, 1897: 191, pl. 7, f. 29 (mf)
Callilepis nubivaga Simon, 1898e: 172 (misidentified per Simon, 1914: 222)
Pterotricha cinerea Simon, 1914: 191, 222, f. 409-410 (mf, S of Pythonissa silacea)
Berlandia cinerea Dalmas, 1921: 275, f. 51, 60 (mf)
Pythonissa cinerea Spassky, 1925: 3, f. 50 (f)
Berlandina cinerea Palmgren, 1943: 81, f. 85 (m)
Berlandina cinerea Tystshenko, 1971: 94, f. 192 (m)
Berlandina cinerea Miller, 1971: 77, pl. V, f. 18-19 (mf)
Berlandina cinerea Staręga, 1972: 83, f. 9-11, photo 7 (mf)
Berlandina cinerea Grimm, 1985: 37, f. 6, 19-25 (mf)
Berlandina cinerea Heimer & Nentwig, 1991: 416, f. 1095 (mf)
Berlandina cinerea Tuneva & Esyunin, 2002: 217, f. 1-6 (mf)
Berlandina cinerea Kovblyuk, 2003a: 881, f. 1.8-9, 12 (m)
Berlandina corcyraea (O. Pickard-Cambridge, 1874) | | Albania, Greece (incl. Corfu) urn:lsid:nmbe.ch:spidersp:026685
Gnaphosa corcyraea O. Pickard-Cambridge, 1874a: 376, pl. 51, f. 5 (Dm) Original description
Pythonissa corcycrea Simon, 1878a: 205
Pterotricha corcyraea Kulczyński, 1915: 917, pl. 66, f. 25-29 (m, Df)
Berlandia corcyraea Dalmas, 1921: 274, f. 39 (m)
Berlandina denisi Roewer, 1961 | | Afghanistan urn:lsid:nmbe.ch:spidersp:026686
Berlandina denisi Roewer, 1961c: 10, f. 10-11, 64 (Df) Original description
Berlandina deserticola (Dalmas, 1921) | | Algeria, Libya urn:lsid:nmbe.ch:spidersp:026687
Berlandia deserticola Dalmas, 1921: 273, f. 49, 58 (Dmf) Original description
Berlandina deserticola Al-Khazali & Fomichev, 2021: 70, f. 1h (f)
Berlandina drassodea (Caporiacco, 1934) | | Pakistan (Karakorum) urn:lsid:nmbe.ch:spidersp:026688
Berlandia drassodea Caporiacco, 1934a: 140, pl. 2, f. 5 (Dm) Original description
Berlandina hui Song, Zhu & Zhang, 2004 | | Kazakhstan, China urn:lsid:nmbe.ch:spidersp:040592
Berlandina potanini Hu & Wu, 1989: 251, f. 206.1-4 (mf; misidentified per Song, Zhu & Zhang, 2004: 26, 309)
Berlandina potanini Zhao, 1993: 116, f. 51a-b (mf; misidentified per Song, Zhu & Zhang, 2004: 26, 309)
Berlandina potanini Zhao, 1995: 834, f. 387a-b (mf; misidentified sensu Song, Zhu & Zhang, 2004: 26, 309)
Berlandina potanini Hu, 2001: 227, f. 122.1-4 (mf; misidentified per Song, Zhu & Zhang, 2004: 26, 309)
Berlandina hui Song, Zhu & Zhang, 2004: 26, 309, f. 13A-D (Dmf) Original description
Berlandina hui Marusik, Fomichev & Omelko, 2014a: 192, f. 17-19 (f)
Berlandina ilika Fomichev & Marusik, 2019 | | Kazakhstan urn:lsid:nmbe.ch:spidersp:051410
Berlandina ilika Fomichev & Marusik, 2019a: 14, f. 1-5, 7, 9-11 (Dm) Original description
Berlandina izzatullaevi Shodmonov & Fomichev, 2025 | | Uzbekistan urn:lsid:nmbe.ch:spidersp:064859
Berlandina izzatullaevi Shodmonov & Fomichev, 2025: 511, f. 1-9, 42 (Dm) Original description
