Gnaphosidae Banks, 1892

Family |   ACCEPTED
N.B.: the names Gnaphosae and Gnaphosi were used firstly at tribe level by Banks, 1892d: 92, 94 and thus Banks, 1892 is the valid authorship and year for the family, not Pocock, 1898c: 219; for the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612; most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae; Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5; Prodidomidae downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893), redefined by Rodrigues & Rheims, 2020b: 694 and elevated to familiy rank again by Azevedo et al., 2022: 12 (without Molycriinae); Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function; Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614); the subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra); Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020; Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7; Ammoxenidae (Ammoxenus, Austrammo, Barrowammo, Rastellus) are synonymized with Gnaphosidae by Azevedo et al., 2022: 12; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 520, f. 19.
Gen. Ammoxenus Simon, 1893 urn:lsid:nmbe.ch:spidergen:02327
N.B.: transferred from the Ammoxenidae to the Gnaphosidae by Azevedo et al., 2022: 12.

In synonymy:

Ammoxenus fallopius Lawrence, 1927 = Ammoxenus coccineus Simon, 1893 (Dippenaar & Meyer, 1980: 44)

Ammoxenus multosignatus Simon, 1910 = Ammoxenus psammodromus Simon, 1910 (Benoit, 1972b: 188)

Ammoxenus amphalodes Dippenaar & Meyer, 1980 | | South Africa urn:lsid:nmbe.ch:spidersp:026156
Ammoxenus amphalodes Dippenaar & Meyer, 1980: 45, f. 5, 11-12, 16-19 (Dmf) Original description
Ammoxenus amphalodes Dippenaar-Schoeman et al., 2020c: 7, p. 3: 2 f., p. 4: 2 f., p. 7: 6 f., p. 8: 6 f. (mf)
Ammoxenus amphalodes Dippenaar-Schoeman, Bird & Webb, 2024: 12, f. 1-14 (f)
Ammoxenus coccineus Simon, 1893 | | Zambia, Namibia, Botswana, South Africa urn:lsid:nmbe.ch:spidersp:026157
Ammoxenus coccineus Simon, 1893a: 455, f. 432-438 (Dmf) Original description
Ammoxenus fallopius Lawrence, 1927: 26, pl. 1, f. 18 (Dmf)
Ammoxenus coccineus Benoit, 1972b: 183, f. 1-7 (mf)
Ammoxenus fallopius Benoit, 1972b: 185, f. 8 (f)
Ammoxenus coccineus Dippenaar & Meyer, 1980: 44, f. 3, 9-10, 15 (mf, S)
Ammoxenus coccineus Ramírez, 2014: 267, f. 182A (f)
Ammoxenus coccineus Azevedo, Griswold & Santos, 2018b: 599, f. 9a, 15a (m)
Ammoxenus coccineus Dippenaar-Schoeman et al., 2020c: 9, 5 f. (mf)
Ammoxenus daedalus Dippenaar & Meyer, 1980 | | South Africa urn:lsid:nmbe.ch:spidersp:026158
Ammoxenus daedalus Dippenaar & Meyer, 1980: 47, f. 6, 13-14, 20-25 (Dmf) Original description
Ammoxenus daedalus Dippenaar-Schoeman et al., 2020c: 10, 3 f. (mf)
Ammoxenus kalaharicus Benoit, 1972 | | Botswana, South Africa urn:lsid:nmbe.ch:spidersp:026159
Ammoxenus kalaharicus Benoit, 1972b: 187, f. 9-11 (Df) Original description
Ammoxenus kalaharicus Dippenaar & Meyer, 1980: 42, f. 1 (f)
Ammoxenus kalaharicus Dippenaar-Schoeman et al., 2020c: 11, 3 f. (f)
Ammoxenus pentheri Simon, 1897 | | Botswana, South Africa urn:lsid:nmbe.ch:spidersp:026160
Ammoxenus pentheri Simon, 1897l: 222 (Df) Original description
Ammoxenus pentheri Benoit, 1972b: 187, f. 12-14 (f)
Ammoxenus pentheri Dippenaar & Meyer, 1980: 43, f. 2, 7-8 (f)
Ammoxenus pentheri Dippenaar-Schoeman et al., 2020c: 12, 5 f. (f)
Ammoxenus pentheri Frétey, 2023: 124 (year of description corrected)
Ammoxenus psammodromus Simon, 1910 | | Namibia, Botswana, South Africa urn:lsid:nmbe.ch:spidersp:026161
Ammoxenus psammodromus Simon, 1910b: 189 (Df) Original description
Ammoxenus multosignatus Simon, 1910b: 189 (Df)
Ammoxenus psammodromus Benoit, 1972b: 188, f. 15-18 (Dm, f, S of Ammoxenus multosignatus)
Ammoxenus psammodromus Dippenaar & Meyer, 1980: 45, f. 4 (f)
Ammoxenus psammodromus Dippenaar-Schoeman et al., 2020c: 13, 4 f. (mf)

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