World Spider Catalog

Zodariidae Thorell, 1881

Family | ACCEPTED

N.B.: the family name Zodariidae is protected by usage over the earlier name Cryptothelidae L. Koch, 1872a: 238 to keep nomenclatorial stability; separation of Cryptothelidae (Murphy & Roberts, 2015: viii) not sufficiently justified and is not accepted here; Jocqué, 1991a stated the subfamilies Cryptothelinae, Cydrelinae Simon, 1893, Cyriocteinae Jocqué, 1991, Lachesaninae Jocqué, 1991, Storeninae Simon, 1893, Storenomorphinae Simon, 1890 and Zodariinae; synonymy of the Cydrelinae with the Cryptothelinae considered by Ramírez et al., 2014: 392.

Gen. Zodariellum Andreeva & Tystshenko, 1968 urn:lsid:nmbe.ch:spidergen:03445

N.B.: considered by Marusik & Koponen, 2001a: 40 to be the earliest available name for Acanthinozodium Denis, 1952g: 58, because the type species of that genus was not designated by Denis, contra Jocqué, 1991a: 128 who designated A. cirrisulcatum Denis, 1952 as the type species and considered Acanthinozodium a senior synonym of Zodariellum Andreeva & Tystshenko, 1968: 688 (type Z. surprisum Tyschchenko, 1968) and contra Jocqué & Henrard, 2015a: 19, who considered Acanthinozodium Denis, 1966 (type A. spinulosum Denis, 1966) as valid and Zodariellum as a separate and monotypic genus; see also Levy, 1992: 85 for further comments on the type species and the validity of the genus.

Transferred to other genera:

Zodariellum bicoloripes (Denis, 1959) -- see Zodarion

Zodariellum buettikeri (Ono & Jocqué, 1986) -- see Zodarion

Zodariellum cirrisulcatum (Denis, 1952) -- see Acanthinozodium

Zodariellum cirrisulcatum longispina (Denis, 1952) -- see Acanthinozodium

Zodariellum inderensis Ponomarev, 2007 -- see Zodarion

Zodariellum lutipes (O. Pickard-Cambridge, 1872) -- see Zodarion

Zodariellum sahariense (Denis, 1959) -- see Acanthinozodium

Zodariellum sericeum (Denis, 1956) -- see Acanthinozodium

Zodariellum serraferum Lin & Li, 2009 -- see Tropizodium

Zodariellum spinulosum (Denis, 1966) -- see Acanthinozodium

Zodariellum subclavatum (Denis, 1952) -- see Acanthinozodium

Zodariellum sungar (Jocqué, 1991) -- see Zodarion

Zodariellum tibesti (Jocqué, 1991) -- see Acanthinozodium

Zodariellum zavattarii (Caporiacco, 1941) -- see Acanthinozodium

In synonymy:

Zodariellum apertum (B. S. Zhang & F. Zhang, 2019, T from Zodarion) = Zodariellum mongolicum Marusik & Koponen, 2001 (Zamani & Marusik, 2022b: 170)

Zodariellum martynovae (Andreeva & Tystshenko, 1968, T from Zodarion) = Zodariellum continentale Andreeva & Tystshenko, 1968 (Zamani & Marusik, 2022b: 169)

Zodariellum asiaticum (Tystshenko, 1970) | | Kazakhstan, Kyrgyzstan urn:lsid:nmbe.ch:spidersp:026036

Zodarion asiaticum Tystshenko in Marikovskii & Tystshenko, 1970: 199, f. 1 (Dmf) Original description

Zodariellum asiaticum Marusik & Koponen, 2001a: 41 (T from Zodarion)

Zodarion asiaticum Jocqué & Henrard, 2015a: 21 (T from Zodariellum)

Zodariellum asiaticum Zamani & Marusik, 2021b: 185 (T from Zodarion)

Zodariellum asiaticum Zamani & Marusik, 2022b: 164, f. 1A, 2A, 3A, 4A, 5A, 6A, 9A (m)

Zodariellum bactrianum (Kroneberg, 1875) | | Kazakhstan, Uzbekistan, Kyrgyzstan, Tadjikistan urn:lsid:nmbe.ch:spidersp:026040

Zodarion bactrianum Kroneberg, 1875: 12, pl. 1, f. 5 (Dmf) Original description

Zodarion bactrianum Andreeva & Tystshenko, 1968: 687, f. 4 (m)

Zodarion bactrianum Andreeva, 1976: 35, f. 42 (m)

