World Spider Catalog

Pisauridae Simon, 1890

Family | ACCEPTED

N.B.: placement of some nursery-web spiders in a separate family Dolomedidae Simon, 1898a: 281, 301 (Lehtinen, 1967) [but Dolomedinae was first used by Simon, 1876a: 226, which makes it older than Pisauridae, overlooked by Bonnet, 1956: 1543] was rejected by Sierwald, 1990: 50, who also recognized a "Trechalea genus group [that] is not a close relative of the nursery-web Pisauridae" and whose members are now listed under Trechaleidae; considered a synonym of Bradystichidae Simon, 1884e: 298 (removed from the synonymy of Lycosidae by Lehtinen, 1967: 293) by Platnick & Forster, 1993: 3 and of Halidae Jocqué, 1994b: 281 by Jocqué & Dippenaar-Schoeman, 2006: 212; probably includes Senoculidae (Murphy & Roberts, 2015: ix); mentioning of "Dolomediens" by Simon, 1864: 342, but the French ending -iens was not accepted as taxon at familiar level by Bonnet and Platnick, and this is continued here; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 518, f. 17-18.

Gen. Pisaura Simon, 1886 urn:lsid:nmbe.ch:spidergen:01725

N.B.: new genus for the members of Ocyale Audouin, 1826 of the mirabilis-group, because the type species O. atalanta Audouin, 1826 is totally different (Simon, 1886g: 354).

Transferred to other genera:

Pisaura affinis (Lessert, 1916) -- see Cispius

Pisaura bobbiliensis Reddy & Patel, 1993 -- see Dendrolycosa

Pisaura camerunensis Roewer, 1955 -- see Afropisaura

Pisaura decorata Patel & Reddy, 1990 -- see Nilus

Pisaura dentifasciata (O. Pickard-Cambridge, 1885) -- see Perenethis

Pisaura ducis Strand, 1913 -- see Afropisaura

Pisaura faradjensis (Lessert, 1928) -- see Afropisaura

Pisaura gitae Tikader, 1970 -- see Dendrolycosa

Pisaura lizhii Zhang, 2000 -- see Dendrolycosa

Pisaura parangbusta Barrion & Litsinger, 1995 -- see Dendrolycosa

Pisaura putiana Barrion & Litsinger, 1995 -- see Dendrolycosa

Pisaura rectifasciata (O. Pickard-Cambridge, 1885) -- see Perenethis

Pisaura rothiformis Strand, 1908 -- see Afropisaura

Pisaura rothiformis orientalis Strand, 1913 -- see Afropisaura

Pisaura songi Zhang, 2000 -- see Dendrolycosa

Pisaura valida Simon, 1885 -- see Afropisaura

Pisaura zonaformis Wang, 1993 -- see Polyboea

In synonymy:

Pisaura anahita Kishida, 1914 = Pisaura anahitiformis Kishida, 1910 (Yaginuma, 1977c: 393)

Pisaura clarivittata Dönitz & Strand, 1906 = Pisaura lama Bösenberg & Strand, 1906 (Yaginuma, 1974a: 170)

Pisaura flavistriata Yaginuma, 1960 = Pisaura lama Bösenberg & Strand, 1906 (Yaginuma, 1974a: 170)

Pisaura lantana Wang, 1993 = Pisaura bicornis Zhang & Song, 1992 (Zhang, 2000a: 2)

Pisaura maderiana Kulczyński, 1895 = Pisaura quadrilineata Lucas, 1838 (Wunderlich, 1992a: 438)

Pisaura mirabilis albida (Franganillo, 1913) = Pisaura mirabilis Clerck, 1757 (Barrientos, 1978b: 18)

Pisaura mirabilis fusca (Franganillo, 1913) = Pisaura mirabilis Clerck, 1757 (Barrientos, 1978b: 18)

Pisaura strandi Kishida, 1914 = Pisaura lama Bösenberg & Strand, 1906 (Paik, 1969a: 44)

Nomen dubium:

Pisaura kishidai Saito, 1936b: 82, pl. 1, f. 3, 20 (m, China) -- Zhang, Zhu & Song, 2004d: 389, after Song, Zhu & Chen, 2001: 268

Pisaura acoreensis Wunderlich, 1992 | | Azores urn:lsid:nmbe.ch:spidersp:019748

Pisaura acoreensis Wunderlich, 1992a: 437, f. 656-659 (Dmf) Original description

Pisaura anahitiformis Kishida, 1910 | | Japan urn:lsid:nmbe.ch:spidersp:019749

Pisaura anahitiformis Kishida, 1910a: 5 (Df) Original description

Pisaura anahita Kishida, 1914b: 127, f. 4 (Df) [urn:lsid:nmbe.ch:spidersp:056382]

