World Spider Catalog

Lycosidae Sundevall, 1833

Family | ACCEPTED

N.B.: according to Brignoli, 1983c: 432, "Guy (1966, 1969) has proposed many synonymies and has downgraded many genera to subgenus rank; as this author based his conclusions more on nomenclature than on the study of material (or even of the original descriptions)", Platnick (WSC 15.0) did not accept Guy's conclusions; the changes by Guy, 1966 are noted (except where more recent work has superseded them); Guy, 1969 is merely a literature compilation that reflects those changes and is not cited separately; several genera of Lycosidae have been found not to be monophyletic (Murphy et al., 2006). Piacentini & Ramírez, 2019: 233-237 confirm ten valid subfamilies of Lycosidae: Venoniinae Lehtinen & Hippa, 1979, Zoicinae Lehtinen & Hippa, 1979 (= Piratinae Zyuzin, 1993), Evippinae Zyuzin, 1985, Sosippinae Dondale, 1986 (= Hygrolycosinae He & Song, 1996), Artoriinae Framenau, 2007, Tricassinae Alderweireldt & Jocqué, 1993 (= Arctosinae He & Song, 1996), Hippasinae Simon, 1898, Allocosinae Dondale, 1986, Pardosinae Simon, 1898 (= Wadicosinae Zyuzin, 1985) and Lycosinae Sundevall, 1833; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 518, f. 17-18.

Gen. Hippasosa Roewer, 1960 urn:lsid:nmbe.ch:spidergen:01623

N.B.: considered a junior synonym of Ocyale Audouin, 1826: 149 [urn:lsid:nmbe.ch:spidergen:01641] (type O. atalanta Audouin, 1826) by Alderweireldt & Jocqué, 2005: 46, 63; synonymy rejected and Ocyale declared a nomen dubium (in Pisauridae) by Sherwood, 2022c: 583.

Transferred to other genera:

Hippasosa huachoi (Mello-Leitão, 1942) -- see Prolycosides

Hippasosa kalpiensis (Gajbe, 2004) -- see Arctosa

In synonymy:

Hippasosa modesta (Giebel, 1863, T from Lycosa) = Hippasosa pelliona Audouin, 1826 (Roewer, 1955c: 280, sub Ocyale)

Hippasosa natalica (Roewer, 1960, T from Hippasa) = Hippasosa guttata Karsch, 1878 (Alderweireldt & Jocqué, 2005: 65, sub Ocyale)

Hippasosa neatalanta (Alderweireldt, 1996) = Hippasosa pilosa Roewer, 1960 (Alderweireldt & Jocqué, 2005: 63, sub Ocyale)

Hippasosa spissa (Bösenberg & Lenz, 1895) = Hippasosa guttata Karsch, 1878 (Alderweireldt, 1996: 1356, sub Ocyale)

Nomina dubia:

Hippasosa conspersa (Karsch, 1879e: 831, j, Gabun, originally in Ocyale) -- Alderweireldt, 1996: 1364, sub Ocyale

Hippasosa ingenua (Karsch, 1879a: 354, j, Angola, originally in Lycosa) -- Alderweireldt, 1996: 1364, sub Ocyale

Hippasosa maculata (L. Koch, 1875b: 78, pl. 7, f. 5-5a, f, Ethiopia, originally in Trochosa) -- Alderweireldt, 1996: 1363, sub Ocyale

Nomen nudum:

Hippasosa spissides (Strand, 1908k: 344, Tanzania, originally in Ocyale, provisional name, taken as nomen dubium by Platnick) [urn:lsid:nmbe.ch:spidersp:039902] -- Alderweireldt, 1996: 1363, sub Ocyale

Hippasosa dewinterae (Alderweireldt, 1996) | | Malawi, Namibia, South Africa urn:lsid:nmbe.ch:spidersp:018465

Ocyale dewinterae Alderweireldt, 1996: 1359, f. 18-19, 22, 28-29 (Dmf) Original description

