World Spider Catalog

Thomisidae Sundevall, 1833

Family | ACCEPTED

N.B.: Includes Aphantochilidae as "a subfamily related to the Strophiinae" (Ono, 1988c: 217-218, after Homann, 1975: 184; see also Benjamin, 2011: 8 and Teixeira, Campos & Lise, 2014: 73). Separation of Borboropactidae Wunderlich, 2004 (Wunderlich, 2004m: 1738, Murphy & Roberts, 2015: viii) not sufficiently justified and is not accepted here. Lehtinen, 2016 splits the family into five families, i.e. Thomisidae s. str., Borboropactidae Wunderlich, 2004, Stephanopidae O. Pickard-Cambridge, 1871, Stiphropidae Simon, 1886 [sub Stiphropodidae; for nomenclatural author and year see Simon, 1886b: 170] and Bomidae Simon, 1886 [for nomenclatural author and year see Simon, 1886b: 173]; because of lack of a phylogenetic analysis and partly unclear limitations this splitting is not accepted; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 518, f. 17-18.

Gen. Psammitis Menge, 1876 urn:lsid:nmbe.ch:spidergen:04530

type Thomisus sabulosus Hahn, 1832; gender feminine

N.B.: type species fixed by F. O. Pickard-Cambridge, 1903b: 43; separated from Xysticus C. L. Koch, 1835 as an own genus by Ono, 1978b: 285, Wunderlich, 1987a: 253, Lehtinen, 2002: 320 and Breitling, 2019a: 203; formerly considered a junior synonym of Xysticus C. L. Koch, 1835 (Ono, 1988c: 78); Wunderlich, 1987a: 253 placed the subgenus Proxysticus Dalmas, 1922 (treated as a genus by some authors, e.g. Jézéquel, 1964c) as a junior synonym of Psammitis; Wunderlich, 1992a: 495 resurrected Proxysticus but this was not followed by Wunderlich, 1995F (see also Lehtinen, 2002: 320, 322 and Jantscher, 2002: 331); Breitling, 2019a: 202, 203 confirmed the synonymy of Proxysticus with Bassaniodes Pocock, 1903 (considered by Lehtinen, 2002: 322).

In synonymy:

Psammitis calcarata (Westring, 1861) = Psammitis sabulosa (Hahn, 1832) (Tullgren, 1944: 101, sub Xysticus)

Psammitis cambridgei (Blackwall, 1858) = Psammitis sabulosa (Hahn, 1832) (O. Pickard-Cambridge, 1900b: 60, sub Xysticus)

Psammitis defecta (O. Pickard-Cambridge, 1873) = Psammitis ninnii (Thorell, 1872) (Simon, 1875a: 180, sub Xysticus)

Psammitis embriki (Kolosváry, 1935) = Psammitis marmorata (Thorell, 1875) (Bosmans et al., 2013: 9, sub Xysticus)

Psammitis furcillifer (Schenkel, 1936) = Psammitis xysticiformis (Caporiacco, 1935) (Marusik & Logunov, 1995a: 161, sub Xysticus)

Psammitis gloveri (Blackwall, 1868, T from Thomisus) = Psammitis sabulosa (Hahn, 1832) (O. Pickard-Cambridge, 1900b: 60, sub Xysticus)

Psammitis gymnocephalus (Strand, 1915) = Psammitis marmorata (Thorell, 1875) (Bosmans et al., 2013: 9, sub Xysticus)

Psammitis jezequeli (Karol, 1966) = Psammitis marmorata (Thorell, 1875) (Assi, 1986: 45, sub Xysticus gymnocephalus)

Psammitis johannislupi (Denis, 1952) = Psammitis bonneti (Denis, 1938) (Heimer & Nentwig, 1991: 482, sub Xysticus johannislupi)

Psammitis jucunda (Simon, 1873) = Psammitis ninnii (Thorell, 1872) (Simon, 1875a: 180, sub Xysticus)

Psammitis knowltoni (Gertsch, 1939) = Psammitis nevadensis (Keyserling, 1880) (Dondale & Redner, 1975a: 134, sub Xysticus)

Psammitis perileuca (Simon, 1876) = Psammitis ninnii (Thorell, 1872) (Simon, 1932: 879, sub Xysticus)

Psammitis rufopicta (O. Pickard-Cambridge, 1873, T from Thomisus) = Psammitis sabulosa (Hahn, 1832) (Simon, 1875a: 189, sub Xysticus)

Psammitis setosa (Westring, 1851) = Psammitis sabulosa (Hahn, 1832) (Tullgren, 1944: 101, sub Xysticus)

Homonyms replaced:

Psammitis bimaculata (Emerton, 1894) -- see Psammitis rugosa Buckle & Redner, 1964 Buckle & Redner, 1964

Psammitis laticeps (Schenkel, 1963) -- see Psammitis gobiensis Marusik & Logunov, 2002

Nomen dubium:

Psammitis abscondita Menge, 1876: 450, pl. 75, f. 255 (mf, Poland) -- Prószyński & Staręga, 1971: 247

Psammitis abramovi (Marusik & Logunov, 1995) | | Turkey, Iran, Tajikistan urn:lsid:nmbe.ch:spidersp:031775

Xysticus abramovi Marusik & Logunov, 1995a: 146, f. 21-23 (Dm) Original description

Xysticus abramovi Demir, Aktaş & Topçu, 2010b: 253, f. 1A-B, 2A-C, 3A-B (m, Df)

Xysticus abramovi Kiany et al., 2017: 5, f. 10a-c (m)

Psammitis abramovi Breitling, 2019a: 203 (T from Xysticus)

Psammitis abuliensis Seropian & Mumladze, 2024 | | Georgia urn:lsid:nmbe.ch:spidersp:060596

Psammitis abuliensis Seropian & Mumladze, 2024: 121, f. 1-5 (Dm) Original description

Psammitis albida (Grese, 1909) | | Northern Europe, Russia (Europe, Siberia, Far East) urn:lsid:nmbe.ch:spidersp:031782

Xysticus albidus Grese, 1909: 328, pl. 7, f. 6-7 (Dmf) Original description

Xysticus albidus Kulczyński, 1916: 22, pl. 1, f. 36-37, pl. 2, f. 38-39 (mf)

Xysticus labradorensis Tullgren, 1944: 129, f. 48 (m; misidentified per Tullgren, 1946: 132)

Xysticus labradoriensis Palmgren, 1950: 20, f. 1, 4 (mf; misidentified sensu Tullgren, 1946: 132)

Xysticus labradorensis Utochkin, 1968: 20, f. 121-123 (mf; misidentified sensu Holm, 197 0: 207)

Xysticus albidus Holm, 1970: 207, f. 41-42 (f, removed from S of Xysticus labradorensis, rejecting Utochkin, 1968: 20 )

Xysticus albidus Ono, 1988c: 87, f. 75-77 (f)

Xysticus labradorensis Utochkin, 1989: 15 (misidentified sensu Holm, 197 0: 207 and Marusik, 1989a: 141)

Xysticus labradorensis Izmailova, 1989: 144, f. 138 (m; misidentified sensu Holm, 197 0: 207 and Marusik, 1989a: 141)

Xysticus albidus Marusik, 1989a: 141, pl. 1, f. 3-5 (m)

Xysticus albidus Almquist, 2007: 499, f. 422a-f (mf)

Psammitis albidus Breitling, 2019a: 203 (T from Xysticus)

Psammitis alpinistus (Ono, 1978) | | Nepal, China urn:lsid:nmbe.ch:spidersp:031787

Xysticus alpinistus Ono, 1978b: 279, f. 42-57, 74-77 (Df) Original description

Xysticus alpinistus Hu & Li, 1987b: 320, f. 43.1-3 (f)

Xysticus alpinistus Song & Zhu, 1997: 71, f. 41A-B (f)

Xysticus alpinistus Song, Zhu & Chen, 1999: 501, f. 284J (f)

Xysticus alpinistus Hu, 2001: 351, f. 212.1-3 (f)

Psammitis alpinistus Marusik, 2024: 262 (T from Xysticus)

Psammitis atevs (Ovtsharenko, 1979) | | Russia (Caucasus), Iran urn:lsid:nmbe.ch:spidersp:031801

Xysticus atevs Ovtsharenko, 1979: 40, f. 5-6, 20 (Dmf) Original description

Psammitis atevs Marusik, 2024: 262 (T from Xysticus)

Psammitis bonneti (Denis, 1938) | | France, Italy, Austria, Bulgaria, Russia (Urals to South Siberia), Kazakhstan urn:lsid:nmbe.ch:spidersp:031827

Xysticus ibex Simon, 1875a: 175 (f; misidentified per Jantscher, 2002b: 13)

Xysticus ibex Simon, 1932: 823, f. 1233 (f; misidentified sensu Jantscher, 2002b: 13)

Xysticus bonneti Denis, 1938b: 584, pl. 1, f. 6-8 (Dmf) Original description

Xysticus johannis-lupi Denis, 1952a: 59, f. 1 (Df) [urn:lsid:nmbe.ch:spidersp:056640]

Xysticus bonneti Thaler, 1981c: 122, f. 46-49 (mf)

Xysticus bonneti Marusik, 1989a: 143, pl. 2, f. 10 (m)

Xysticus johannislupi Heimer & Nentwig, 1991: 482, f. 1270 (mf, S of Xysticus bonneti; used younger name without explanation)

Xysticus johannislupi Esyunin, 1992a: 40, f. 6.1-7 (mf)

Psammitis bonneti Breitling, 2019a: 203 (T from Xysticus)

Psammitis bonneti Steinwandter et al., 2022: 143, f. 2A-F (m)

