Halonoproctidae Pocock, 1901

Family |   ACCEPTED
N.B.: described as a subfamily of Ctenizidae by Pocock, 1901e: 207 (type genus Halonoproctus, which is a synonym of Cyclocosmia), elevated to family level by Pocock, 1903h: 352; included in Ctenizidae by Simon, 1903a: 884 (sub Ctenizinae, Cyclocosmieae) and listed there by the subsequent catalogues; recircumscribed and splitted from Ctenizidae by Godwin et al., 2018: 307; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 487, f. 5.

In synonymy:

Pachylomerides Strand, 1934 = Ummidia Thorell, 1875 (Roewer, 1955c: 1715)

Pachylomerus Ausserer, 1871 = Ummidia Thorell, 1875 (Roewer, 1955c: 1715, automatically with its replacement name Pachylomerides)

Sterrhochrotus Simon, 1892 = Ummidia Thorell, 1875 (Decae et al., 2019: 519, by transfer of the type species, rejecting Raven, 1985a: 159, syn. of Cteniza)

Homonym replaced:

Pachylomerus Ausserer, 1871 -- see Ummidia Thorell, 1875 - in Strand, 1934

Gen. Ummidia Thorell, 1875 urn:lsid:nmbe.ch:spidergen:00071
N.B.: transferred from the Ctenizidae to the Halonoproctidae by Godwin et al., 2018: 307; Ummidia is the type genus of the subfamily Ummidiinae (see Ortiz, 2007: 395); considered a senior synonym of Pachylomerides Strand, 1934: 273 [urn:lsid:nmbe.ch:spidergen:05052] by Roewer, 1955c: 1715; Pachylomerides is a replacement name for Pachylomerus Ausserer, 1871a: 145 [urn:lsid:nmbe.ch:spidergen:05051] (type P. glaber Doleschall, 1871), preoccupied in Coleoptera by Bertolini, 1849; considered a senior synonym of Sterrhochrotus Simon, 1892a: 97 [urn:lsid:nmbe.ch:spidergen:05044] (type Cteniza ferghanensis Kroneberg, 1875) by Decae et al., 2019: 519 (by transfer of the type species), rejecting the synonymy with Cteniza Latreille, 1829, considered by Raven, 1985a: 159; Conothele Thorell, 1878 is considered indistinguishable from Ummidia by Decae, 2010: 329.

Transferred to other genera:

Ummidia doenitzi (Kishida, 1913) -- see Conothele

Ummidia fragaria (Dönitz, 1887) -- see Conothele

Ummidia granulosa (Kishida, 1927) -- see Conothele

Ummidia gressitti Roewer, 1963 -- see Conothele

Ummidia midzusawai (Kishida, 1921) -- see Conothele

Ummidia mirandus (Kishida, 1921) -- see Conothele

Ummidia nawai (Kishida, 1921) -- see Conothele

Ummidia taiwanensis Tso, Haupt & Zhu, 2003 -- see Conothele

In synonymy:

Ummidia absoluta (Gertsch & Mulaik, 1940) = Ummidia audouini Lucas, 1836 (Godwin & Bond, 2021: 16)

Ummidia carabivora emarginata (Atkinson, 1886) = Ummidia carabivora Atkinson, 1886 (Godwin & Bond, 2021: 37)

Ummidia celsa (Gertsch & Mulaik, 1940) = Ummidia funerea Gertsch, 1936 (Godwin & Bond, 2021: 63)

Ummidia lanthanus (Valerio, 1988, T from Hebestatis) = Ummidia rugosa Karsch, 1880 (Godwin & Bond, 2021: 128)

Ummidia occidentalis (Simon, 1909) = Ummidia aedificatoria Westwood, 1840 (Decae, 2010: 337)

Ummidia pygmaea (Chamberlin & Ivie, 1945) = Ummidia beatula Gertsch & Mulaik, 1940 (Godwin & Bond, 2021: 55)

Ummidia tuobita (Chamberlin, 1917) = Ummidia audouini Lucas, 1836 (Godwin & Bond, 2021: 16)

Nomen dubium:

Ummidia oaxacana (Chamberlin, 1925d: 212, f; Mexico, originally in Pachylomerus) -- Godwin & Bond, 2021: 16

Nomen nudum:

Ummidia tsushimensis (Kishida, sub Pachylomerus) -- Brignoli, 1983c: 111

Ummidia aedificatoria (Westwood, 1840) | | Morocco urn:lsid:nmbe.ch:spidersp:000605
Actinopus aedificatorius Westwood, 1840: 175, pl. 10, f. 1-25 (Df) Original description
Sphodros aedificatorius Walckenaer, 1841: 438
Cteniza aedificatoria Ausserer, 1871a: 155
Pachylomerus aedificatorius Simon, 1892a: 88, f. 85-87
Pachylomerus occidentalis Simon, 1909b: 8 (Df) [urn:lsid:nmbe.ch:spidersp:000621]
Pachylomerus occidentalis Frade & Bacelar, 1931c: 512, f. 5-6 (f)
Ummidia aedificatoria Decae, 2010: 337, f. 15, 19 (f, S of Ummidia occidentalis)
Ummidia algarve Decae, 2010 | | Portugal urn:lsid:nmbe.ch:spidersp:043497
Pachylomerus aedificatorius O. Pickard-Cambridge, 1908b: 817, pl. 1, f. 1-6 (f, misidentified)
Pachylomerus piceus Frade & Bacelar, 1931c: 510, f. 3-4 (mf, misidentified)
Pachylomerus piceus Bacelar, 1937: 1568, f. 1-2 (mf, misidentified)
Ummidia algarve Decae, 2010: 329, f. 2-3, 10-11, 17, 21, 23 (Dmf) Original description
Ummidia algeriana (Lucas, 1846) | | Algeria, Tunisia urn:lsid:nmbe.ch:spidersp:043499
Actinopus algerianus Lucas, 1846: 96, pl. 1, f. 5 (Df) Original description
Pachylomerus aedificatorius Simon, 1892a: 88, f. 86 (f, misidentified)
Pachylomerus aedificatorius Simon, 1903a: 887, f. 1048 (f, misidentified)
Pachylomerus aedificatorius Frade & Bacelar, 1931c: 509, f. 1-2 (f, misidentified)
Ummidia algeriana Decae, 2010: 337, f. 14, 18 (removed f from S of U. aedificatoria)
Ummidia anaya Godwin & Bond, 2021 | | Mexico urn:lsid:nmbe.ch:spidersp:052971
Ummidia anaya Godwin & Bond, 2021: 105, f. 3I, 42A-E (Dm) Original description
Ummidia armata (Ausserer, 1875) | | Unknown urn:lsid:nmbe.ch:spidersp:000606
Pachylomerus armatus Ausserer, 1875: 143 (Df) Original description
Ummidia asperula (Simon, 1889) | | Venezuela urn:lsid:nmbe.ch:spidersp:000607
Pachylomerus asperulus Simon, 1889m: 179 (Df) Original description
Ummidia asperula Godwin & Bond, 2021: 152, f. 68A-B, 69A-E (Dmf)
Ummidia asperula Dupérré & Tapia, 2021: 270, f. 12A-B, 13A-D (f)
Ummidia audouini (Lucas, 1836) | | USA urn:lsid:nmbe.ch:spidersp:000608
Pachyloscelis audouinii Lucas, 1836f: 5 (Df) Original description
Actinopus audouini Lucas, 1837: 387, pl. 13, f. 1-5 (f)
Sphodros audouini Walckenaer, 1841: 436
Mygale solstitialis Hentz, 1842: 56, pl. 7, f. 2 (Df)
Mygale carolinensis Hentz, 1842: 56, pl. 7, f. 3 (Df)
Pachylomerus carolinensis Ausserer, 1871a: 147
Pachylomerus turris Atkinson, 1886: 136, pl. 5, f. 5 (Df)
Pachylomerus audouini Simon, 1891f: 312 (S of Mygale carolinensis and M. solstitialis)
Pachylomerus audouini Banks, 1910: 2 (S of Pachylomerus turris)
Pachylomerus tuobitus Chamberlin, 1917: 33, pl. 1, f. 6-8 (Dm) [urn:lsid:nmbe.ch:spidersp:000626]
