Araneidae Clerck, 1757

Family |   ACCEPTED

N.B.: family name and authorship as per ICZN, 2009 Opinion 2224, after application by Kluge, 2007; Arkyinae L. Koch, 1872 (genera Arkys Walckenaer, 1837 and Demadiana Strand, 1929) elevated to Arkyidae by Dimitrov et al., 2017: 244; monophyly of Araneidae stated by Kallal et al., 2018: 138; Nephilinae Simon, 1894 [urn:lsid:nmbe.ch:spiderfam:0111] elevated to family and relimited by Kuntner, 2006: 24, re-synonymized with Araneidae by Dimitrov et al., 2017: 244; Nephilinae and Phonognathinae Simon, 1894 (Simon, 1894a: 746 as Phonognatheae), the valid name for Zygiellinae Simon, 1929 sensu Wunderlich, 2004f, elevated to family by Kuntner et al., 2019: 563, not accepted by Scharff et al., 2020 and Kallal et al., 2020, and confirmed as family by Kuntner et al., 2023: 969-970 along with the separation of Paraplectanoididae Kuntner, Coddington, Agnarsson & Bond, 2023; Nephilinae and Phonognathinae downgraded to subfamily and Paraplectanoididae considered a synonym of Araneidae by Hormiga et al., 2023: 803; family status of Paraplectanoididae, Nephilidae and Phonognathidae revalidated by Kuntner et al., 2025: 17, 23, but not accepted here for reasons of stability; Oarcinae (Gnolus and Oarces) transferred from the Mimetidae to the Araneidae by Dimitrov et al., 2012: Suppl. 1, p. 15, re-transfer to the Mimetidae by Eskov & Marusik, 2023: 340 not accepted here (by majority vote of the WSC boards); Eskov & Marusik, 2023: 241 and Eskov & Marusik, 2024: 52 consider Zygiellidae as the valid name instead of Phonognathidae and accept the family status of Nephilidae; the status of the Nephilinae and Phonognathinae (= Zygiellidae) as subfamilies is accepted by majority vote of the WSC boards; Eskov & Marusik, 2024: 52 list ten subfamilies and tribes as valid (besides Nephilidae and Zygiellidae): Araneinae, Argiopinae Simon, 1890a: 81 (sub Argiopidae), Cyrtarachninae Simon, 1895a: 876 (sub Cyrtarachneae), Сyrtophorinae Simon, 1895a: 876 (sub Cyrtophoreae), Gasteracanthinae O. Pickard-Cambridge, 1871e: 217 (sub Gasteracanthides), Micratheninae Simon, 1895a: 848 (sub Micratheneae), Guizygiellinae Zhu, Song & Zhang, 2003: 42, Caerostrini Simon, 1895a: 831 (sub Cerostreae), Hypognathini Simon, 1895a: 871 (sub Hypognatheae) and Poltyini Simon, 1895a: 888 (sub Poltyeae).

Gen. Pararaneus Caporiacco, 1940 urn:lsid:nmbe.ch:spidergen:01538
type Epeira striata Bösenberg & Lenz, 1895; gender masculine; syn. of Pararaneus cyrtoscapus (Pocock, 1898)
N.B.: Archer, 1951a: 21 implicitly synonymized this genus by transferring its type species to Eriophora Simon, 1864, but this change was not adopted by Grasshoff, 1968.

In synonymy:

Pararaneus asbenicus (Denis, 1955, T from Araneus) = Pararaneus spectator (Karsch, 1885) (Grasshoff, 1968: 92)

Pararaneus confusionis (Strand, 1906, T from Araneus) = Pararaneus spectator (Karsch, 1885) (Grasshoff, 1968: 91)

Pararaneus crassicaudex (Thorell, 1899, T from Araneus) = Pararaneus perforatus (Thorell, 1899) (Grasshoff, 1968: 89)

Pararaneus darolicola (Strand, 1906, T from Araneus) = Pararaneus cyrtoscapus (Pocock, 1898) (Grasshoff, 1968: 86)

Pararaneus hoplophallus (Pocock, 1899, T from Araneus) = Pararaneus cyrtoscapus (Pocock, 1898) (Grasshoff, 1968: 86)

Pararaneus lunulifera (Reimoser, 1937, T from Araneus) = Pararaneus spectator (Karsch, 1885) (Levy, 1998a: 344)

Pararaneus manicola (Strand, 1906, T from Araneus) = Pararaneus spectator (Karsch, 1885) (Grasshoff, 1968: 91)