Berlandina jabborovi Shodmonov & Fomichev, 2025 | | Uzbekistan urn:lsid:nmbe.ch:spidersp:064860
Berlandina jabborovi Shodmonov & Fomichev, 2025: 518, f. 24-32, 44 (Dm) Original description
Berlandina khalimovi Shodmonov & Fomichev, 2025 | | Uzbekistan urn:lsid:nmbe.ch:spidersp:064861
Berlandina khalimovi Shodmonov & Fomichev, 2025: 520, f. 33-41, 45 (Dm) Original description
Berlandina kolosvaryi Caporiacco, 1947 | j | Ethiopia urn:lsid:nmbe.ch:spidersp:026689
Berlandina kolosvaryi Caporiacco, 1947c: 190 (Dj) Original description
Berlandina kolosvaryi Sherwood, 2024b: 207 (immature, species inquirenda)
Berlandina koponeni Marusik, Fomichev & Omelko, 2014 | | Mongolia, China urn:lsid:nmbe.ch:spidersp:047059
Berlandina koponeni Marusik, Fomichev & Omelko, 2014a: 192, f. 30-37 (Dm) Original description
Berlandina koponeni Fomichev & Marusik, 2017b: 70, f. 1-4 (Df)
Berlandina koponeni Al-Khazali, 2020b: 446, f. 3K-L (m)
Berlandina meruana (Dalmas, 1921) | | Tanzania urn:lsid:nmbe.ch:spidersp:026690
Pythonissa plumalis Tullgren, 1910: 110, pl. 1, f. 26a (f; misidentified per Dalmas, 1921: 270)
Berlandia meruana Dalmas, 1921: 270 (Df) Original description
Berlandina mesopotamica Al-Khazali, 2020 | | Iraq, Iran urn:lsid:nmbe.ch:spidersp:052075
Berlandina mesopotamica Al-Khazali, 2020b: 444, f. 2A-C, 3A-H (Dm) Original description
Berlandina mesopotamica Al-Khazali & Fomichev, 2021: 70, f. 1a-e (f)
Berlandina mesopotamica Zamani et al., 2021a: 76, f. 3E-G, 4A-C (f)
Berlandina mishenini Marusik, Fomichev & Omelko, 2014 | | Mongolia urn:lsid:nmbe.ch:spidersp:047058
Berlandina mishenini Marusik, Fomichev & Omelko, 2014a: 192, f. 20-29 (Dm) Original description
Berlandina nabozhenkoi Ponomarev & Tsvetkov, 2006 | | Turkey, Russia (Europe), Georgia, Azerbaijan, Iran urn:lsid:nmbe.ch:spidersp:041364
Berlandina nabozhenkoi Ponomarev & Tsvetkov, 2006: 5, f. 1-2 (Dm) Original description
Berlandina nabozhenkoi Ponomarev, 2008b: 79, f. 1-2 (m, Df)
Berlandina nabozhenkoi Marusik, Fomichev & Omelko, 2014a: 210, f. 105-110 (m)
Berlandina nabozhenkoi Ponomarev et al., 2018a: 246, f. 1-2 (f)
Berlandina nabozhenkoi Seyyar & Demir, 2020d: 93, f. 2A-B, 3A-D, 4A-C (m)
Berlandina nabozhenkoi Seropian et al., 2024b: 187, f. 1-2 (f)
Berlandina nabozhenkoi Nuruyeva, Snegovaya & Aliyeva, 2026: 134, f. 1 (f)
Berlandina nakonechnyi Marusik, Fomichev & Omelko, 2014 | | Mongolia urn:lsid:nmbe.ch:spidersp:047061
Berlandina nakonechnyi Marusik, Fomichev & Omelko, 2014a: 204, f. 74-82 (Dm) Original description
Berlandina nakonechnyi Fomichev & Marusik, 2017b: 72, f. 5-8 (Df)
Berlandina nenilini Ponomarev & Tsvetkov, 2006 | | Kazakhstan urn:lsid:nmbe.ch:spidersp:041365