Pachylomerides absolutus Gertsch & Mulaik, 1940: 311, f. 10 (Df) [urn:lsid:nmbe.ch:spidersp:000604]
Pachylomerides tuobitus Chamberlin & Ivie, 1945a: 556, f. 13-15 (m, Df)
Ummidia audouini Roddy, 1957: 286, f. 5-6 (Dm)
Ummidia absoluta Brignoli, 1983c: 117
Ummidia audouini Godwin & Bond, 2021: 16, f. 3A-B 4A-B, 5A-E (mf; S of Ummidia absoluta and U. tuobita)
Ummidia beatula (Gertsch & Mulaik, 1940) | | USA urn:lsid:nmbe.ch:spidersp:000609
Pachylomerides beatulus Gertsch & Mulaik, 1940: 312, f. 11 (Df) Original description
Pachylomerides pygmaeus Chamberlin & Ivie, 1945a: 558, f. 11-12 (Dm) [urn:lsid:nmbe.ch:spidersp:000623]
Ummidia beatula Brignoli, 1983c: 117
Ummidia pygmaea Brignoli, 1983c: 117
Ummidia beatula Godwin & Bond, 2021: 55, f. 18A-B, 19A-E (Dmf, S of Ummidia pygmaea)
Ummidia brandicarlileae Godwin & Bond, 2021 | | Mexico urn:lsid:nmbe.ch:spidersp:052973
Ummidia brandicarlileae Godwin & Bond, 2021: 110, f. 45A-B, 46A-E (Dmf) Original description
Ummidia carabivora (Atkinson, 1886) | | USA urn:lsid:nmbe.ch:spidersp:000610
Pachylomerus carabivorus Atkinson, 1886: 133, pl. 5, f. 1-4 (Df) Original description
Pachylomerus quadrispinosus Atkinson, 1886: 133, pl. 5, f. 21 (D)
Pachylomerus carabivorus emarginatus Atkinson, 1886: 134 (Df) [urn:lsid:nmbe.ch:spidersp:000611]
Pachylomerus carabivorus Baerg, 1928: 1, pl. 1
Pachylomerus carabivorus Petrunkevitch, 1929a: 511, pl. 1, f. 1-9
Ummidia carabivora Godwin & Bond, 2021: 37, f. 10A-B, 11A-E (Dm, f, S of Ummidia carabivora emarginata)
Ummidia carlosviquezi Godwin & Bond, 2021 | | Nicaragua, Costa Rica urn:lsid:nmbe.ch:spidersp:052979
Ummidia carlosviquezi Godwin & Bond, 2021: 132, f. 57A-B, 58A-E (Dmf) Original description
Ummidia cerrohoya Godwin & Bond, 2021 | | Panama urn:lsid:nmbe.ch:spidersp:052982
Ummidia cerrohoya Godwin & Bond, 2021: 144, f. 63A-B (Df) Original description
Ummidia colemanae Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052960
Ummidia colemanae Godwin & Bond, 2021: 61, f. 20A-E (Dm) Original description
Ummidia cuicatec Godwin & Bond, 2021 | | Mexico urn:lsid:nmbe.ch:spidersp:052972
Ummidia cuicatec Godwin & Bond, 2021: 107, f. 43A-B, 44A-E (Dmf) Original description
Ummidia erema (Chamberlin, 1925) | | Panama urn:lsid:nmbe.ch:spidersp:000613
Pachylomerus eremus Chamberlin, 1925d: 211 (Df) Original description
Ummidia erema Godwin & Bond, 2021: 145, f. 64A-B, 65A-E (Dmf)
Ummidia ferghanensis (Kroneberg, 1875) | j | Central Asia urn:lsid:nmbe.ch:spidersp:000535
Cteniza ferghanensis Kroneberg, 1875: 27, pl. 3, f. 18 (Dj) Original description
Sterrhochrotus ferghanensis Simon, 1892a: 97
Ummidia ferghanensis Decae et al., 2019: 519 (T from Cteniza)
Ummidia frankellerae Godwin & Bond, 2021 | | Belize urn:lsid:nmbe.ch:spidersp:052975
Ummidia frankellerae Godwin & Bond, 2021: 118, f. 49A-B (Df) Original description