Pararaneus paracymbifer (Strand, 1906) = Pararaneus cyrtoscapus (Pocock, 1898) (Grasshoff, 1968: 86)

Pararaneus randiae (Strand, 1917, T from Araneus) = Pararaneus spectator (Karsch, 1885) (Grasshoff, 1968: 92)

Pararaneus sanguipes (Thorell, 1899, T from Araneus) = Pararaneus cyrtoscapus (Pocock, 1898) (Grasshoff, 1986: 78)

Pararaneus streptoceros (Pocock, 1898) = Pararaneus spectator (Karsch, 1885) (Grasshoff, 1968: 91)

Pararaneus striatellus Roewer, 1942 = Pararaneus cyrtoscapus (Pocock, 1898) (Grasshoff, 1968: 86)

Pararaneus striatellus intermedius (Pavesi, 1897) = Pararaneus spectator (Karsch, 1885) (Grasshoff, 1968: 91)

Pararaneus striatellus similis (Bösenberg & Lenz, 1895) = Pararaneus spectator (Karsch, 1885) (Grasshoff, 1968: 91)

Pararaneus striatus (Bösenberg & Lenz, 1895) = Pararaneus cyrtoscapus (Pocock, 1898) (Grasshoff, 1968: 86)

Pararaneus suedicola (Simon, 1890, T from Araneus) = Pararaneus spectator (Karsch, 1885) (Grasshoff, 1968: 91)

Pararaneus suedicola pardalis (Simon, 1907, T from Araneus) = Pararaneus spectator (Karsch, 1885) (Grasshoff, 1968: 92)

Pararaneus tyloscapus (Pocock, 1900, T from Araneus) = Pararaneus perforatus (Thorell, 1899) (Grasshoff, 1968: 89)