Psammitis courti (Marusik & Omelko, 2014) | | China urn:lsid:nmbe.ch:spidersp:047181

Xysticus sibiricus Schenkel, 1963: 236, f. 132 (m; misidentified per Marusik & Omelko, 2014a: 283)

Xysticus courti Marusik & Omelko, 2014a: 283, f. 30-35 (Dm) Original description

Psammitis courti Breitling, 2019a: 203 (T from Xysticus)

Psammitis courti Marusik, 2024: 264, f. 3A-C (m)

Psammitis daisetsuzana (Ono, 1988) | | Japan urn:lsid:nmbe.ch:spidersp:031862

Xysticus daisetsuzanus Ono, 1988c: 85, f. 70-74 (Dmf) Original description

Xysticus daisetsuzanus Ono, 2009a: 513, f. 146-149 (mf)

Psammitis daisetsuzanus Breitling, 2019a: 203 (T from Xysticus)

Psammitis deichmanni (Sørensen, 1898) | | Alaska, Canada, Greenland urn:lsid:nmbe.ch:spidersp:031864

Xysticus deichmanni Sørensen, 1898: 228 (Dmf) Original description

Xysticus labradorensis Gertsch, 1939b: 401, f. 248-249 (m; misidentified per Buckle & Redner, 196 4: 1139)

Xysticus deichmanni Brændegaard, 1940: 22, f. 20-21 (mf)

Xysticus deichmanni Holm, 1958a: 533, f. 9 (m, removed from S of Xysticus labradorensis, rejecting Gertsch, 1939b: 401)

Xysticus deichmanni Buckle & Redner, 1964: 1139, f. 9-13 (mf)

Xysticus deichmanni Turnbull, Dondale & Redner, 1965: 1263, f. 84-85, 161, 164 (mf)

Xysticus deichmanni Leech, 1966: 194, f. 40-41 (mf)

Xysticus deichmanni Dondale & Redner, 1978b: 239, f. 721-725 (mf)

Xysticus deichmanni Marusik, 1989a: 141, pl. 1, f. 6 (m)

Xysticus deichmanni Marusik, 2015: 701, f. 2E-F, 25A-B, 26A (mf)

Psammitis deichmanni Breitling, 2019a: 203 (T from Xysticus)

Psammitis demirsoyi (Demir, Topçu & Türkes, 2006) | | Turkey urn:lsid:nmbe.ch:spidersp:040760

Xysticus demirsoyi Demir, Topçu & Türkeş, 2006: 46, f. 1-8 (Dmf) Original description

Psammitis demirsoyi Breitling, 2019a: 203 (T from Xysticus)

Psammitis gobiensis (Marusik & Logunov, 2002) | | Russia (South Siberia), Mongolia, China urn:lsid:nmbe.ch:spidersp:038198

Xysticus laticeps Schenkel, 1963: 238, f. 133 (Df; preoccupied by Bryant, 1933, see Psammitis laticeps) [urn:lsid:nmbe.ch:spidersp:056563]

Xysticus laticeps Song & Haupt, 1996: 316, f. 5a-b (f; may refer to P. khamenkovae per Marusik, 2024: 264)

Xysticus laticeps Song & Zhu, 1997: 93, f. 60A-B (f)

Xysticus laticeps Song, Zhu & Chen, 1999: 503, f. 286E (f)

Xysticus laticeps Hu, 2001: 361, f. 222.1-3 (f; may refer to P. courti per Marusik, 2024: 264)

Xysticus gobiensis Marusik & Logunov, 2002b: 316, f. 1-5 (Dmf, removed Xysticus laticeps from S of X. sibiricus [rejecting Marusik, 1989a: 142], replacement name) Original description

Xysticus gobiensis Marusik & Kovblyuk, 2011: 253, f. 37.21, 27 (mf)

Xysticus gobiensis Marusik & Omelko, 2014a: 280, f. 28-29, 36-38 (mf)

Psammitis gobiensis Breitling, 2019a: 203 (T from Xysticus)

Psammitis gobiensis Marusik, 2024: 264, f. 3D-F, 4G-H (mf)

Psammitis khamenkovae Marusik, 2024 | | Kazakhstan urn:lsid:nmbe.ch:spidersp:060667

Psammitis khamenkovae Marusik, 2024: 262, f. 1A-E, 2A-F, 3G, 4A-F (Dmf) Original description

Psammitis labradorensis (Keyserling, 1887) | | North America, Greenland urn:lsid:nmbe.ch:spidersp:031957

Xysticus labradorensis Keyserling, 1887b: 479, pl. 6, f. 39 (Df) Original description

Xysticus bimaculatus Emerton, 1919b: 6, pl. 2, f. 18-20 (mf; misidentified per Buckle & Redner, 1964: 1138)

Xysticus labradorensis Gertsch, 1939b: 401, f. 268 (f, S of Xysticus deichmanni [revalidated by Holm, 1958a: 533], m f. 248-249 see Psammitis deichmanni)

