World Spider Catalog

Gnaphosidae Banks, 1892

Family | ACCEPTED

N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.

Gen. Marinarozelotes Ponomarev, 2020 urn:lsid:nmbe.ch:spidergen:04642

N.B.: the genus was firstly named Simonizelotes by Marinaro, 1967: 702, but as there was no type species designated this is a nomen nudum; nevertheless Platnick & Murphy, 1984: 3 synonymzed it with Trachyzelotes; Levy, 1998c: 102 tried to validate the name, but failed as he clearly attributed the authorship to Marinaro; finally Ponomarev (in Ponomarev & Shmatko, 2020: 128) made it valid, but with the new name Marinarozelotes; regarded as subgenus of Zelotes Gistel, 1848 by Wunderlich, 2023: 62.

In synonymy:

Marinarozelotes acanthognatha (Purcell, 1907, T from Camillina) = Marinarozelotes jaxartensis Kroneberg, 1875 (Platnick & Murphy, 1984: 10, sub Trachyzelotes)

Marinarozelotes agilis (Bryant, 1936, T from Drassyllus) = Marinarozelotes lyonneti Audouin, 1826 (Platnick & Murphy, 1984: 6, sub Trachyzelotes)

Marinarozelotes barbaranus (Chamberlin, 1922, T from Nodocion) = Marinarozelotes jaxartensis Kroneberg, 1875 (Ubick & Roth, 1973a: 3, sub Drassyllus peninsulanus)

Marinarozelotes brasilianus (Keyserling, 1891, T from Zelotes) = Marinarozelotes lyonneti Audouin, 1826 (Platnick & Murphy, 1984: 6, sub Trachyzelotes)

Marinarozelotes cavaleriei (Schenkel, 1963, T from Zelotes) = Marinarozelotes jaxartensis Kroneberg, 1875 (Platnick & Murphy, 1984: 11, sub Trachyzelotes)

Marinarozelotes chamberlini (Roewer, 1951, T from Nodocion) = Marinarozelotes lyonneti Audouin, 1826 (Ubick & Roth, 1973a: 2, sub Drassyllus agilis)

Marinarozelotes chohanius (Patel & Patel, 1975, T from Scotophaeus) = Marinarozelotes jaxartensis Kroneberg, 1875 (Platnick & Murphy, 1984: 11, sub Trachyzelotes)

Marinarozelotes costatus (Denis, 1952) = Marinarozelotes bardiae Caporiacco, 1928 (Levy, 1998c: 110, sub Trachyzelotes)

Marinarozelotes errans (Benoit, 1977, T from Zelotes) = Marinarozelotes lyonneti Audouin, 1826 (Platnick & Murphy, 1984: 7, sub Trachyzelotes)

Marinarozelotes foveolatus (Simon, 1880, T from Zelotes) = Marinarozelotes jaxartensis Kroneberg, 1875 (Platnick & Song, 1986: 18, sub Trachyzelotes)

Marinarozelotes indraprastha (Tikader & Gajbe, 1975, T from Drassodes) = Marinarozelotes jaxartensis Kroneberg, 1875 (Platnick & Murphy, 1984: 11, sub Trachyzelotes)

Marinarozelotes insipiens (Simon, 1886, T from Zelotes) = Marinarozelotes jaxartensis Kroneberg, 1875 (Platnick & Murphy, 1984: 10, sub Trachyzelotes)

Marinarozelotes iugans (Chamberlin, 1922, T from Nodocion) = Marinarozelotes jaxartensis Kroneberg, 1875 (Ubick & Roth, 1973a: 3, sub Drassyllus peninsulanus)

Marinarozelotes microbarbatus (Marinaro, 1968, T from Zelotes) = Marinarozelotes mutabilis Simon, 1878 (Platnick & Murphy, 1984: 18, sub Trachyzelotes)

Marinarozelotes peninsulanus (Banks, 1898, T from Drassyllus) = Marinarozelotes jaxartensis Kroneberg, 1875 (Platnick & Murphy, 1984: 10, sub Trachyzelotes)

Marinarozelotes philippsoni (Denis, 1956, T from Zelotes) = Marinarozelotes lyonneti Audouin, 1826 (Platnick & Murphy, 1984: 7, sub Trachyzelotes)

