Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.

In synonymy:

Macedoniella Drensky, 1935 = Anagraphis Simon, 1893 (Bosmans, 2014b: 38)

Gen. Anagraphis Simon, 1893 urn:lsid:nmbe.ch:spidergen:02362
N.B.: transferred from the Prodidomidae to the Gnaphosidae by Platnick & Baehr, 2006: 5; could not be placed in a subfamily by Rodrigues & Rheims, 2020b: 661; considered a senior synonym of Macedoniella Drensky, 1935: 107 [urn:lsid:nmbe.ch:spidergen:02190] (type M. karamani Drensky, 1935 = Anagraphis ochracea (L. Koch, 1867)) by Bosmans, 2014b: 38.

In synonymy:

Anagraphis aculeatus (Charitonov, 1946) = Anagraphis pallens Simon, 1893 (Khasayeva & Huseynov, 2017: 126, removed from S of Berinda amabilis)

Anagraphis karamani (Drensky, 1935, T from Macedoniella=Anagraphis) = Anagraphis ochracea L. Koch, 1867 (Bosmans, 2014b: 38)

Anagraphis pallida (Hadjissarantos, 1940, T from Talanites) = Anagraphis ochracea L. Koch, 1867 (Bosmans, 2014b: 38)

Anagraphis vitellina Simon, 1897 = Anagraphis pallens Simon, 1893 (Levy, 1999c: 218)

Anagraphis incerta Caporiacco, 1941 | | Ethiopia urn:lsid:nmbe.ch:spidersp:026416
Anagraphis incerta Caporiacco, 1941b: 89, f. 31-32 (Dmf) Original description
Anagraphis maculosa Denis, 1958 | | Afghanistan urn:lsid:nmbe.ch:spidersp:026417
Anagraphis maculosa Denis, 1958b: 84, f. 1 (Df) Original description
Anagraphis minima Caporiacco, 1947 | | East Africa urn:lsid:nmbe.ch:spidersp:026418
Anagraphis minima Caporiacco, 1947c: 188, pl. 2, f. 39 (Df) Original description
Anagraphis ochracea (L. Koch, 1867) | | Italy, Albania, North Macedonia, Greece, Turkey (Europe), Russia (Caucasus)? urn:lsid:nmbe.ch:spidersp:024239
Liocranum ochraceum L. Koch, 1867c: 864 (Df) Original description
Macedoniella karamani Drensky, 1935: 109, f. 6 (Df) [urn:lsid:nmbe.ch:spidersp:047515]
Talanites pallidus Hadjissarantos, 1940: 79, f. 23-25 (Dmf) [urn:lsid:nmbe.ch:spidersp:028015]
Anagraphis pallida Chatzaki, Thaler & Mylonas, 2002b: 605, f. 3-8 (mf,T from Talanites)
Macedoniella karamani Deltshev, 2003: 143 (species inquirenda)
Macedoniella karamani Wunderlich, 2011: 42, f. 150-152 (f)
Anagraphis ochracea Bosmans, 2014b: 38, f. 1 (f, T from Liocranum, S of Anagraphis pallida and Macedoniella karamani)
Anagraphis ochracea Demircan & Topçu, 2015: 177, f. 1A-C (f)
Anagraphis ochracea Ponomarev & Shmatko, 2020b: 304, f. 17-18 (f; N.B.: probably misidentified)
Anagraphis ochracea Caria et al., 2021: 322, f. 2A-B (m)
Anagraphis ochracea Lenzini et al., 2022: 84, f. 1a-d, 3a-c, 4a-b, 5a-b, 6, 7a-b (mf)
Anagraphis pallens Simon, 1893 | | Libya, Malta, Greece, Turkey, Israel, Syria, Russia (Europe, Caucasus), Georgia, Azerbaijan, Iran, Kazakhstan, Central Asia urn:lsid:nmbe.ch:spidersp:026419
Anagraphis pallens Simon, 1893e: 308 (Dm) Original description
Anagraphis vitellina Simon, 1897b: 465 (Df) [urn:lsid:nmbe.ch:spidersp:054331]
Talanites aculeatus Charitonov, 1946: 26, f. 43 (Dm; see also Charitonov, 1969: 108) [urn:lsid:nmbe.ch:spidersp:028003]
Anagraphis pallens Lawrence, 1947: 11 (Df)
Anagraphis pallens Levy, 1999c: 216, f. 1-7 (mf, S of Anagraphis vitellina)
Anagraphis pallens Chatzaki, Thaler & Mylonas, 2002a: 567, f. 2-7 (mf)
Anagraphis pallens Topçu, Seyyar & Kunt, 2005: 85, f. 1-2 (mf)
Anagraphis pallens Platnick & Baehr, 2006: 5 (type locality in South Africa considered erroneous)
Anagraphis pallens Murphy, 2007: 41, f. 230-231 (mf)
Anagraphis pallens Wunderlich, 2011: 37, f. 32-34 (mf)
Anagraphis pallens Ramírez, 2014: 268, f. 183C (f)
Anagraphis pallens Khasayeva & Huseynov, 2017: 126, f. 1-8 (mf, S of Talanites aculeatus, which is removed from S of Berinda amabilis, contra Chatzaki, Thaler & Mylonas, 2002a: 570)
Anagraphis pallens Seropian et al., 2023: 238, f. 14-15, S1 (mf)
Anagraphis pluridentata Simon, 1897 | | Syria urn:lsid:nmbe.ch:spidersp:026420
Anagraphis pluridentata Simon, 1897b: 465 (Df) Original description
Anagraphis pori Levy, 1999 | | Israel urn:lsid:nmbe.ch:spidersp:026421
Anagraphis pori Levy, 1999c: 219, f. 8-9 (Dm) Original description

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