Pisaura anahitiformis Yaginuma, 1977c: 393 (S of Pisaura anahita)

Pisaura ancora Paik, 1969 | | Russia (South Siberia, Far East), China, Korea urn:lsid:nmbe.ch:spidersp:019750

Pisaura ancora Paik, 1969a: 49, f. 14, 18, 54-64, text-f. 2 (Dmf) Original description

Pisaura ancora Paik, 1978e: 377, f. 171.1-10 (mf)

Pisaura ancora Song & Zheng, 1982c: 155, f. 1 (f)

Pisaura ancora Hu, 1984: 261, f. 275.1-2, 276.1-2 (f, not male)

Pisaura ancora Guo, 1985: 134, f. 2-70.1-3 (mf)

Pisaura ancora Zhu & Shi, 1985: 147, f. 130a-c (f)

Pisaura ancora Song, 1987: 210, f. 169 (f)

Pisaura ancora Zhang, 1987: 167, f. 141.1-2 (mf)

Pisaura ancora Logunov, 1990: 37, f. 2-3 (mf)

Pisaura ancora Feng, 1990: 162, f. 137.1-3 (f)

Pisaura ancora Chen & Gao, 1990: 137, f. 171a-b (mf)

Pisaura ancora Chen & Zhang, 1991: 227, f. 235.1-2 (f)

Pisaura ancora Song, Zhu & Chen, 1999: 348, f. 204A (f)

Pisaura ancora Zhang, 2000a: 2, f. 1A-E (mf)

Pisaura ancora Hu, 2001: 217, f. 115.1-4 (mf)

Pisaura ancora Song, Zhu & Chen, 2001: 268, f. 168A-B (f)

Pisaura ancora Namkung, 2002: 350, f. 21.5a-b (mf)

Pisaura ancora Kim & Cho, 2002: 191, f. 397-402 (mf)

Pisaura ancora Namkung, 2003: 352, f. 21.5a-b (mf)

Pisaura ancora Zhang, Zhu & Song, 2004d: 391, f. 111-117, 220-225 (mf)

Pisaura ancora Zhu & Zhang, 2011: 296, f. 215A-G (mf)

Pisaura ancora Marusik & Kovblyuk, 2011: 219, f. 31.1-3 (m)

Pisaura ancora Yin et al., 2012: 888, f. 446a-f (mf)

Pisaura ancora Kim & Lee, 2013: 79, f. 52A-D (mf)

Pisaura ancora Dong, 2018: 72, f. 10-4.A-G, pl. 16 (mf)

Pisaura ancora Zhang, Peng & Zhang, 2022: 157, f. 116A-G (mf)

Pisaura bicornis Zhang & Song, 1992 | | China, Japan urn:lsid:nmbe.ch:spidersp:019751

Pisaura bicornis Zhang & Song, 1992: 17, f. 1A-D (Dmf) Original description

Pisaura lantanus Wang, 1993a: 158, f. 11 (Df)

Pisaura bicornis Song, Zhu & Chen, 1999: 348, f. 204B-C, I-J (mf)

Pisaura lantanus Song, Zhu & Chen, 1999: 348, f. 204E (f)

Pisaura bicornis Zhang, 2000a: 2, f. 2A-F (mf, S)

Pisaura bicornis Tanikawa, 2003: 41, f. 27-30 (f)

Pisaura bicornis Zhang, Zhu & Song, 2004d: 392, f. 118-123, 226-231 (mf)

Pisaura bicornis Tanikawa & Ono, 2009: 220, f. 33-34 (mf)

Pisaura bicornis Yin et al., 2012: 890, f. 447a-c (f)

Pisaura bicornis Suguro, 2013c: 14, f. 1, 3 (m)

Pisaura bicornis Zhang & Wang, 2017: 558, 7 f. (mf)

Pisaura consocia (O. Pickard-Cambridge, 1872) | | Cyprus, Turkey, Israel, Lebanon, Syria urn:lsid:nmbe.ch:spidersp:019753

Dolomedes consocius O. Pickard-Cambridge, 1872: 320 (Dmf) Original description

Pisaura consocia Simon, 1898a: 289

Ocyale consocia Kulczyński, 1911a: 48, pl. 2, f. 57-59 (mf)

Pisaura consocia Blandin, 1976c: 922, f. 11, 14 (mf)

Pisaura consocia Brignoli, 1984c: 39, f. 12, 14, 16 (mf)

Pisaura consocia Levy, 1999a: 58, f. 31A-B, 32A-B (mf)

Pisaura consocia Kunt et al., 2012: 641, f. 5a-c, 6a-c, 7a-d (mf)

Pisaura lama Bösenberg & Strand, 1906 | | Russia (Far East), China, Korea, Japan urn:lsid:nmbe.ch:spidersp:019757