Marinarozelotes postrema (Tucker, 1923, T from Camillina) = Marinarozelotes jaxartensis Kroneberg, 1875 (Platnick, 1997: 97, sub Trachyzelotes)

Marinarozelotes rubicundulus (Simon, 1878, T from Zelotes) = Marinarozelotes fuscipes L. Koch, 1866 (Platnick & Murphy, 1984: 15, sub Trachyzelotes)

Marinarozelotes samoensis (Berland, 1934, T from Zelotes) = Marinarozelotes kulczynskii Bösenberg, 1902 (Platnick & Murphy, 1984: 13, sub Trachyzelotes)

Marinarozelotes semibadius (L. Koch, 1878, T from Zelotes) = Marinarozelotes jaxartensis Kroneberg, 1875 (Khasayeva & Huseynov, 2017: 129, sub Trachyzelotes)

Marinarozelotes solaster (Chamberlin, 1937, T from Nodocion) = Marinarozelotes lyonneti Audouin, 1826 (Ubick & Roth, 1973a: 2, sub Drassyllus agilis)

Marinarozelotes sorex (Denis, 1945, T from Zelotes) = Marinarozelotes jaxartensis Kroneberg, 1875 (Platnick & Murphy, 1984: 11, sub Trachyzelotes)

Marinarozelotes spinibarbis (Simon, 1897, T from Camillina) = Marinarozelotes jaxartensis Kroneberg, 1875 (Platnick & Murphy, 1984: 10, sub Trachyzelotes)

Marinarozelotes ursinus (O. Pickard-Cambridge, 1872, T from Zelotes) = Marinarozelotes lyonneti Audouin, 1826 (Platnick & Murphy, 1984: 6, sub Trachyzelotes)

Marinarozelotes zagistus (Ponomarev, 1981) = Marinarozelotes adriaticus Caporiacco, 1951 (Ponomarev & Tsvetkov, 2004: 94, sub Trachyzelotes)

Marinarozelotes achaemenes Zamani, Chatzaki, Esyunin & Marusik, 2021 | | Iran urn:lsid:nmbe.ch:spidersp:053232

Marinarozelotes achaemenes Zamani et al., 2021a: 91, f. 15A-F (Df) Original description

Marinarozelotes adriaticus (Caporiacco, 1951) | | Italy to China urn:lsid:nmbe.ch:spidersp:028023

Zelotes adriaticus Caporiacco, 1951a: 87, f. 8a-b (Dm) Original description

Zelotes zagistus Ponomarev, 1981: 62, f. 4-5 (Dm) [urn:lsid:nmbe.ch:spidersp:028038]

Trachyzelotes adriaticus Platnick & Murphy, 1984: 9, f. 11-14 (m, Df, T from Zelotes)

Trachyzelotes adriaticus Hu & Wu, 1989: 283, f. 228.1-2 (f)

Trachyzelotes adriaticus Song, Zhu & Chen, 1999: 455, f. 265E, O (mf)

Trachyzelotes adriaticus Tuneva & Esyunin, 2002: 223, f. 30-34 (mf)

Trachyzelotes adriaticus Chatzaki, Thaler & Mylonas, 2003: 54, f. 30-35, 38-39 (f, S of Trachyzelotes stubbsi, revalidated by Levy, 2009: 25, m see Marinarozelotes stubbsi)

Trachyzelotes adriaticus Song, Zhu & Zhang, 2004: 232, f. 135A-D (mf)

Trachyzelotes adriaticus Ponomarev & Tsvetkov, 2004: 94 (S of Zelotes zagistus)

Trachyzelotes adriaticus Chatzaki, 2010b: 45, f. 2-5, 13 (mf)

Marinarozelotes adriaticus Ponomarev & Shmatko, 2020a: 132, f. 3-6, 9-10, 28-29, 35-36, 57-58 (mf, T from Trachyzelotes; misidentified Marinarozelotes lyonneti per Chatzaki, in litt.)