Xysticus labradorensis Holm, 1958a: 533, f. 10 (m)

Xysticus labradorensis Buckle & Redner, 1964: 1138, f. 1-6 (mf)

Xysticus labradorensis Turnbull, Dondale & Redner, 1965: 1261, f. 80, 83, 160, 163, 166 (mf)

Xysticus labradorensis Utochkin, 1968: 20 (S of Xysticus albidus [revalidated by Holm, 1970: 207], f. 121-123 see Psammitis albida)

Xysticus labradorensis Dondale & Redner, 1978b: 237, f. 715-720 (mf)

Xysticus labradorensis Marusik, 1989a: 141, pl. 1, f. 7 (m)

Xysticus labradorensis Paquin & Dupérré, 2003: 234, f. 2642-2645 (mf)

Psammitis labradorensis Breitling, 2019a: 203 (T from Xysticus)

Psammitis laticeps (Bryant, 1933) | | USA, Cuba urn:lsid:nmbe.ch:spidersp:031965

Xysticus laticeps Bryant, 1933a: 178, pl. 3, f. 25 (Dm) Original description

Xysticus laticeps Gertsch, 1934b: 13, f. 14 (m)

Xysticus laticeps Gertsch, 1939b: 370, f. 182-183, 191 (m, Df)

Xysticus laticeps Bryant, 1940: 415, f. 198 (mf)

Xysticus laticeps Gertsch, 1953: 434, f. 34-35 (mf)

Psammitis laticeps Breitling, 2019a: 203 (T from Xysticus)

Psammitis lindbergi (Roewer, 1962) | | Afghanistan urn:lsid:nmbe.ch:spidersp:031969

Xysticus lindbergi Roewer, 1962a: 20, f. 104-105 (Df) Original description

Xysticus lindbergi Marusik & Logunov, 1995a: 163, f. 63-64 (f)

Psammitis lindbergi Breitling, 2019a: 203 (T from Xysticus)

Psammitis marmorata (Thorell, 1875) | | Central Europe, Balkans, Greece, Ukraine, Russia (Europe), Turkey, Caucasus, Kazakhstan urn:lsid:nmbe.ch:spidersp:031983

Xysticus marmoratus Thorell, 1875b: 92 (Dmf) Original description

Xysticus marmoratus Thorell, 1875c: 135 (Dmf)

Xysticus marmoratus Chyzer & Kulczyński, 1891: 94, pl. 3, f. 34 (f)

Xysticus gymnocephalus Strand, 1915c: 148 (Df) [urn:lsid:nmbe.ch:spidersp:031914]

Xysticus embriki Kolosváry, 1935c: 37, f. 3-4 (Dm) [urn:lsid:nmbe.ch:spidersp:031878]

Xysticus embriki Kolosváry, 1935d: 285, f. a-b (m)

Xysticus sabulosus embriki Kolosváry, 1936a: 95, 112, f. 3-4, 7-10 (m, Df)

Xysticus embriki Kolosváry, 1936b: 46, f. II (m)

Xysticus embriki Šilhavý, 1938: 89, f. 1-3 (mf)

Xysticus jézéqueli Karol, 1966a: 1, f. 1-3 (Dm) [urn:lsid:nmbe.ch:spidersp:055473]

Xysticus marmoratus Utochkin, 1968: 19, f. 113-114 (m; probably misidentified per Marusik & Logunov, 1995a: 154)

Xysticus embriki Miller, 1971: 121, pl. XV, f. 2, pl. LXII, f. 19 (mf)

Xysticus gymnocephalus Levy, 1976: 32, f. 57-58 (f)

Xysticus jezequeli Levy, 1976: 33, f. 59-60 (m)

Xysticus gymnocephalus Levy, 1985b: 103, f. 162-163 (f)

Xysticus jezequeli Levy, 1985b: 104, f. 164-165 (m)

Xysticus gymnocephalus Assi, 1986: 44 (S of Xysticus jezequeli)

Xysticus embriki Marusik & Logunov, 1995a: 154, f. 40-41 (m)

Xysticus marmoratus Mcheidze, 1997: 163, f. 305-306 (m)

Xysticus embriki Hepner et al., 2011: 38, f. 5-10 (mf)

Xysticus marmoratus Bosmans et al., 2013: 9 (S of Xysticus embriki and X. gymnocephalus)

Xysticus marmoratus Bosmans & Gavalas, 2023: 85, f. 45A-D (mf)

Xysticus marmoratus Geci et al., 2023: 546, f. 25 (m)

Psammitis marmoratus Seropian & Mumladze, 2024: 123 (T from Xysticus)

Psammitis marmorata Ponomarev, Turbanov & Shmatko, 2025a: 423, 25 (m)

Psammitis minor (Charitonov, 1946) | | Kazakhstan, Iran, Central Asia urn:lsid:nmbe.ch:spidersp:031985