Zodariellum bactrianum Zamani & Marusik, 2021b: 185, f. 28D-F (m, T from Zodarion)

Zodariellum bactrianum Zamani & Marusik, 2022b: 167, f. 1B, 2B, 2F, 3B, 4B, 5B, 6B, 7A-C, 8A-C, 9B-C (m)

Zodariellum bactrianum Fomichev & Zamani, 2024: 1057, f. 11-13 (m)

Zodariellum bekuzini (Nenilin, 1985) | | Uzbekistan urn:lsid:nmbe.ch:spidersp:026041

Zodarion bekuzini Nenilin, 1985b: 1584, f. 1-5 (Dmf) Original description

Zodariellum bekuzini Marusik & Koponen, 2001a: 41 (T from Zodarion)

Zodarion bekuzini Jocqué & Henrard, 2015a: 21 (T from Zodariellum)

Zodariellum bekuzini Zamani & Marusik, 2021b: 185 (T from Zodarion)

Zodariellum chaoyangense (C. D. Zhu & S. F. Zhu, 1983) | | China urn:lsid:nmbe.ch:spidersp:026047

Zodarium chaoyangensis Zhu & Zhu, 1983a: 137, f. a-h (Dmf) Original description

Zodarium chaoyangense Zhang, 1987: 60, f. 44.1-5 (mf)

Zodarium chaoyangense Zhu, 1988: 353, f. 1-4 (mf)

Zodarion chaoyangense Hu & Wu, 1989: 79, f. 58.1-6 (mf)

Zodarion chaoyangense Song, Zhu & Chen, 1999: 431, f. 257P-Q, 258A (mf)

Zodarion chaoyangense Song, Zhu & Chen, 2001: 327, f. 210A-D (mf)

Zodariellum chaoyangense Marusik & Koponen, 2001a: 41 (T from Zodarion)

Zodarion chaoyangense Jocqué & Henrard, 2015a: 21 (T from Zodariellum)

Zodarion chaoyangense Zhang & Zhang, 2019a: 81, f. 9A-D, 10A-E (mf)

Zodariellum chaoyangense Zamani & Marusik, 2021b: 185 (T from Zodarion)

Zodarion chaoyangense Zhang, Peng & Zhang, 2022: 209, f. 152A-G (mf)

Zodariellum continentale (Andreeva & Tystshenko, 1968) | | Uzbekistan, Tadjikistan urn:lsid:nmbe.ch:spidersp:026049

Zodarion continentalis Andreeva & Tystshenko, 1968: 687, f. 5 (Dm; N.B.: in heading, this species was referred to Zodariellum, apparently a lapsus, judging by legend and abstract) Original description

Zodarion martynovae Andreeva & Tystshenko, 1968: 684, f. 1-2 (Df) [urn:lsid:nmbe.ch:spidersp:026092]

Zodarion continentalis Andreeva, 1976: 35, f. 43 (m)

Zodarion martynovae Andreeva, 1976: 37, f. 45-46 (f)

Zodarion continentale Brignoli, 1983c: 212

Zodariellum continentale Marusik & Koponen, 2001a: 41 (T from Zodarion)

Zodarion continentale Jocqué & Henrard, 2015a: 21 (T from Zodariellum)

Zodariellum continentalis Zamani & Marusik, 2021b: 185 (T from Zodarion)

Zodariellum continentale Zamani & Marusik, 2022b: 169 (S of Zodarion martynovae)

Zodariellum dagestanum Ponomarev & Shmatko, 2023 | | Russia (Caucasus) urn:lsid:nmbe.ch:spidersp:059517

Zodariellum dagestanum Ponomarev in Ponomarev & Shmatko, 2023b: 216, f. 25, 27-28 (Dm) Original description

Zodariellum doroshkini Fomichev & Zamani, 2024 | | Tajikistan urn:lsid:nmbe.ch:spidersp:061324

Zodariellum doroshkini Fomichev & Zamani, 2024: 1055, f. 1-5 (Dm) Original description

Zodariellum furcum (Zhu, 1988) | | China urn:lsid:nmbe.ch:spidersp:026065

Zodarium furcum Zhu, 1988: 354, f. 5-9 (Dmf) Original description

Zodarion furcum Song, Zhu & Chen, 1999: 431, f. 257R-S, 258B (mf)

Zodarion furcum Song, Zhu & Chen, 2001: 328, f. 211A-E (mf)

Zodariellum furcum Marusik & Koponen, 2001a: 41 (T from Zodarion)