Marinarozelotes adriaticus Seyyar, Türkeş & Demir, 2022b: 346, f. 2, 3A-B (f)

Marinarozelotes cf. adriaticus Lecigne, 2023: 11, f. 6A-C (f)

Zelotes (Marinarozelotes) adriaticus Wunderlich, 2023: 155, f. 155 (m)

Marinarozelotes ansimensis (Seo, 2002) | | Korea urn:lsid:nmbe.ch:spidersp:048112

Trachyzelotes ansimensis Seo, 2002b: 59, f. 5-8 (Df) Original description

Marinarozelotes ansimensis Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Marinarozelotes baiyuensis (Xu, 1991) | | China urn:lsid:nmbe.ch:spidersp:040934

Trachyzelotes baiyuensis Xu, 1991b: 39, f. 22-23 (Df) Original description

Trachyzelotes baiyuensis Song, Zhu & Zhang, 2004: 233, f. 136A-B (f)

Marinarozelotes baiyuensis Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Marinarozelotes barbatus (L. Koch, 1866) | | Mediterranean to Caucasus. Introduced to USA urn:lsid:nmbe.ch:spidersp:028024

Melanophora barbata L. Koch, 1866: 161, pl. 6, f. 101-103 (Dmf) Original description

Prosthesima barbata Simon, 1878a: 45

Prosthesima barbata Chyzer & Kulczyński, 1897: 208, pl. 7, f. 46, pl. 8, f. 23 (mf)

Zelotes barbatus Simon, 1914: 153, 213, f. 282-283 (mf)

Zelotes barbatus Denis, 1952f: 115, f. 1-3 (mf)

Zelotes barbatus Jézéquel, 1962a: 522, f. 3 (f)

Trachyzelotes barbatus Platnick & Murphy, 1984: 15, f. 27-30 (Tmf from Zelotes)

Zelotes barbatus Pesarini, 2000: 385, f. 18-19 (mf)

Trachyzelotes barbatus Chatzaki, Thaler & Mylonas, 2003: 53, f. 20-21, 26-27 (mf)

Trachyzelotes barbatus Kovblyuk & Tuneva, 2009: 157, f. 10-16 (mf)

Trachyzelotes barbatus Gaymard & Lecigne, 2018: 18, f. 13A-D (f)

Marinarozelotes barbatus Ponomarev & Shmatko, 2020a: 130, f. 12, 17, 23-24, 39-40, 48-49, 52, 60-62 (mf, T from Trachyzelotes)

Zelotes (Marinarozelotes) barbatus Wunderlich, 2023: 68, f. 156-156a (m)

Marinarozelotes bardiae (Caporiacco, 1928) | | Mediterranean urn:lsid:nmbe.ch:spidersp:028025

Zelotes bardiae Caporiacco, 1928b: 87, f. 4 (Dm) Original description

Zelotes costatus Denis, 1952f: 118, f. 4 (Df) [urn:lsid:nmbe.ch:spidersp:056923]

Trachyzelotes costatus Platnick & Murphy, 1984: 20, f. 43-46 (Dm, f, T from Zelotes)

Trachyzelotes bardiae Levy, 1998c: 110, f. 31-32 (mf, T from Zelotes, S of Trachyzelotes costatus)

Marinarozelotes bardiae Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Zelotes (Marinarozelotes) costatus Wunderlich, 2023: 68, f. 162-163 (m)

Marinarozelotes chybyndensis (Tuneva & Esyunin, 2002) | | Russia (Europe), Kazakhstan urn:lsid:nmbe.ch:spidersp:038081

Trachyzelotes chybyndensis Tuneva & Esyunin, 2002: 224, f. 35-38 (Dmf) Original description

Marinarozelotes chybyndensis Ponomarev & Shmatko, 2020a: 134 (T from Trachyzelotes)

Marinarozelotes cumensis (Ponomarev, 1979) | | Greece, Ukraine, Russia (Europe), Azerbaijan, Kazakhstan urn:lsid:nmbe.ch:spidersp:028170

Zelotes cumensis Ponomarev, 1979a: 923, f. 8 (Df) Original description

Trachyzelotes cumensis Ponomarev & Tsvetkov, 2004: 94, f. 11-12 (Dm, f, T from Zelotes)

Trachyzelotes cumensis Ponomarev, 2008b: 82, f. 4 (m)

Trachyzelotes cumensis Ponomarev et al., 2018a: 248, f. 10-11 (f)

Marinarozelotes cumensis Ponomarev & Shmatko, 2020a: 135, f. 13, 18, 41-42, 63 (mf, T from Trachyzelotes)