Xysticus minor Charitonov, 1946: 28, f. 47 (Dm; see also Charitonov, 1969: 120) Original description

Xysticus minor Utochkin, 1968: 19, f. 115-117 (m)

Xysticus minor Marusik & Logunov, 1990: 44, f. 32-36 (m, Df)

Psammitis minor Breitling, 2019a: 203 (T from Xysticus)

Psammitis minor Nekhaeva, Kim & Yeszhanov, 2024: 403, f. 7E-F (m)

Psammitis mugur (Marusik, 1990) | | Russia (South Siberia) urn:lsid:nmbe.ch:spidersp:031989

Xysticus mugur Marusik in Marusik & Chevrizov, 1990: 91, f. 7-8 (Dm) Original description

Xysticus mugur Logunov & Marusik, 1994b: 187, f. 6-7 (Df)

Xysticus mugur Marusik & Kovblyuk, 2011: 253, f. 37.20 (m)

Psammitis mugur Marusik, 2024: 262 (T from Xysticus)

Psammitis nenilini (Marusik, 1989) | | Russia (Middle and South Siberia), Mongolia, China urn:lsid:nmbe.ch:spidersp:031997

Xysticus nenilini Marusik, 1989a: 140, pl. 1, f. 1-2 (Dm) Original description

Xysticus nenilini Logunov & Marusik, 1994b: 188, f. 8-9 (Df)

Xysticus nenilini Liu, Guo & Zhang, 2015: 59, f. 2a-c (f)

Psammitis nenilini Breitling, 2019a: 203 (T from Xysticus)

Psammitis nepalhimalaica (Ono, 1978) | | Nepal urn:lsid:nmbe.ch:spidersp:031998

Xysticus nepalhimalaicus Ono, 1978b: 276, f. 36-38, 69 (Dm) Original description

Xysticus nepalhimalaicus Marusik & Logunov, 1990: 45, f. 37-38 (m)

Psammitis nepalhimalaicus Breitling, 2019a: 203 (T from Xysticus)

Psammitis nevadensis (Keyserling, 1880) | | USA urn:lsid:nmbe.ch:spidersp:031999

Oxyptila nevadensis Keyserling, 1880: 50, pl. 1, f. 25 (Df) Original description

Xysticus knowltoni Gertsch, 1939b: 399, f. 244-245 (Dm) [urn:lsid:nmbe.ch:spidersp:056641]

Xysticus knowltoni Gertsch, 1953: 452, f. 58 (Df)

Ozyptila nevadensis Gertsch, 1953: 467, f. 84 (mf)

Xysticus knowltoni Schick, 1965: 167, f. 247-249 (mf)

Xysticus nevadensis Dondale & Redner, 1975a: 134 (mf, T from Ozyptila, S of Xysticus knowltoni)

Psammitis nevadensis Breitling, 2019a: 203 (T from Xysticus)

Psammitis ninnii (Thorell, 1872) | | Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Central Asia urn:lsid:nmbe.ch:spidersp:032004

Xysticus ninnii Thorell, 1872a: 246 (Dm) Original description

Xysticus jucundus Simon, 1873c: 328, pl. 10, f. 2 (Dm; f see Xysticus cristatus) [urn:lsid:nmbe.ch:spidersp:065911]

Xysticus defectus O. Pickard-Cambridge, 1873g: 541, pl. 15, f. 9 (Dm) [urn:lsid:nmbe.ch:spidersp:065912]

Xysticus ninnii Simon, 1875a: 180 (Df, S of Xysticus defectus and X. jucundus)

Xysticus perileucus Simon, 1876e: 180 (Df) [urn:lsid:nmbe.ch:spidersp:065913]

Xysticus ninnii Chyzer & Kulczyński, 1891: 94, pl. 3, f. 35 (mf)

Xysticus ninnii Simon, 1932: 829, 833, 879, f. 1246, 1256 (mf, S of Xysticus perileucus)

Xysticus ninnii Crome, 1965: 424, f. 1-7, 22-23 (mf)

Xysticus ninnii Utochkin, 1968: 19, f. 107-109 (mf)

Xysticus ninnii Azheganova, 1968: 122, f. 266, 270 (mf)

Xysticus ninnii Wunderlich, 1969: 381, f. 1-4 (f)

Xysticus ninni Tystshenko, 1971: 122, f. 299 (f)

Xysticus ninnii Miller, 1971: 120, pl. XV, f. 8-9 (mf)

Xysticus ninnii Thaler, 1981c: 123, f. 54-56 (m)

Xysticus ninnii Izmailova, 1989: 146, f. 143 (f)

Xysticus ninnii Heimer & Nentwig, 1991: 482, F. 1273 (mf)

Xysticus ninnii Mcheidze, 1997: 162, f. 299-301 (mf)

Xysticus ninnii Roberts, 1998: 171, f. (mf)

Xysticus ninnii Pesarini, 2000: 389, f. 31-32 (mf)