Zodarion furcum Jocqué & Henrard, 2015a: 21 (T from Zodariellum)

Zodarion furcum Zhang & Zhang, 2019a: 81, f. 11A-D, 12A-E (mf)

Zodariellum furcum Zamani & Marusik, 2021b: 185 (T from Zodarion)

Zodariellum hunanense (Yin, 2012) | | China urn:lsid:nmbe.ch:spidersp:046212

Zodarion hunanensis Yin in Yin et al., 2012: 1148, f. 611a-e (Df) Original description

Zodariellum hunanense Zamani & Marusik, 2022b: 170 (T from Zodarion)

Zodariellum kattakum Fomichev & Zamani, 2024 | | Uzbekistan urn:lsid:nmbe.ch:spidersp:061325

Zodariellum kattakum Fomichev & Zamani, 2024: 1056, f. 6-10 (Dm) Original description

Zodariellum mongolicum Marusik & Koponen, 2001 | | Mongolia, China urn:lsid:nmbe.ch:spidersp:037901

Zodariellum mongolicum Marusik & Koponen, 2001a: 45, f. 3-5, 8-9, 17-18, 26 (Dm) Original description

Zodarion mongolicum Jocqué & Henrard, 2015a: 21 (T from Zodariellum)

Zodarion apertum Zhang & Zhang, 2019a: 69, f. 1A-D, 2A-E, 3A-E (Dmf) [urn:lsid:nmbe.ch:spidersp:050828]

Zodariellum mongolicum Zamani & Marusik, 2021b: 185 (T from Zodarion)

Zodariellum mongolicum Zamani & Marusik, 2022b: 170 (S of Zodarion apertum)

Zodariellum mongolicum Fomichev & Zamani, 2024: 1057, f. 14-16 (m)

Zodariellum nenilini (Eskov, 1995) | | Kazakhstan, Mongolia, Russia (Urals)? urn:lsid:nmbe.ch:spidersp:025590

Zodarion nenilini Eskov in Eskov & Marusik, 1995: 62, f. 27-29 (Dmf) Original description

Acanthinozodium nenilini Marusik & Logunov, 1999: 253 (T from Zodarion)

Zodariellum nenilini Marusik & Koponen, 2001a: 42, f. 1, 6, 10-13, 19-24 (mf)

Zodariellum nenilini Esyunin, Tuneva & Farzalieva, 2007: 57, f. 21-23 (m)

Zodariellum nenilini Marusik & Kovblyuk, 2011: 277, f. 44.2. 4-5 (mf)

Zodarion nenilini Jocqué & Henrard, 2015a: 21 (T from Zodariellum)

Zodariellum nenilini Zamani & Marusik, 2021b: 185 (T from Zodarion)

Zodariellum nenilini Zamani & Marusik, 2022b: 172, f. 1D-E, 2C, 3C, 4C, 5C, 6C, 7D-F, 8D-F, 10A (mf, possibly the record from the south Urals belongs to Z. volgouralense)

Zodariellum planum (B. S. Zhang & F. Zhang, 2019) | | China urn:lsid:nmbe.ch:spidersp:050829

Zodarion planum Zhang & Zhang, 2019a: 73, f. 4A-E, 5A-C (Dm) Original description

Zodariellum planum Zamani & Marusik, 2022b: 174 (T from Zodarion)

Zodariellum proszynskii (Nenilin & Fet, 1985) | | Iran, Turkmenistan urn:lsid:nmbe.ch:spidersp:026108

Zodarion proszynskii Nenilin & Fet, 1985: 618, f. 1-4, 9 (Dmf) Original description

Zodariellum proszynskii Marusik & Koponen, 2001a: 41 (T from Zodarion)

Zodarion proszynskii Jocqué & Henrard, 2015a: 21 (T from Zodariellum)

Zodariellum proszynskii Zamani & Marusik, 2021b: 185, f. 31A-C (f, T from Zodarion)

Zodariellum proszynskii Zamani & Marusik, 2022b: 172, f. 10B-D (f)

Zodariellum schmidti Marusik & Koponen, 2001 | | Mongolia urn:lsid:nmbe.ch:spidersp:037902

Zodariellum schmidti Marusik & Koponen, 2001a: 44, f. 2, 7, 14-16, 25 (Dmf) Original description

Zodarion schmidti Jocqué & Henrard, 2015a: 21 (T from Zodariellum)

Zodariellum schmidti Zamani & Marusik, 2021b: 185 (T from Zodarion)