Marinarozelotes fuscipes (L. Koch, 1866) | | Mediterranean, Iran, Kazakhstan, Uzbekistan, China urn:lsid:nmbe.ch:spidersp:028026

Melanophora fuscipes L. Koch, 1866: 189, pl. 8, f. 127-129 (Dmf) Original description

Prosthesima fuscipes Simon, 1878a: 51

Prosthesima rubicundula Simon, 1878a: 89 (Dmf) [urn:lsid:nmbe.ch:spidersp:056933]

Zelotes rubicundulus Simon, 1914: 159, 166, 216, f. 311, 340 (mf)

Zelotes fuscipes Simon, 1914: 175, 216, f. 365 (f)

Zelotes rubicundulus Jézéquel, 1962a: 527, f. 11 (f)

Zelotes rubicundulus Miller, 1967: 272, pl. III, f. 19 (m)

Trachyzelotes fuscipes Platnick & Murphy, 1984: 15, f. 31-34 (mf, T from Zelotes, S of Zelotes rubicundulus)

Trachyzelotes fuscipes Hu & Wu, 1989: 284, f. 229.1-4 (mf)

Trachyzelotes fuscipes Song, Zhu & Chen, 1999: 455, f. 265F, P (mf)

Trachyzelotes fuscipes Song, Zhu & Zhang, 2004: 234, f. 137A-D (mf)

Trachyzelotes fuscipes Akpınar et al., 2011: 16377, f. 2b (f)

Trachyzelotes fuscipes Gaymard & Lecigne, 2018: 37, f. 17A-C (f)

Trachyzelotes fuscipes Zamani et al., 2020: 579, f. 3H (f)

Marinarozelotes fuscipes Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Zelotes (Marinarozelotes) fuscipes Wunderlich, 2023: 67, f. 164-165 (m)

Marinarozelotes glossus (Strand, 1915) | | Turkey, Israel urn:lsid:nmbe.ch:spidersp:028027

Prosthesima glossa Strand, 1915c: 140 (Dm) Original description

Trachyzelotes glossus Levy, 1998c: 109, f. 29-30 (Tm from Zelotes)

Marinarozelotes glossus Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Marinarozelotes holosericeus (Simon, 1878) | | Mediterranean urn:lsid:nmbe.ch:spidersp:028028

Prosthesima holosericea Simon, 1878a: 47, pl. 14, f. 10 (Dmf) Original description

Zelotes holosericeus Simon, 1914: 153, 213, f. 280-281 (mf)

Zelotes holosericeus Jézéquel, 1962a: 522, f. 2 (f)

Trachyzelotes holosericeus Platnick & Murphy, 1984: 17, f. 35-38 (Tmf from Zelotes)

Trachyzelotes holosericeus Lecigne, 2017: 33, f. 15K-L, 16A-C (mf)

Marinarozelotes holosericeus Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Zelotes (Marinarozelotes) holosericeus Wunderlich, 2023: 67, f. 157-158 (m, at figs. holosericaeus)

Marinarozelotes huberti (Platnick & Murphy, 1984) | | Algeria, Italy, Albania urn:lsid:nmbe.ch:spidersp:028029

Trachyzelotes huberti Platnick & Murphy, 1984: 21, f. 47-50 (Dmf) Original description

Trachyzelotes huberti IJland, van Helsdingen & Miller, 2012: 8, f. 8a-c (m)

Marinarozelotes huberti Boucherit et al., 2020: 68, f. 5a-f (mf)

Marinarozelotes huberti Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Marinarozelotes jaxartensis (Kroneberg, 1875) | | Northern Africa to Caucasus, Russia (Europe) to Central Asia, Iran. Introduced to Hawaii, USA, Mexico, Ascension Is., South Africa, India, China, Japan urn:lsid:nmbe.ch:spidersp:028030

Melanophora jaxartensis Kroneberg, 1875: 23, pl. 2, f. 1 (Df) Original description

Prosthesima semibadia L. Koch, 1878b: 42, pl. 1, f. 3 (Df) [urn:lsid:nmbe.ch:spidersp:028383]

Prosthesima foveolata Simon, 1880b: 117, pl. 3, f. 17 (Df) [urn:lsid:nmbe.ch:spidersp:056925]

Prosthesima insipiens Simon, 1886g: 382 (Dmf) [urn:lsid:nmbe.ch:spidersp:056927]