Psammitis ninnii Breitling, 2019a: 203 (T from Xysticus)

Psammitis ninnii fusciventris (Crome, 1965) | | Eastern Europe to Mongolia urn:lsid:nmbe.ch:spidersp:032005

Xysticus ninnii fusciventris Crome, 1965: 428, f. 8-21, 24-31 (Dmf) Original description

Xysticus ninnii fusciventris Utochkin & Savelyeva, 1995: 67, f. 2A-E (mf)

Xysticus ninnii fusciventris Ono & Martens, 2005: 110, f. 5-12 (mf)

Psammitis ninnii fusciventris Breitling, 2019a: 203 (T from Xysticus)

Psammitis novokhatskyii (Fomichev, 2015) | | Russia (Altai) urn:lsid:nmbe.ch:spidersp:047560

Xysticus novokhatskyii Fomichev, 2015a: 92, f. 1-2, 6-9, 16-17, 20-23 (Dmf) Original description

Psammitis novokhatskyii Breitling, 2019a: 203 (T from Xysticus)

Psammitis parallela (Simon, 1873) | | Spain, France, Italy urn:lsid:nmbe.ch:spidersp:032020

Xysticus parallelus Simon, 1873c: 330, pl. 10, f. 6-7 (Dmf) Original description

Xysticus parallelus Simon, 1932: 829, 836, 879, f. 1247, 1262 (mf)

Xysticus parallelus Lissner, 2016c: 6, f. 3A-B, 7-8 (mf)

Psammitis parallela Marusik, 2024: 262 (T from Xysticus)

Psammitis potamon (Ono, 1978) | | Nepal urn:lsid:nmbe.ch:spidersp:032030

Xysticus potamon Ono, 1978b: 277, f. 33-35, 70 (Dm) Original description

Psammitis potamon Breitling, 2019a: 203 (T from Xysticus)

Psammitis rugosa (Buckle & Redner, 1964) | | Russia (Middle Siberia to Far East), Canada, USA urn:lsid:nmbe.ch:spidersp:032053

Xysticus bimaculatus Emerton, 1894: 416, pl. 4, f. 5 (Df; preoccupied by L. Koch, 1867, see Xysticus bimaculatus) [urn:lsid:nmbe.ch:spidersp:065914]

Xysticus rugosus Buckle & Redner, 1964: 1141, f. 7-8 (f, replacement name) Original description

Xysticus rugosus Turnbull, Dondale & Redner, 1965: 1264, f. 162, 165 (f)

Xysticus rugosus Leech, 1969a: 633, f. 1-2 (Dm)

Xysticus rugosus Dondale & Redner, 1978b: 236, f. 711-714 (mf)

Xysticus rugosus Marusik, 1989a: 142, pl. 2, f. 9 (m)

Xysticus rugosus Logunov & Marusik, 1994b: 191, f. 20-22 (mf)

Xysticus rugosus Marusik & Logunov, 1996: 134, f. 17-23 (mf)

Psammitis rugosus Breitling, 2019a: 203 (T from Xysticus)

Psammitis sabulosa (Hahn, 1832) | | Europe, North Africa, Turkey, Caucasus, Russia (Europe to Middle Siberia), Afghanistan urn:lsid:nmbe.ch:spidersp:032054 TYPE SPECIES

Thomisus sabulosus Hahn, 1832: 28, f. 24 (Dmf) Original description

Xysticus sabulosus C. L. Koch, 1845: 64, f. 999-1000 (mf)

Thomisus setosus Westring, 1851: 50 (Df) [urn:lsid:nmbe.ch:spidersp:065916]

Thomisus cambridgei Blackwall, 1858a: 426 (Df) [urn:lsid:nmbe.ch:spidersp:056638]

Thomisus calcaratus Westring, 1861: 420 (Dm) [urn:lsid:nmbe.ch:spidersp:065917]

Thomisus sabulosus Blackwall, 1861a: 72, f. 41 (mf)

Thomisus cinereus Blackwall, 1861a: 74, pl. 4, f. 43 (f; misidentified per O. Pickard-Cambridge, 1900b: 60)

Thomisus cambridgei Blackwall, 1861a: 81, pl. 4, f. 47 (f)

Thomisus gloveri Blackwall, 1868: 405 (Df) [urn:lsid:nmbe.ch:spidersp:065918]

Xysticus calcaratus Thorell, 1872a: 242

Xysticus sabulosus Thorell, 1872a: 249

Xysticus setosus Thorell, 1872a: 250

Xysticus cambridgei Thorell, 1873: 425

Thomisus rufopictus O. Pickard-Cambridge, 1873b: 436 (Df) [urn:lsid:nmbe.ch:spidersp:065919]

Xysticus sabulosus Simon, 1875a: 189, pl. 7, f. 13 (mf, S of Thomisus rufopictus)

Psammitis sabulosa Menge, 1876: 449, pl. 75, f. 254 (mf)