Ummidia funerea (Gertsch, 1936) | | USA urn:lsid:nmbe.ch:spidersp:000615
Pachylomerus funereus Gertsch, 1936a: 1, f. 1-2 (Dm) Original description
Pachylomerides celsus Gertsch & Mulaik, 1940: 313, f. 14-15 (Dm) [urn:lsid:nmbe.ch:spidersp:000612]
Ummidia celsa Brignoli, 1983c: 117
Ummidia funerea Godwin & Bond, 2021: 63, f. 21A-B, 22A-E (m, Df, S of Ummidia celsa)
Ummidia gabrieli Godwin & Bond, 2021 | | Mexico urn:lsid:nmbe.ch:spidersp:052967
Ummidia gabrieli Godwin & Bond, 2021: 90, f. 34A-B, 35A-E (Dmf) Original description
Ummidia gandjinoi (Andreeva, 1968) | | Tajikistan urn:lsid:nmbe.ch:spidersp:000616
Pachylomerus gandjinoi Andreeva, 1968: 68, f. 1 (Dm) Original description
Pachylomerus ganjinoi Andreeva, 1976: 13, f. 1-5 (m)
Ummidia gandjinoi Brignoli, 1983c: 117
Ummidia gandjinoi Zonstein, 2008: 151, f. 1-6 (Df)
Ummidia gandjinoi Zonstein, 2018a: 17, f. 1, 3, 5-8 (m)
Ummidia gertschi Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052965
Ummidia gertschi Godwin & Bond, 2021: 80, f. 30A-B, 31A-F (Dmf) Original description
Ummidia gingoteague Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052955
Ummidia gingoteague Godwin & Bond, 2021: 33, f. 3D-E, 8A-B, 9A-E (Dmf) Original description
Ummidia glabra (Doleschall, 1871) | | Brazil urn:lsid:nmbe.ch:spidersp:000617
Pachylomerus glaber Doleschall in Ausserer, 1871a: 146 (Df) Original description
Ummidia hondurena Godwin & Bond, 2021 | | Honduras, El Salvador urn:lsid:nmbe.ch:spidersp:052976
Ummidia hondurena Godwin & Bond, 2021: 122, f. 3D, 51A-B, 52A-E (Dmf) Original description
Ummidia huascazaloya Godwin & Bond, 2021 | | Mexico urn:lsid:nmbe.ch:spidersp:052970
Ummidia huascazaloya Godwin & Bond, 2021: 101, f. 3F, 40A-B, 41A-E (Dmf) Original description
Ummidia insularis Santos, Ortiz & Sánchez-Ruiz, 2022 | | Dominican Rep. urn:lsid:nmbe.ch:spidersp:053711
Ummidia insularis Santos, Ortiz & Sánchez-Ruiz, 2022: 443, f. 1A-F, 2A-F, 3A-G, 4A-F, 5A-F (Dmf) Original description
Ummidia macarthuri Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052959
Ummidia macarthuri Godwin & Bond, 2021: 51, f. 16A-B, 17A-E (Dmf) Original description
Ummidia matagalpa Godwin & Bond, 2021 | | Nicaragua urn:lsid:nmbe.ch:spidersp:052978
Ummidia matagalpa Godwin & Bond, 2021: 127, f. 54A-B (Df) Original description
Ummidia mercedesburnsae Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052962
Ummidia mercedesburnsae Godwin & Bond, 2021: 70, f. 24A-B, 25A-E (Dmf) Original description
Ummidia mischi Zonstein, 2014 | | Afghanistan urn:lsid:nmbe.ch:spidersp:047173
Ummidia mischi Zonstein, 2014: 269, f. 1-6 (Dm) Original description
Ummidia mischi Zonstein, 2018a: 18, f. 2, 4, 9 (m)
Ummidia modesta (Banks, 1901) | | USA urn:lsid:nmbe.ch:spidersp:000618
Pachylomerus modestus Banks, 1901b: 570, pl. 33, f. 1 (Dm) Original description
Ummidia modesta Godwin & Bond, 2021: 75, f. 27A-B, 28A-E (m, Df)