Echemus spinibarbis Simon, 1897h: 97 (Df) [urn:lsid:nmbe.ch:spidersp:056937]

Prosthesima peninsulana Banks, 1898b: 217, pl. 13, f. 16 (Df) [urn:lsid:nmbe.ch:spidersp:056930]

Prosthesima iaxartensis Kulczyński, 1901b: 323

Melanophora acanthognathus Purcell, 1907: 333, pl. 15, f. 66 (Df) [urn:lsid:nmbe.ch:spidersp:056916]

Zelotes peninsulanus Petrunkevitch, 1911: 150

Nodocion barbaranus Chamberlin, 1922: 154 (Df) [urn:lsid:nmbe.ch:spidersp:056918]

Nodocion iugans Chamberlin, 1922: 154 (Dm) [urn:lsid:nmbe.ch:spidersp:056928]

Camillina acanthognathus Tucker, 1923: 335

Camillina postrema Tucker, 1923: 344, pl. 9, f. 61 (Dm) [urn:lsid:nmbe.ch:spidersp:056932]

Nodocion barbaranus Chamberlin, 1936a: 12, f. 16 (f)

Zelotes sorex Denis, 1945b: 46, f. 12 (Df) [urn:lsid:nmbe.ch:spidersp:056936]

Zelotes cavaleriei Schenkel, 1963: 50, f. 25a-b (Df) [urn:lsid:nmbe.ch:spidersp:056920]

Drassyllus peninsulanus Ubick & Roth, 1973a: 3 (f, T from Zelotes, S of Nodocion barbaranus and N. iugans)

Drassodes indraprastha Tikader & Gajbe, 1975: 276, f. 11-14 (Df) [urn:lsid:nmbe.ch:spidersp:056926]

Scotophaeus chohanius Patel & Patel, 1975a: 35, f. 2a-e (Df) [urn:lsid:nmbe.ch:spidersp:056922]

Drassodes indraprastha Tikader, 1982b: 419, f. 283-287 (f)

Zelotes cavaleriei Zhang & Zhu, 1983: 166, f. 4-6 (mf)

Zelotes cavaleriei Hayashi, 1983a: 11, f. 11-15 (f)

Zelotes cavalierei Brignoli, 1983c: 578

Zelotes cavaleriei Hu, 1984: 282, f. 297.1-4 (mf)

Trachyzelotes jaxartensis Platnick & Murphy, 1984: 10, f. 19-22 (mf, T from Zelotes, S of Camillina acanthognatha, C. spinibarbis, Drassodes indraprastha, Drassyllus peninsulanus, Scotophaeus chohanius, Zelotes cavaleriei, Z. insipiens and Z. sorex)

Zelotes cavaleriei Guo, 1985: 145, f. 1-3 (mf)

Zelotes cavaleriei Zhu & Shi, 1985: 154, f. 137a-c (f)

Trachyzelotes jaxartensis Platnick & Song, 1986: 18, f. 71-74 (mf, S of Zelotes foveolatus)

Zelotes jaxartensis Yaginuma, 1986a: 192, f. 106.7 (f)

Trachyzelotes jaxartensis Paik, 1986d: 33, f. 44-52 (mf)

Trachyzelotes jaxartensis Song, 1987: 343, f. 297 (mf)

Zelotes cavaleriei Zhang, 1987: 186, f. 158.1-4 (mf)

Trachyzelotes jaxartensis Hu & Wu, 1989: 286, f. 228.3-4 (f)

Zelotes jaxartensis Chikuni, 1989b: 120, f. 11 (mf)

Trachyzelotes jaxartensis Chen & Zhang, 1991: 238, f. 248.1-4 (mf)

Trachyzelotes jaxartensis Xu, 1991b: 39, f. 24 (f)

Zelotes cavaleriei Zhao, 1993: 127, f. 61a-c (mf)

Zelotes cavaleriei Barrion & Litsinger, 1995: 184, f. 105a-b (f, in part; not m, f. 106a-g, which belongs to Setaphis)

Zelotes cavaleriei Zhao, 1995: 844, f. 397a-c (mf)

Trachyzelotes jaxartensis Platnick, 1997: 97 (S of Camillina postrema)

Trachyzelotes jaxartensis Levy, 1998c: 107, f. 25-28 (mf)