Ocyale dewinterae Jocque et al., 2017: 4, f. a-b, q (mf)

Ocyale dewinterae Dippenaar-Schoeman et al., 2021l: 15, 6 f. (mf)

Hippasosa dewinterae Sherwood, 2022c: 583 (T from Ocyale)

Hippasosa discrepans (Roewer, 1960) | | Ethiopia urn:lsid:nmbe.ch:spidersp:018466

Ocyale discrepans Roewer, 1960d: 809, f. 449a-b (Df) Original description

Ocyale discrepans Alderweireldt, 1996: 1362, f. 24 (f)

Hippasosa discrepans Sherwood, 2022c: 583 (T from Ocyale)

Hippasosa fera (Strand, 1908) | | Madagascar urn:lsid:nmbe.ch:spidersp:018467

Lycosa robusta Lenz, 1891: 171, pl. 1, f. 7 (Df; preoccupied by Simon, 1876, see Trochosa robusta)

Ocyale fera Strand, 1908a: 155 (replacement name) Original description

Ocyale fera Roewer, 1960d: 814, f. 452a-b (f)

Ocyale fera Alderweireldt, 1996: 1364 (does not belong to Ocyale, probably Lycosa s. lat.)

Hippasosa fera Sherwood, 2022c: 583 (T from Ocyale)

Hippasosa ghost (Jocque & Jocqué, 2017) | | Madagascar urn:lsid:nmbe.ch:spidersp:049637

Ocyale ghost Jocque & Jocqué in Jocque et al., 2017: 5, f. e-f, n, 3A-F, 4A-D, 5A-F, 6A-C, 7A-C (Dmf) Original description

Hippasosa ghost Sherwood, 2022c: 583 (T from Ocyale)

Hippasosa grandis (Alderweireldt, 1996) | | Togo, Gabun, Congo, Namibia urn:lsid:nmbe.ch:spidersp:018468

Ocyale ingenua Roewer, 1960d: 817, f. 453a-b (m; misidentified per Alderweireldt, 1996: 1359)

Ocyale grandis Alderweireldt, 1996: 1358, f. 16-17, 23, 30-31 (Dmf) Original description

Ocyale grandis Jocque et al., 2017: 4, f. c-d, i, p (mf)

Hippasosa grandis Sherwood, 2022c: 583 (T from Ocyale)

Hippasosa guttata (Karsch, 1878) | | Rwanda, Tanzania, Zambia, Namibia, Botswana, Mozambique, South Africa urn:lsid:nmbe.ch:spidersp:018469

Lycosa guttata Karsch, 1878a: 329, pl. 2, f. 1 (Dm) Original description

Trochosa guttata Pavesi, 1881: 552

Trochosa spissa Bösenberg & Lenz, 1895: 39, pl. 2, f. 18 (Dj) [urn:lsid:nmbe.ch:spidersp:037916]

Ocyale spissa Strand, 1908k: 342 (m, also using the provisional name O. spissides, see [urn:lsid:nmbe.ch:spidersp:039902])

Ocyale spissa Lessert, 1925b: 336, f. 7C (Df)

Ocyale spissa Lessert, 1927: 407, f. 1 (mf)

Ocyale spissa Roewer, 1960d: 810, f. 450a-c (mf)

Hippasa natalica Roewer, 1960d: 991, f. 548a-d (Dm) [urn:lsid:nmbe.ch:spidersp:017917]

Crocodilosa guttata Roewer, 1960d: 850, f. 471 (m, removed from S of Ocyale atalanta)

Ocyale guttata Alderweireldt, 1996: 1356, f. 4, 14-15, 21, 26-27 (mf, T from Crocodilosa, S of Ocyale spissa)

Ocyale guttata Alderweireldt & Jocqué, 2005: 65 (S of Hippasa natalica)

Ocyale guttata Jocque et al., 2017: 4, f. g-h, j-k (m)