Pararaneus cyrtoscapus (Pocock, 1898) | | Central, East, Southern Africa, Yemen (Socotra) urn:lsid:nmbe.ch:spidersp:016883 TYPE SPECIES
Epeira striata Bösenberg & Lenz, 1895: 43, pl. 2, f. 24-25 (Dmf; preoccupied by Doleschall, 1857, see Argiope aemula) [urn:lsid:nmbe.ch:spidersp:062183]
Araneus cyrtoscapus Pocock, 1898c: 206, pl. 8, f. 2 (Dmf) Original description
Aranea sanguipes Thorell, 1899: 48 (Df) [urn:lsid:nmbe.ch:spidersp:062175]
Araneus hoplophallus Pocock, 1899c: 40 (Dm) [urn:lsid:nmbe.ch:spidersp:062171]
Aranea darolicola Strand, 1906b: 623 (Dm) [urn:lsid:nmbe.ch:spidersp:062170]
Aranea paracymbifera Strand, 1906b: 624 (Dmf) [urn:lsid:nmbe.ch:spidersp:062174]
Aranea striata Tullgren, 1910: 163, pl. 4, f. 109 (f)
Araneus striatus Lessert, 1930: 639, f. 15-16 (mf)
Araneus sanguipes Lessert, 1930: 645, f. 20 (Dm, f, S of Araneus haematocnemis; rejected by Grasshoff, 1986: 78, see Neoscona penicillipes)
Pararaneus striatus Caporiacco, 1940b: 299
Pararaneus paracymbifer Caporiacco, 1941b: 90 (T from Araneus)
Pararaneus striatellus Roewer, 1942a: 869 (replacement name for Epeira striata Bösenberg & Lenz) [urn:lsid:nmbe.ch:spidersp:062177]
Pararaneus cyrtoscalus Caporiacco, 1947c: 160 (T from Araneus)
Pararaneus cyrtoscapus Grasshoff, 1968: 86, f. 48, 52-54, 60b, 63a-f, 67-73, 82 (mf, S of Araneus darolicola, A. hoplophallus, Pararaneus paracymbifer, P. striatus Bösenberg & Lenz and P. striatellus)
Pararaneus cyrtoscapus Grasshoff, 1986: 78 (S or Araneus sanguipes)
Pararaneus cyrtoscapus Dippenaar-Schoeman et al., 2022f: 31, p. 31: 7 f., p. 32: 6 f. (mf)
Pararaneus perforatus (Thorell, 1899) | | Central, East, Southern Africa urn:lsid:nmbe.ch:spidersp:016884
Aranea perforata Thorell, 1899: 46 (Dm) Original description
Aranea crassicaudex Thorell, 1899: 44 (Df) [urn:lsid:nmbe.ch:spidersp:062169]
Araneus tyloscapus Pocock, 1900g: 851, pl. 55, f. 3 (Df) [urn:lsid:nmbe.ch:spidersp:062182]
Pararaneus perforatus Grasshoff, 1968: 89, f. 47, 49-51, 60a, 62a-g, 67-73, 82-84 (mf, T from Araneus, S of Araneus crassicaudex and A. tyloscapus)
Pararaneus perforatus Dippenaar-Schoeman et al., 2022f: 33, p. 30: 1 f., p. 33: 5 f. (f)
Pararaneus pseudostriatus (Strand, 1908) | | Central, East Africa urn:lsid:nmbe.ch:spidersp:016885
Aranea striata pseudostriata Strand, 1908g: 380 (Dj) Original description
Pararaneus striatus substriatus Caporiacco, 1940b: 299 (lapsus for pseudostriatus, per Grasshoff, 1968: 93)
Araneus striatus pseudostriata Caporiacco, 1940c: 826 (Df)
Pararaneus pseudostriatus Grasshoff, 1968: 93, f. 58-59, 60d, 65a-d, 74-82 (Dm, f, elevated from subspecies)
Pararaneus spectator (Karsch, 1885) | | Africa, Middle East urn:lsid:nmbe.ch:spidersp:016886
Epeira spectator Karsch, 1885: 134, f. 3-4 (Df) Original description
Epeira suedicola Simon, 1890a: 103 (Df) [urn:lsid:nmbe.ch:spidersp:062180]
Epeira similis Bösenberg & Lenz, 1895: 44, pl. 2, f. 26 (Df; preoccupied by Blackwall, 1844, see Zygiella x-notata) [urn:lsid:nmbe.ch:spidersp:062179]
Epeira striata similis Pavesi, 1897b: 164
Epeira striata intermedia Pavesi, 1897b: 164 (Dmf) [urn:lsid:nmbe.ch:spidersp:062178]
Araneus streptoceros Pocock, 1898e: 436, pl. 13, f. 2 (Dmf) [urn:lsid:nmbe.ch:spidersp:062176]
Aranea confusionis Strand, 1906b: 619 (Df) [urn:lsid:nmbe.ch:spidersp:062168]
Aranea manicola Strand, 1906b: 623 (Dm) [urn:lsid:nmbe.ch:spidersp:062173]
Araneus suedicola pardalis Simon, 1907e: 6 (Df) [urn:lsid:nmbe.ch:spidersp:062181]
Aranea streptocera Strand, 1908g: 379
Aranea randiae Strand, 1917c: 71 (replacement name for Epeira similis Bösenberg & Lenz) [urn:lsid:nmbe.ch:spidersp:062184]
Aranea lunulifera Reimoser, 1937b: 20, f. 2 (Df) [urn:lsid:nmbe.ch:spidersp:062172]
Araneus striatus intermedius Caporiacco, 1940c: 826
Pararaneus striatellus intermedius Roewer, 1942a: 870
Pararaneus streptoceros Caporiacco, 1949a: 411, f. 53 (f, T from Araneus)
Araneus asbenicus Denis, 1955b: 134, f. 40 (Df) [urn:lsid:nmbe.ch:spidersp:062167]
Pararaneus spectator Grasshoff, 1968: 91, f. 55-57, 60c, 64a-d, 74-82 (mf, T from Araneus, S of Araneus asbenicus, A. confusionis, A. manicola, A. randiae, A. suedicola, A. suedicola pardalis, Pararaneus striatellus intermedius and P. striatellus similis)
Pararaneus spectator Levy, 1998a: 342, f. 127-134 (mf, S of Araneus lunulifer)
Pararaneus spectator Dippenaar-Schoeman et al., 2022f: 34, p. 30: 1 f., p. 34: 7 f. (f habitus)
Pararaneus uncivulva (Strand, 1907) | | Madagascar urn:lsid:nmbe.ch:spidersp:016887
Aranea uncivulva Strand, 1907a: 535 (Df) Original description
Aranea uncivulva Strand, 1907n: 634 (Df)
Aranea uncivulva Strand, 1908f: 469, f. K (f)
Pararaneus uncivulva Grasshoff, 1968: 94, f. 66a-d, 74-82 (f, T from Araneus)

Sponsors