Trachyzelotes jaxartensis Song, Zhu & Chen, 1999: 455, f. 265G, Q (mf)

Trachyzelotes jaxartensis Song, Zhu & Chen, 2001: 357, f. 233A-D (mf)

Trachyzelotes jaxartensis Namkung, 2002: 484, f. 37.25a-b (mf)

Trachyzelotes jaxartensis Namkung, 2003: 487, f. 37.25a-b (mf)

Trachyzelotes jaxartensis Song, Zhu & Zhang, 2004: 236, f. 138A-D (mf)

Trachyzelotes jaxartensis Jung et al., 2005: 196, f. 38, 76, 97, 136-137 (mf)

Trachyzelotes jaxartensis Jocqué & Dippenaar-Schoeman, 2006: 128, f. 41i-k (mf)

Trachyzelotes jaxartensis Kamura, 2009: 493, f. 150-152 (mf)

Trachyzelotes jaxartensis Zhu & Zhang, 2011: 403, f. 287A-D (mf)

Trachyzelotes jaxartensis Yin et al., 2012: 1204, f. 642a-e (mf)

Trachyzelotes jaxartensis Kim & Lee, 2013: 133, f. 93A-E (mf)

Trachyzelotes jaxartensis Zamani et al., 2016: 107, f. 19-20, 25 (mf)

Trachyzelotes jaxartensis Khasayeva & Huseynov, 2017: 128, f. 9-12 (mf, S of Prosthesima semibadia = Zelotes semibadius)

Marinarozelotes jaxartensis Ponomarev & Shmatko, 2020a: 135, f. 7-8, 11, 30, 37-38, 50, 59 (mf, T from Trachyzelotes)

Trachyzelotes jaxartensis Suzuki, 2020b: 59, f. 1a-e (mf)

Marinarozelotes jaxartensis Zamani, Al-Yacoub & Najim, 2022: 144, f. 1C, 4C (f)

Marinarozelotes jaxartensis Zhang, Peng & Zhang, 2022: 242, f. 180A-G (mf)

Marinarozelotes kulczynskii (Bösenberg, 1902) | | North Macedonia, Bulgaria. Introduced to USA, Caribbean, Colombia, Brazil, Japan, Samoa urn:lsid:nmbe.ch:spidersp:028031

Prosthesima kulczynskii Bösenberg, 1902: 313, pl. 29, f. 463 (Df) Original description

Zelotes samoensis Berland, 1934b: 325, f. 7-8 (Df) [urn:lsid:nmbe.ch:spidersp:056934]

Trachyzelotes kulczynskii Platnick & Murphy, 1984: 13, f. 23-26 (Dm, f, T from Zelotes, S of Zelotes samoensis, removed from S of T. barbatus contra Braun, 1982: 377)

Zelotes kulczynski Grimm, 1985: 287 (nomen dubium)

Trachyzelotes kulczynskii Kamura, 1997: 79, f. 5-8 (mf)

Trachyzelotes kulczynskii Brescovit, Freitas & Vasconcelos, 2008: 329, f. 3-6 (mf)

Trachyzelotes kulczynskii Kamura, 2009: 493, f. 153-155 (mf)

Marinarozelotes kulczynskii Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Trachyzelotes kulczynskii Suzuki & Serita, 2021a: 234, f. 7-8 (f)

Marinarozelotes lyonneti (Audouin, 1826) | | Macaronesia, Mediterranean to Central Asia. Introduced to USA, Mexico, Peru, Brazil urn:lsid:nmbe.ch:spidersp:028032

Drassus lyonetii Audouin, 1826: 383, pl. 5, f. 6 (Dm) Original description

Melanophora ursina O. Pickard-Cambridge, 1872: 245, pl. 16, f. 25 (Df) [urn:lsid:nmbe.ch:spidersp:028037]

Drassus secretus Thorell, 1875c: 92 (Df)

Prosthesima lyonneti Simon, 1878a: 98

Prosthesima ursina Simon, 1878a: 99

Prosthesima oceanica Simon, 1883: 273, pl. 8, f. 6 (Df)

Drassus brasilianus Keyserling, 1891: 34, pl. 1, f. 13 (Df) [urn:lsid:nmbe.ch:spidersp:056919]

Prosthesima lyonneti Chyzer & Kulczyński, 1897: 208, pl. 7, f. 34, 45 (mf)