Berlandina nenilini Ponomarev & Tsvetkov, 2006: 7, f. 3 (Df) Original description
Berlandina nenilini Marusik, Fomichev & Omelko, 2014a: 191, f. 13-16 (f)
Berlandina nigromaculata (Blackwall, 1865) | | Cape Verde urn:lsid:nmbe.ch:spidersp:026691
Drassus nigromaculatus Blackwall, 1865a: 86 (Dmf) Original description
Pythonissa nigromaculata Simon, 1883: 304, pl. 8, f. 22 (f)
Berlandia atlantica Dalmas, 1921: 271, f. 47 (Df) [urn:lsid:nmbe.ch:spidersp:059676]
Berlandina nigromaculata Berland, 1936: 73, f. 9-11 (mf, S of Berlandina atlantica)
Berlandina nigromaculata Schmidt, 1999e: 5, f. 1 (m)
Berlandina nubivaga (Simon, 1878) | | Alps (France, Italy, Switzerland), North Macedonia, Bulgaria urn:lsid:nmbe.ch:spidersp:026692
Pythonissa nubivaga Simon, 1878a: 197 (Df) Original description
Pterotricha nubivaga Simon, 1914: 190, 222
Berlandia nubivaga Dalmas, 1921: 274, f. 50 (f)
Berlandina nubivaga Grimm, 1985: 40, f. 26-28 (f)
Berlandina nubivaga Heimer & Nentwig, 1991: 416, f. 1096 (f)
Berlandina nubivaga Isaia, Negro & Rolando, 2008: 180, f. 1-4 (f, Dm)
Berlandina obscurata Caporiacco, 1947 | | Tanzania urn:lsid:nmbe.ch:spidersp:026693
Berlandina obscurata Caporiacco, 1947c: 191, pl. 2, f. 41 (Df) Original description
Berlandina ovtsharenkoi Marusik, Fomichev & Omelko, 2014 | | Mongolia urn:lsid:nmbe.ch:spidersp:047060
Berlandina ovtsharenkoi Marusik, Fomichev & Omelko, 2014a: 195, f. 38-48 (Dmf) Original description
Berlandina piephoi Schmidt, 1994 | | Cape Verde urn:lsid:nmbe.ch:spidersp:026694
Berlandina piephoi Schmidt in Schmidt, Geisthardt & Piepho, 1994: 99, f. 12 (Df) Original description
Berlandina plumalis (O. Pickard-Cambridge, 1872) | | West Africa, Mediterranean to Central Asia, Iran, Afghanistan, Myanmar urn:lsid:nmbe.ch:spidersp:026695 TYPE SPECIES
Gnaphosa plumalis O. Pickard-Cambridge, 1872: 225, pl. 15, f. 3 (Dm) Original description
Gnaphosa rhodopis L. Koch, 1875b: 40, pl. 4, f. 4 (Df) [urn:lsid:nmbe.ch:spidersp:059682]
Pythonissa plumalis Simon, 1878a: 203 (Df)
Pythonissa cinereo-plumosa Simon, 1878a: 203 (Df) [urn:lsid:nmbe.ch:spidersp:059683]
Gnaphosa cinereo-plumosa Pavesi, 1880b: 358
Pythonissa passerina Simon, 1884n: 350, f. 7 (Df) [urn:lsid:nmbe.ch:spidersp:059684]
Callilepis plumalis Simon, 1893a: 382
Pythonissa plumalis Kulczyński, 1901a: 65, pl. l, f. 8-9, 11 (f)
Callilepis passerina Simon, 1905a: 170
Pythonissa plumalis Tullgren, 1910: 110, pl. 1, f. 26 (f; f f. 26a see Berlandina meruana)
Pterotricha plumalis Kulczyński, 1911a: 24, f. 19-20 (m)
Pterotricha plumalis Simon, 1914: 190, 222, f. 406-408 (mf)