Ocyale guttata Dippenaar-Schoeman et al., 2021l: 16, 4 f. (mf)

Hippasosa guttata Sherwood, 2022c: 583 (T from Ocyale)

Hippasosa guttata Dippenaar-Schoeman & Webb, 2023a: 20, f. 1-15 (mf)

Hippasosa kumari (Dyal, 1935) | | Pakistan urn:lsid:nmbe.ch:spidersp:018471

Ocyale kumari Dyal, 1935: 147, pl. 13, f. 47 (Df) Original description

Hippasosa kumari Sherwood, 2022c: 583 (T from Ocyale)

Hippasosa lanca (Karsch, 1879) | | Sri Lanka urn:lsid:nmbe.ch:spidersp:018472

Lycosa lanca Karsch, 1879d: 551 (Dm; N.B.: not = O. atalanta, per Roewer) Original description

Lycosa lanca Karsch, 1892: 297, pl. 12, f. 21 (m, Df)

Ocyale lanca Roewer, 1955c: 281

Hippasosa lanca Sherwood, 2022c: 583 (T from Ocyale)

Hippasosa pelliona (Audouin, 1826) | | Algeria, Egypt urn:lsid:nmbe.ch:spidersp:018474

Lycosa pelliona Audouin, 1826: 368, pl. 4, f. 5 (Df) Original description

Lycosa modesta Giebel, 1863: 314 (Df) [urn:lsid:nmbe.ch:spidersp:055203]

Leimonia pelliona Simon, 1864: 351

Ocyale pelliona Roewer, 1955c: 280 (S of Lycosa modesta)

Hippasosa pelliona Sherwood, 2022c: 583 (T from Ocyale)

Hippasosa pilosa Roewer, 1960 | | Turkey, Africa to Saudi Arabia, India, Sri Lanka, Myanmar urn:lsid:nmbe.ch:spidersp:017924

Dolomedes ocyale Walckenaer, 1837: 353 (merely refers to O. atalanta Audouin)

Ocyale ocyale Simon, 1864: 382

Ocyale atalanta Simon, 1886g: 358 (m, S of Trochosa lactea)

Ocyale atalanta Lessert, 1925b: 336, f. 7A-B (f; doubts the S of Trochosa lactea)

Ocyale atalanta Roewer, 1960d: 807, f. 448a-c (mf)

Hippasosa pilosa Roewer, 1960d: 1003, f. 555a-c (Dm) Original description

Ocyale atalanta Tikader & Malhotra, 1980: 364, f. 231-235 (mf)

Ocyale neatalanta Alderweireldt, 1996: 1353, f. 1-3, 5-13, 20, 25 (Dmf) [urn:lsid:nmbe.ch:spidersp:018473]

Ocyale pilosa Alderweireldt & Jocqué, 2005: 63 (S of Ocyale neatalanta)

Ocyale pilosa Siyam, Dunlop & El-Hennawy, 2015: 267, f. 13-15 (m)

Ocyale pilosa Jocque et al., 2017: 4, f. l-m, o, r (mf)

Hippasosa pilosa Sherwood, 2022c: 583 (T from Ocyale)

Hippasosa pilosa Sankaran & Caleb, 2023b: 541, f. 6A-B (f)

Hippasosa qiongzhongensis (Yin & Peng, 1997) | | China (Hainan) urn:lsid:nmbe.ch:spidersp:018475

Ocyale qiongzhongensis Yin & Peng, 1997: 6, f. 1-3 (Dm) Original description

Ocyale qiongzhongensis Yin et al., 1997c: 22, f. 8a-f (m)

Ocyale qiongzhongensis Song, Zhu & Chen, 1999: 329, f. 192O (m)

Ocyale qiongzhongensis Wang et al., 2021a: 42, f. 2A-B, 33A-H, 34A-F, 35A-D (m, Df)

Hippasosa qiongzhongensis Sherwood, 2022c: 583 (T from Ocyale)