Latonigena brasiliana Mello-Leitão, 1918a: 72, f. 42 (Dm)

Nodocion agilis Bryant, 1936a: 93, pl. 3, f. 3 (Dm) [urn:lsid:nmbe.ch:spidersp:056917]

Nodocion zelotoides Chamberlin, 1936b: 6, f. 20 (Df; preoccupied by Worley, 1928, see Drassyllus lamprus)

Nodocion solaster Chamberlin, 1937c: 170 (replacement name for N. zelotoides) [urn:lsid:nmbe.ch:spidersp:056935]

Zelotes brasilianus Mello-Leitão, 1943a: 216, f. 42 (mf)

Nodocion chamberlini Roewer, 1951: 443 (superfluous replacement name for Nodocion zelotoides) [urn:lsid:nmbe.ch:spidersp:056921]

Zelotes philippsoni Denis, 1956b: 275, f. 1 (Df) [urn:lsid:nmbe.ch:spidersp:056931]

Zelotes lyonneti Denis, 1962a: 38, f. 26-28 (mf)

Drassyllus agilis Ubick & Roth, 1973a: 2 (T from Nodocion, S of Nodocion chamberlini and N. solaster)

Zelotes errans Benoit, 1977c: 55, f. 19a-f (Df) [urn:lsid:nmbe.ch:spidersp:056924]

Trachyzelotes lyonneti Platnick & Murphy, 1984: 6, f. 7-10 (mf, T from Zelotes, S of Drassyllus agilis, Zelotes brasilianus, Z. errans, Z. philippsoni and Z. ursinus)

Trachyzelotes lyonneti Levy, 1998c: 105, f. 19-22 (mf)

Trachyzelotes lyonneti Tuneva, 2001: 1521, f. 1-2 (f)

Trachyzelotes lyonneti Chatzaki, Thaler & Mylonas, 2003: 54, f. 24-25, 29 (mf)

Trachyzelotes lyonneti Senglet, 2004: 101, f. 42, 45-46 (mf)

Trachyzelotes lyonneti Kovblyuk, 2004b: 141, f. 5-9 (mf)

Trachyzelotes lyonneti Lecigne, 2011: 18, f. p. 22 (m)

Trachyzelotes lyonneti Jiménez, Bizuet-Flores & Corcuera-Martínez del Río, 2012: 145, f. 1-3 (mf)

Marinarozelotes lyonneti Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Marinarozelotes lyonneti Seyyar, Türkeş & Demir, 2022b: 347, f. 3C-D (f)

Zelotes (Marinarozelotes) lyonneti Wunderlich, 2023: 67, f. 160-161 (m)

Marinarozelotes lyonneti Rehfeldt & Cassar, 2024: 23, f. 9d-e, 40e (f)

Marinarozelotes malkini (Platnick & Murphy, 1984) | | Romania, Albania, North Macedonia, Bulgaria, Greece, Ukraine, Russia (Europe, Caucasus), Turkey, Iran, Kazakhstan urn:lsid:nmbe.ch:spidersp:028033

Zelotes barbatus Miller, 1967: 271, pl. III, f. 17-18, pl. VI, f. 5 (mf, misidentified, per Kovblyuk & Tuneva, 2009: 159)

Trachyzelotes malkini Platnick & Murphy, 1984: 22, f. 51-54 (Dmf) Original description

Trachyzelotes malkini Chatzaki, Thaler & Mylonas, 2003: 53, f. 22-23, 28 (mf)

Trachyzelotes malkini Kovblyuk, 2004b: 143, f. 10-14 (mf)

Trachyzelotes malkini Kovblyuk & Tuneva, 2009: 159, f. 17-21 (mf)

Trachyzelotes malkini Ponomarev et al., 2016: 10, f. 9 (m)

Trachyzelotes malkini Türkeş, 2019b: 52, f. 1, 2A-B, 3-4 (mf)

Marinarozelotes malkini Ponomarev & Shmatko, 2020a: 135, f. 14-15, 19, 25-26, 43-44, 64-65 (mf, T from Trachyzelotes)

Marinarozelotes manytchensis (Ponomarev & Tsvetkov, 2006) | | Ukraine, Russia (Europe) urn:lsid:nmbe.ch:spidersp:041369