Berlandia plumalis Dalmas, 1921: 268, f. 45, 52-53 (mf, S of Callilepis passerina, Gnaphosa cinereoplumosa and G. rhodopis)
Berlandina plumalis Dalmas, 1922: 85
Berlandina chopardi Denis, 1955b: 112, f. 10-11 (Dm) [urn:lsid:nmbe.ch:spidersp:059677]
Berlandina paludani Denis, 1958b: 91, f. 10-13 (Dm) [urn:lsid:nmbe.ch:spidersp:059680]
Berlandina plumalis Roewer, 1961c: 8, f. 8-9, 62 (f)
Berlandina paludani Roewer, 1961c: 9, f. 63 (m)
Berlandina macrostigma Denis, 1966e: 108, f. 10 (Df) [urn:lsid:nmbe.ch:spidersp:059679]
Berlandina plumalis Levy, 1995: 940, f. 53-56 (mf, S of Berlandina afghana [revalidated by Marusik et al., 2014b: 416], B. afghana spinitarsis [rejected by Marusik et al., 2014b: 416, see B. afghana], B. chopardi, B. macrostigma and B. paludani)
Berlandina plumalis Chatzaki, Thaler & Mylonas, 2002b: 609, f. 13-16 (mf)
Berlandina plumalis Murphy, 2007: 32, f. 102-103 (mf)
Berlandina plumalis Wunderlich, 2011: 39, f. 54, 54a (mf)
Berlandina plumalis Marusik et al., 2014b: 418, f. 7a-b, 8 (f)
Berlandina plumalis Zamani et al., 2015: 340, f. 2a-c (f)
Berlandina collisarenosa Wunderlich, 2023: 55, f. 119-121 (Df) [urn:lsid:nmbe.ch:spidersp:059445]
Berlandina plumalis Wunderlich, 2024b: 28, f. 59-50 (m, S of Berlandina collisarenosa; sometimes misspelled as collisarenaria)
Berlandina plumalis Shodmonov & Fomichev, 2025: 514, f. 10-23, 43 (mf)
Berlandina potanini Schenkel, 1963 | | Russia (South Siberia), Mongolia, China urn:lsid:nmbe.ch:spidersp:026696
Berlandia plumalis Schenkel, 1936b: 264, f. 86 (f; misidentified per Song, Zhu & Chen, 1999: 446)
Berlandia potanini Schenkel, 1963: 97, f. 56 (Df) Original description
Berlandina potanini Brignoli, 1983c: 565
Berlandina potanini Marusik & Logunov, 1995b: 179, f. 1-3 (f)
Berlandina potanini Song, Zhu & Chen, 1999: 446, f. 259A (f)
Berlandina potanini Song, Zhu & Zhang, 2004: 28, 310, f. 14A-E (f, Dm)
Berlandina potanini Marusik, Fomichev & Omelko, 2014a: 208, f. 94-100 (mf)
Berlandina propinqua Roewer, 1961 | | Kazakhstan, Uzbekistan, Afghanistan urn:lsid:nmbe.ch:spidersp:026697
Berlandina propinqua Roewer, 1961c: 10, f. 12-14, 65 (Df) Original description
Berlandina pulchra (Nosek, 1905) | | Turkey urn:lsid:nmbe.ch:spidersp:026698
Pterotricha pulchra Nosek, 1905b: 127, f. 9 (Df) Original description
Nomisia pulchra Dalmas, 1921: 289, f. 71 (f)
Berlandina pulchra Levy, 1995: 945 (T from Nomisia)
Berlandina punica (Dalmas, 1921) | | Algeria, Tunisia, Libya urn:lsid:nmbe.ch:spidersp:026699
Berlandia punica Dalmas, 1921: 270, f. 46, 54-55 (Dmf) Original description
Berlandina punica Caporiacco, 1936b: 77
Berlandina saraevi Ponomarev, 2008 | | Russia (Caucasus), Kazakhstan urn:lsid:nmbe.ch:spidersp:041584