Xysticus cinereus O. Pickard-Cambridge, 1881a: 536

Xysticus impavidus Hansen, 1882: 68, pl. 4, f. 15 (mf)

Xysticus sabulosus Becker, 1882b: 184, pl. 18, f. 1 (mf)

Xysticus sabulosus Chyzer & Kulczyński, 1891: 93, pl. 3, f. 32 (mf)

Xysticus sabulosus O. Pickard-Cambridge, 1900b: 60 (S of Thomisus gloveri and Xysticus cambridgei)

Xysticus sabulosus Bösenberg, 1902: 348, pl. 32, f. 514 (mf)

Xysticus sabulosus Kulczyński, 1916: 23, pl. 2, f. 40, 43-44 (mf)

Xysticus sabulosus Simon, 1932: 830, 835, 879, f. 1248, 1261 (mf)

Xysticus sabulosus Kolosváry, 1936b: 46, f. I (m)

Xysticus calcaratus Tullgren, 1942: 226, 233

Xysticus setosus Tullgren, 1942: 226, 233

Xysticus sabulosus Tullgren, 1944: 101, f. 37c, pl. 13, f. 194-196 (mf, S of Xysticus calcaratus and X. setosus)

Xysticus sabulosus Palmgren, 1950: 18, f. 3-5 (mf)

Xysticus sabulosus Locket & Millidge, 1951: 185, f. 93A, C (mf, S of Xysticus cambridgei confirmed)

Xysticus sabulosus Roewer, 1962a: 17, f. 96-98 (mf)

Xysticus sabulosus Vilbaste, 1969: 70, f. 59, 60A-B (mf)

Xysticus sabulosus Tystshenko, 1971: 122, f. 289 (m)

Xysticus sabulosus Miller, 1971: 121, pl. XV, f. 3, 7 (mf)

Xysticus sabulosus Brændegaard, 1972: 88, f. 46-48 (mf)

Xysticus sabulosus Thaler, 1981c: 122, f. 52, 57 (m)

Xysticus sabulosus Roberts, 1985: 102, f. 40d (mf)

Xysticus sabulosus Heimer & Nentwig, 1991: 482, f. 1272 (mf)

Xysticus sabulosus Roberts, 1995: 162, f. (mf)

Xysticus sabulosus Mcheidze, 1997: 162, f. 302-304 (mf)

Xysticus sabulosus Roberts, 1998: 170, f. (mf)

Psammitis sabulosus Lehtinen, 2002: 319, f. 8 (m)

Xysticus sabulosus Jantscher, 2002a: 332, f. 5-6 (m)

Xysticus sabulosus Almquist, 2007: 517, f. 435a-g (mf)

Psammitis sabulosus Breitling, 2019a: 203 (T from Xysticus)

Psammitis sabulosus Lecigne, 2020: 53, pl. 2F (m)

Psammitis sabulosa occidentalis (Kulczyński, 1916) | | France urn:lsid:nmbe.ch:spidersp:032055

Xysticus sabulosus occidentalis Kulczyński, 1916: 24, pl. 2, f. 41-42, 45 (Dmf) Original description

Psammitis sabulosus occidentalis Breitling, 2019a: 203 (T from Xysticus)

Psammitis secedens (L. Koch, 1876) | | Alps (Austria, Italy), North Macedonia, Turkey urn:lsid:nmbe.ch:spidersp:032060

Xysticus secedens L. Koch, 1876b: 329 (Dm) Original description

Xysticus secedens Thaler, 1981c: 123, f. 50-51, 53, 58 (mf)

Xysticus secedens Heimer & Nentwig, 1991: 482, f. 1271 (mf)

Xysticus secedens Lazarov, 2004: 163, f. 6-7 (f)

Psammitis secedens Breitling, 2019a: 203 (T from Xysticus)

Psammitis secedens Coşar, Danışman & Karahasan, 2023: 1420, f. 2A-C, 3A-B, 4A-B (f)

Psammitis seserlig (Logunov & Marusik, 1994) | | Russia (South Siberia, Far East), Mongolia urn:lsid:nmbe.ch:spidersp:032062

Xysticus seserlig Logunov & Marusik, 1994b: 191, f. 10-11, 11A (Dm) Original description

Xysticus seserlig Marusik & Logunov, 2002b: 316, f. 6-10 (m, Df)

Psammitis seserlig Breitling, 2019a: 203 (T from Xysticus)

Psammitis setiger (O. Pickard-Cambridge, 1885) | | Pakistan urn:lsid:nmbe.ch:spidersp:032063

Xysticus setiger O. Pickard-Cambridge, 1885b: 67 (Df) Original description

Xysticus setiger Marusik, 1993b: 457, f. 2-3 (f)

Psammitis setiger Breitling, 2019a: 203 (T from Xysticus)

Psammitis sibirica (Kulczyński, 1908) | | Russia (Middle Siberia to Far East), China urn:lsid:nmbe.ch:spidersp:032066