Trachyzelotes sp. Ponomarev & Tsvetkov, 2004: 95, f. 2.13-14 (mf; here per Esyunin & Tuneva, 2020: 113)

Trachyzelotes manytchensis Ponomarev & Tsvetkov, 2006: 11, f. 18-19 (Dmf) Original description

Zelotes manytchensis Senglet, 2011: 527, f. 59-64, 77 (mf, T from Trachyzelotes; misidentified per Ponomarev & Shmatko, 2020: 137)

Trachyzelotes manytchensis Ponomarev et al., 2016: 10, f. 5-8 (Tmf from Zelotes)

Trachyzelotes manytchensis Esyunin & Tuneva, 2020: 113, f. 22, 41-46 (mf)

Marinarozelotes manytchensis Ponomarev & Shmatko, 2020a: 137, f. 16, 20, 27, 45-46, 53, 66-67 (mf, T from Trachyzelotes)

Marinarozelotes miniglossus (Levy, 2009) | | Israel, Iran urn:lsid:nmbe.ch:spidersp:042300

Trachyzelotes miniglossus Levy, 2009: 27, f. 59-62 (Dmf) Original description

Trachyzelotes miniglossus Zamani & Mozaffarian, 2017: 11, f. 3 (f)

Marinarozelotes miniglossus Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Marinarozelotes minutus (Crespo, 2010) | | Portugal urn:lsid:nmbe.ch:spidersp:043618

Trachyzelotes minutus Crespo in Crespo & Mendes, 2010: 588, f. 1-9 (Dmf) Original description

Marinarozelotes minutus Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Zelotes (Marinarozelotes) minutus Wunderlich, 2023: 67, f. 159 (m)

Marinarozelotes mutabilis (Simon, 1878) | | Mediterranean, Romania urn:lsid:nmbe.ch:spidersp:028034

Prosthesima mutabilis Simon, 1878a: 46, pl. 14, f. 7 (Dmf) Original description

Zelotes microbarbatus Marinaro, 1968: 693, f. 10-11 (Dm) [urn:lsid:nmbe.ch:spidersp:056929]

Zelotes mutabilis Simon, 1914: 153, 213, f. 282, 284 (mf)

Trachyzelotes mutabilis Platnick & Murphy, 1984: 18, f. 39-42 (mf, T from Zelotes, S of Zelotes microbarbatus)

Zelotes mutabilis Jézéquel, 1962b: 597, f. 1 (f)

Zelotes (Trachyzelotes) cf. mutabilis Celano & Hansen, 1999: 83, f. 3-4 (f)

Trachyzelotes mutabilis Breitling, 2020: 343, f. 11J (m)

Marinarozelotes mutabilis Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Marinarozelotes ponticus Ponomarev, 2022 | | Russia (Europe) urn:lsid:nmbe.ch:spidersp:057129

Marinarozelotes ponticus Ponomarev in Ponomarev & Shmatko, 2022b: 287, f. 1-4 (Dm) Original description

Marinarozelotes ravidus (L. Koch, 1875) | | Ethiopia urn:lsid:nmbe.ch:spidersp:028036

Prosthesima ravida L. Koch, 1875b: 48, pl. 5, f. 4 (Df) Original description

Trachyzelotes ravidus Platnick & Murphy, 1984: 17, f. 17-18 (Tf from Zelotes)

Marinarozelotes ravidus Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)

Marinarozelotes stubbsi (Platnick & Murphy, 1984) | | Greece, Cyprus, Israel urn:lsid:nmbe.ch:spidersp:042299

Trachyzelotes stubbsi Platnick & Murphy, 1984: 9, f. 15-16 (Dm) Original description

Trachyzelotes stubbsi Levy, 1998c: 106, f. 23-24 (m)

Trachyzelotes adriaticus Chatzaki, Thaler & Mylonas, 2003: 54, f. 36-37 (m; misidentified per Levy, 2009: 25)

Trachyzelotes stubbsi Levy, 2009: 25, f. 57-58 (Df, removed from S of Trachyzelotes adriaticus, contra Chatzaki, Thaler & Mylonas, 2003: 54)

Trachyzelotes stubbsi Chatzaki, 2010b: 48, f. 6-12 (mf)

Marinarozelotes stubbsi Ponomarev & Shmatko, 2020a: 129 (T from Trachyzelotes)