Berlandina saraevi Ponomarev, 2008a: 55, f. 23-24 (Dm) Original description
Berlandina saraevi Marusik, Fomichev & Omelko, 2014a: 202, f. 69-73 (m)
Berlandina saraevi Ponomarev et al., 2019: 314, f. 6 (m)
Berlandina schenkeli Marusik & Logunov, 1995 | | Russia (South Siberia) urn:lsid:nmbe.ch:spidersp:026700
Berlandina schenkeli Marusik & Logunov, 1995b: 180, f. 4-7 (Dmf) Original description
Berlandina schenkeli Marusik, Fomichev & Omelko, 2014a: 208, f. 101-104 (mf)
Berlandina shnitnikovi (Spassky, 1934) | | Kazakhstan urn:lsid:nmbe.ch:spidersp:027808
Pterotricha shnitnikovi Spassky, 1934c: 137, f. 4-5 (Dmf) Original description
Berlandina shnitnikovi Ponomarev & Dvadnenko, 2012: 44 (T from Pterotricha)
Berlandina shumskyi Kovblyuk, 2003 | | Ukraine urn:lsid:nmbe.ch:spidersp:038994
Berlandina shumskyi Kovblyuk, 2003a: 880, f. 1.1-7, 10-11 (Dmf) Original description
Berlandina spasskyi Ponomarev, 1979 | | Russia (Europe), Kazakhstan, Mongolia, China urn:lsid:nmbe.ch:spidersp:026701
Berlandina spasskyi Ponomarev, 1979a: 922, f. 7 (Df) Original description
Berlandina xinjiangensis Hu & Wu, 1989: 253, f. 207.1-4 (Dmf) [urn:lsid:nmbe.ch:spidersp:026704]
Berlandina xinjiangensis Song, Zhu & Zhang, 2004: 30, f. 15A-E (mf)
Berlandina spasskyi Tuneva, 2004: 327, f. 28-29 (f, S of Berlandina xinjiangensis)
Berlandina spasskyi Ponomarev & Tsvetkov, 2006: 7, f. 4-5 (m)
Berlandina spasskyi Marusik, Fomichev & Omelko, 2014a: 200, f. 57-68 (mf)
Berlandina ubsunurica Marusik & Logunov, 1995 | | Russia (South Siberia), Mongolia urn:lsid:nmbe.ch:spidersp:026702
Berlandina ubsunurica Marusik & Logunov, 1995b: 181, f. 8-10 (Dm) Original description
Berlandina ubsunurica Marusik, Fomichev & Omelko, 2014a: 189, f. 1-12 (m, Df)
Berlandina venatrix (O. Pickard-Cambridge, 1874) | | Libya, Egypt urn:lsid:nmbe.ch:spidersp:026703
Gnaphosa venatrix O. Pickard-Cambridge, 1874a: 375, pl. 51, f. 4 (Dm) Original description
Pythonissa venatrix Simon, 1878a: 205
Berlandia venatrix Dalmas, 1921: 272, f. 48, 56-57 (m, Df)
Berlandina jovia Denis, 1947a: 63, pl. 3, f. 11 (Df) [urn:lsid:nmbe.ch:spidersp:059678]
Berlandina venatrix Levy, 1995: 942, f. 57-60 (mf, S of Berlandina jovia)
Berlandina venatrix Al-Khazali, 2020b: 446, f. 3I-J (m)
Berlandina venatrix Al-Khazali & Fomichev, 2021: 70, f. 1f-g (f)
Berlandina venatrix Abdel-Ghani, Hassan & Sallam, 2023: 33, f. 7a-b (mf)
Berlandina yakovlevi Marusik, Fomichev & Omelko, 2014 | | Mongolia urn:lsid:nmbe.ch:spidersp:047062
Berlandina yakovlevi Marusik, Fomichev & Omelko, 2014a: 204, f. 83-90, 91a-b, 92-93 (Dmf) Original description
Berlandina yakovlevi Fomichev & Marusik, 2019a: 14, f. 6, 8 (m)

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