Zodariellum spasskyi (Charitonov, 1946) | | Uzbekistan, Kazakhstan? urn:lsid:nmbe.ch:spidersp:026124

Zodarion spasskyi Charitonov, 1946: 22, f. 11 (Df; see also Charitonov, 1969: 72) Original description

Zodariellum spasskyi Zamani & Marusik, 2022b: 175 (T from Zodarion, possibly the record from Kazakhstan belongs to another species)

Zodariellum spasskyi Esyunin, Zamani & Marusik, 2023: 704, f. 1-2, 4-5 (f)

Zodariellum surprisum (Andreeva & Tystshenko, 1968) | | Kazakhstan, Tadjikistan urn:lsid:nmbe.ch:spidersp:025596

Zodariellum surprisum Andreeva & Tystshenko, 1968: 688, f. 7-8 (Dm; N.B.: in heading, this name was referred only to Tystshenko, but two other references in the paper, and Andreeva, 1976 refer it to both authors) Original description

Zodariellum surprisum Andreeva, 1976: 41, f. 48-49 (m)

Zodariellum sytchevskajae (Nenilin & Fet, 1985) | | Turkmenistan urn:lsid:nmbe.ch:spidersp:026127

Zodarion sytchevskajae Nenilin & Fet, 1985: 619, f. 5-8, 10 (Dmf) Original description

Zodariellum sytchevskajae Marusik & Koponen, 2001a: 41 (T from Zodarion)

Zodarion sytchevskajae Jocqué & Henrard, 2015a: 21 (T from Zodariellum)

Zodariellum sytchevskajae Zamani & Marusik, 2021b: 185 (T from Zodarion)

Zodariellum tadzhikum (Andreeva & Tystshenko, 1968) | | Tajikistan urn:lsid:nmbe.ch:spidersp:026128

Zodarion tadzhikum Andreeva & Tystshenko, 1968: 686, f. 3 (Df) Original description

Zodarion tadzhikum Andreeva, 1976: 39, f. 47 (f)

Zodariellum tadzhikum Zamani & Marusik, 2022b: 174 (T from Zodarion)

Zodariellum testaceofasciatum (Spassky, 1941) | | Tajikistan urn:lsid:nmbe.ch:spidersp:026130

Zodarion testaceo-fasciatum Spassky, 1941: 14, f. 3 (Df) Original description

Zodarion testaceo-fasciatum Spassky & Luppova, 1945: 45 (Df)

Zodarion testaceofasciatum Brignoli, 1983c: 213

Zodariellum testaceofasciatum Zamani & Marusik, 2022b: 174 (T from Zodarion)

Zodariellum turanicum Zamani & Marusik, 2022 | | Uzbekistan, Turkmenistan urn:lsid:nmbe.ch:spidersp:055094

Zodariellum turanicum Zamani & Marusik, 2022b: 175, f. 1C, 2D, 3D, 4D, 5D, 6D, 9D (Dm) Original description

Zodariellum turanicum Fomichev & Zamani, 2024: 1058, f. 17-19 (m)

Zodariellum turkmenicum Zamani & Marusik, 2022 | | Turkmenistan urn:lsid:nmbe.ch:spidersp:055095

Zodariellum turkmenicum Zamani & Marusik, 2022b: 175, f. 1F, 2E, 3E, 4E, 5E, 6E (Dm) Original description

Zodariellum volgouralense Ponomarev, 2007 | | Russia (Europe), Kazakhstan urn:lsid:nmbe.ch:spidersp:041389

Zodariellum volgouralensis Ponomarev, 2007b: 95, f. 39-40 (Dmf) Original description

Zodariellum volgouralensis Ponomarev, Belosludtsev & Dvadnenko, 2008: 173, f. 13-16 (m)

Zodarion volgouralensis Jocqué & Henrard, 2015a: 21 (T from Zodariellum)

Zodariellum volgouralensis Zamani & Marusik, 2021b: 185 (T from Zodarion)

Zodariellum volgouralense Ponomarev & Shmatko, 2023b: 216, f. 26, 29 (m)

Zodariellum zebra (Charitonov, 1946) | | Uzbekistan urn:lsid:nmbe.ch:spidersp:026142

Zodarion zebra Charitonov, 1946: 21, f. 10 (Df; see also Charitonov, 1969: 71) Original description

Zodariellum zebra Zamani & Marusik, 2022b: 176 (T from Zodarion)

Zodariellum zebra Esyunin, Zamani & Marusik, 2023: 705, f. 3, 6-7 (f)