Xysticus sibiricus Kulczyński, 1908b: 54, pl. 2, f. 66-67 (Dm) Original description

Xysticus sibiricus Utochkin, 1968: 20, f. 124-125 (m)

Xysticus sibiricus Izmailova, 1978b: 18, f. 1 (f)

Xysticus sibiricus Izmailova, 1989: 148, f. 145 (f)

Xysticus sibiricus Marusik, 1989a: 142, pl. 2, f. 5-8 (m, S of Xysticus laticeps [misspelled leviceps]; revalidated by Marusik & Logunov, 2002b: 316, see Psammitis gobiensis)

Xysticus sibiricus Ono, Marusik & Logunov, 1990: 8, f. 3-4 (m)

Xysticus sibiricus Logunov & Marusik, 1994b: 191, f. 23-24 (mf)

Xysticus sibiricus Marusik & Logunov, 1996: 135, f. 24-26 (mf, S of Xysticus laticeps confirmed; revalidated by Marusik & Logunov, 2002b: 316, see Psammitis gobiensis)

Xysticus sibiricus Marusik & Omelko, 2014a: 285, f. 39 (m)

Psammitis sibiricus Breitling, 2019a: 203 (T from Xysticus)

Psammitis simplicipalpata (Ono, 1978) | | Nepal, Bhutan, China urn:lsid:nmbe.ch:spidersp:032072

Xysticus simplicipalpatus Ono, 1978b: 272, f. 8-32, 67-68 (Dmf) Original description

Psammitis simplicipalpatus Breitling, 2019a: 203 (T from Xysticus)

Psammitis simplicipalpatus Liu & Zhang, 2024d: 30, f. 1A-D, 2A-E (mf)

Psammitis torsiva (Tang & Song, 1988) | | China urn:lsid:nmbe.ch:spidersp:032095

Xysticus torsivus Tang & Song, 1988a: 253, f. 28-32 (Dmf) Original description

Xysticus torsivus Song & Zhu, 1997: 109, f. 74A-E (mf)

Xysticus torsivus Song, Zhu & Chen, 1999: 504, f. 287N, 288C (f)

Xysticus torsivus Hu, 2001: 365, f. 226.1-5 (mf)

Psammitis torsivus Breitling, 2019a: 203 (T from Xysticus)

Psammitis tyshchenkoi (Marusik & Logunov, 1995) | | Kazakhstan, Central Asia urn:lsid:nmbe.ch:spidersp:032105

Xysticus sabulosus Utochkin, 1968: 19, f. 110-112 (mf; misidentified per Marusik & Logunov, 1995a: 150)

Xysticus tyshchenkoi Marusik & Logunov, 1995a: 150, f. 34-39 (Dmf) Original description

Xysticus tyshchenkoi Fomichev, 2015a: 95, f. 3-4, 10-12, 18-19 (mf)

Psammitis tyshchenkoi Breitling, 2019a: 203 (T from Xysticus)

Psammitis wuae (Song & Zhu, 1995) | | China urn:lsid:nmbe.ch:spidersp:032119

Xysticus ninnii Hu & Wu, 1989: 348, f. 277.1-2 (m; misidentified per Song & Zhu, 1995: 120)

Xysticus wuae Song & Zhu, 1995: 120, f. 6A-B (Dm) Original description

Xysticus wuae Song & Zhu, 1997: 111, f. 76A-B (m)

Xysticus wuae Song, Zhu & Chen, 1999: 504, f. 287P (m)

Psammitis wuae Breitling, 2019a: 203 (T from Xysticus)

Psammitis xysticiformis (Caporiacco, 1935) | | Central Asia, China urn:lsid:nmbe.ch:spidersp:032122

Oxyptila xysticiformis Caporiacco, 1935b: 188, pl. 4, f. 8 (Dmf) Original description

Xysticus furcillifer Schenkel, 1936b: 140, f. 49 (Df) [urn:lsid:nmbe.ch:spidersp:056639]

Xysticus xysticiformis Marusik & Logunov, 1990: 50 (T from Ozyptila)

Xysticus xysticiformis Marusik & Logunov, 1995a: 161, f. 57-60 (mf, S of Xysticus furcillifer)

Xysticus xysticiformis Marusik, Fritzén & Song, 2007: 275, f. 21-24, 53-55 (mf)

Psammitis xysticiformis Breitling, 2019a: 203 (T from Xysticus)

Psammitis zonshteini (Marusik, 1989) | | Kyrgyzstan, Tajikistan urn:lsid:nmbe.ch:spidersp:032123

Xysticus zonshteini Marusik, 1989a: 141, pl. 2, f. 1-4 (Dm) Original description

Xysticus zonshteini Marusik & Logunov, 1995a: 162, f. 61-62 (Df)

Psammitis zonshteini Breitling, 2019a: 203 (T from Xysticus)