Ummidia neblina Godwin & Bond, 2021 | | Venezuela urn:lsid:nmbe.ch:spidersp:052985
Ummidia neblina Godwin & Bond, 2021: 155, f. 70A-B (Df) Original description
Ummidia neilgaimani Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052954
Ummidia neilgaimani Godwin & Bond, 2021: 27, f. 3G-H, 6A-B, 7A-E (Dmf) Original description
Ummidia nidulans (Fabricius, 1787) | | Jamaica, West Indies urn:lsid:nmbe.ch:spidersp:000619
Aranea nidulans Fabricius, 1787: 343 (Df) Original description
Mygale nidulans Latreille, 1804a: 166
Cteniza nidulans Sells, 1837: 207, pl. 70, f. 1-9 (f)
Cteniza venatoria C. L. Koch, 1838: 12, f. 345 (Df)
Actinopus nidulans Westwood, 1840: 173
Actinopus venatorius C. L. Koch, 1850: 75
Mygalodonta nidulans Simon, 1864: 75
Pachylomerus nidulans Ausserer, 1871a: 147
Pachylomerus nidulans Simon, 1892a: 87, f. 88-89
Ummidia nidulans Rudloff, 1996b: 48, f. 1-6, 9 (f)
Ummidia nidulans Godwin & Bond, 2021: 159, f. 73A-B (f)
Ummidia okefenokee Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052957
Ummidia okefenokee Godwin & Bond, 2021: 46, f. 14A-E (Dm) Original description
Ummidia paulacushingae Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052963
Ummidia paulacushingae Godwin & Bond, 2021: 73, f. 26A-E (Dm) Original description
Ummidia pesiou Godwin & Bond, 2021 | | Mexico urn:lsid:nmbe.ch:spidersp:052968
Ummidia pesiou Godwin & Bond, 2021: 94, f. 36A-B (Df) Original description
Ummidia picea Thorell, 1875 | | Spain urn:lsid:nmbe.ch:spidersp:043498
Ummidia picea Thorell, 1875a: 102 (Dm) Original description
Ummidia picea Thorell, 1875c: 121 (Dm)
Pachylomerus piceus Frade & Bacelar, 1931c: 510, f. 4bis (not f. 3-4)
Ummidia picea Decae, 2010: 335, f. 12-13, 16, 20, 22 (removed m from S of U. aedificatoria, Df)
Ummidia aedificatoria Le Peru, 2011: 72, f. 35 (m; adapted from Frade & Bacelar, 1931c: 510, f. 4bis)
Ummidia pustulosa (Becker, 1879) | | Mexico urn:lsid:nmbe.ch:spidersp:000622
Pachylomerus pustulosus Becker, 1879d: 140 (Df) Original description
Pachylomerus pustulosus Becker, 1881b: 44, pl. 2, f. 2 (f)
Ummidia pustulosa Godwin & Bond, 2021: 99, f. 39A-B (f)
Ummidia quepoa Godwin & Bond, 2021 | | Costa Rica urn:lsid:nmbe.ch:spidersp:052981
Ummidia quepoa Godwin & Bond, 2021: 141, f. 61A-B, 62A-E (Dmf) Original description
Ummidia quijichacaca Godwin & Bond, 2021 | | Colombia urn:lsid:nmbe.ch:spidersp:052983
Ummidia quijichacaca Godwin & Bond, 2021: 149, f. 66A-B (Df) Original description
Ummidia richmond Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052958
Ummidia richmond Godwin & Bond, 2021: 48, f. 15A-E (Dm) Original description
Ummidia riverai Godwin & Bond, 2021 | | Guatemala urn:lsid:nmbe.ch:spidersp:052974
Ummidia riverai Godwin & Bond, 2021: 116, f. 48A-E (Dm) Original description
Ummidia rodeo Godwin & Bond, 2021 | | Mexico urn:lsid:nmbe.ch:spidersp:052969
Ummidia rodeo Godwin & Bond, 2021: 95, f. 37A-B, 38A-E (Dmf) Original description
Ummidia rongodwini Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052956
Ummidia rongodwini Godwin & Bond, 2021: 41, f. 12A-B, 13A-E (Dmf) Original description
Ummidia rosillos Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052961
Ummidia rosillos Godwin & Bond, 2021: 67, f. 23A-E (Dm) Original description
Ummidia rugosa (Karsch, 1880) | | Nicaragua, Costa Rica urn:lsid:nmbe.ch:spidersp:000624
Pachylomerus rugosus Karsch, 1880c: 388 (Df) Original description
Hebestatis lanthanus Valerio, 1988: 228, f. 1-9 (Dmf) [urn:lsid:nmbe.ch:spidersp:000544]
Ummidia rugosa Godwin & Bond, 2021: 128, f. 55A-C, 56A-E (mf, S of Hebestatis lanthanus)
Ummidia salebrosa (Simon, 1892) | | St. Vincent urn:lsid:nmbe.ch:spidersp:000625
Pachylomerus salebrosus Simon, 1892e: 530 (Df) Original description
Ummidia salebrosa Godwin & Bond, 2021: 158, f. 72A-B (f)
Ummidia solana Echeverri, Gómez Torres, Pinel & Perafán, 2023 | | Colombia urn:lsid:nmbe.ch:spidersp:058313
Ummidia solana Echeverri et al., 2023: 54, f. 3A-B,4A-E, 5A-B, 6A-F, 7A-E, 8A-E, 9A-D, 10A-E (Dmf) Original description
Ummidia tibacuy Godwin & Bond, 2021 | | Colombia urn:lsid:nmbe.ch:spidersp:052984
Ummidia tibacuy Godwin & Bond, 2021: 150, f. 67A-B (Df) Original description
Ummidia timcotai Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052966
Ummidia timcotai Godwin & Bond, 2021: 86, f. 32A-B, 33A-E (Dmf) Original description
Ummidia tunapuna Godwin & Bond, 2021 | | Trinidad and Tobago urn:lsid:nmbe.ch:spidersp:052986
Ummidia tunapuna Godwin & Bond, 2021: 157, f. f71A-B (Df) Original description
Ummidia varablanca Godwin & Bond, 2021 | | Costa Rica urn:lsid:nmbe.ch:spidersp:052980
Ummidia varablanca Godwin & Bond, 2021: 137, f. 59A-B, 60A-E (Dmf) Original description
Ummidia waunekaae Godwin & Bond, 2021 | | USA urn:lsid:nmbe.ch:spidersp:052964
Ummidia waunekaae Godwin & Bond, 2021: 78, f. 29A-E (Dm) Original description
Ummidia yojoa Godwin & Bond, 2021 | | Honduras urn:lsid:nmbe.ch:spidersp:052977
Ummidia yojoa Godwin & Bond, 2021: 125, f. 53A-E (Dm) Original description
Ummidia zebrina (F. O. Pickard-Cambridge, 1897) | | Mexico, Guatemala urn:lsid:nmbe.ch:spidersp:000627
Pachylomerus zebrinus F. O. Pickard-Cambridge, 1897a: 9, pl. 1, f. 8 (Dm) Original description
Ummidia zebrina Ibarra-Núñez, Maya-Morales & Chamé-Vázquez, 2011: 1186, f. 5e (m)
Ummidia zebrina Godwin & Bond, 2021: 114, f. 47A-E (m)
Ummidia zilchi Kraus, 1955 | | Mexico, El Salvador, Belize urn:lsid:nmbe.ch:spidersp:000628
Ummidia zilchi Kraus, 1955b: 7, f. 1-2 (Dm; f is misidentified Ummidia hondurena per Godwin & Bond, 2021: 119) Original description
Ummidia zilchi Bücherl, 1957: 382, f. 1-1a (m)
Ummidia zilchi Godwin & Bond, 2021: 119, f. 50A-E (m)

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