Pisaura lama Bösenberg & Strand, 1906: 306, pl. 13, f. 430 (Df) Original description

Pisaura clarivittata Dönitz & Strand in Bösenberg & Strand, 1906: 389, pl. 6, f. 76 (Df) [urn:lsid:nmbe.ch:spidersp:056383]

Pisaura strandi Kishida, 1914b: 127, f. 1 (Df) [urn:lsid:nmbe.ch:spidersp:056385]

Pisaura lama Saito, 1939: 67, f. 8(6) (f)

Pisaura lama Saito, 1959: 45, f. 20a-b (f)

Pisaura strandi Yaginuma, 1960: 81, f. 74.2 (f)

Pisaura lama Yaginuma, 1960: 81, f. 74.3 (f)

Pisaura flavistriata Yaginuma, 1960: append. 6, pl. 38, f. 212, f. 74.4 (Df) [urn:lsid:nmbe.ch:spidersp:056384]

Pisaura lama Paik, 1969a: 44, f. 13, 15, 48-53, text-f. 1 (mf, S of Pisaura strandi)

Pisaura strandi Yaginuma, 1971: 81, f. 74.2 (f)

Pisaura lama Yaginuma, 1971: 81, f. 74.3 (f)

Pisaura flavistriata Yaginuma, 1971: 81, f. 74.4 (f)

Pisaura lama Yaginuma, 1974a: 170 (S of Pisaura clarivittata and P. flavistriata)

Pisaura lama Arita & Yaginuma, 1975: 29, f. 1-4 (mf)

Pisaura lama Paik, 1978e: 379, f. 172.1-6 (mf)

Pisaura laura Qiu, 1981: 14, f. 3A-C (f)

Pisaura lama Song & Zheng, 1982c: 156, f. 2 (f)

Pisaura ancora Hu, 1984: 261, f. 275.1-2, 276.1-2 (m, misidentified)

Pisaura lama Hu, 1984: 262, f. 275.3 (f)

Pisaura lama Guo, 1985: 135, f. 2-71.1-2 (f)

Pisaura lama Yaginuma, 1986a: 174, f. 96 (mf)

Pisaura lama Song, 1987: 210, f. 170 (f)

Pisaura lama Zhang, 1987: 168, f. 142.1-4 (mf)

Pisaura lama Chikuni, 1989b: 105, f. 1 (mf)

Pisaura lama Logunov, 1990: 37, f. 2-3 (mf)

Pisaura lama Feng, 1990: 163, f. 138.1-5 (mf)

Pisaura lama Chen & Gao, 1990: 137, f. 172 (f)

Pisaura lama Chen & Zhang, 1991: 227, f. 236 (f)

Pisaura lama Barrion & Litsinger, 1994: 305, f. 1579-1582 (mf)

Pisaura lama Song, Zhu & Chen, 1999: 348, f. 204D, K (mf)

Pisaura lama Song, Zhu & Chen, 2001: 269, f. 169A-D (mf)

Pisaura lama Namkung, 2002: 351, f. 21.6a-b (mf)

Pisaura lama Kim & Cho, 2002: 192, f. 403-408 (f)

Pisaura lama Namkung, 2003: 353, f. 21.6a-b (mf)

Pisaura lama Zhang, Zhu & Song, 2004d: 393, f. 124-127, 232-237 (mf)

Pisaura lama Tanikawa & Ono, 2009: 220, f. 35-37 (mf)

Pisaura lama Zhu & Zhang, 2011: 297, f. 216A-D (mf)

Pisaura lama Marusik & Kovblyuk, 2011: 219, f. 31.4 (f)

Pisaura lama Yin et al., 2012: 891, f. 448a-e (mf)

Pisaura lama Kim & Lee, 2013: 80, f. 53A-F, pl. 26-27 (mf)

Pisaura lama Suguro, 2013c: 14, f. 2, 4 (m)

Pisaura lama Baba & Tanikawa, 2015: 78, 6 f. (mf)

Pisaura lama Dong, 2018: 72, f. 10-5.A-G, pl. 16 (mf)

Pisaura mirabilis (Clerck, 1757) | | Europe, Turkey, Middle East, Caucasus, Russia (Europe to Middle Siberia), Central Asia, China urn:lsid:nmbe.ch:spidersp:019759

Araneus mirabilis Clerck, 1757: 108, pl. 5, f. 10 (Dmf) Original description

Aranea listeri Scopoli, 1763: 397 (D)

Aranea arcuato-lineata Martini & Goeze, 1778: 294 (D)

Aranea flavo-striata Martini & Goeze, 1778: 296 (D)

Aranea tripunctata Martini & Goeze, 1778: 297 (D)

Aranea rufo-fasciata De Geer, 1783: 269, pl. 16, f. 1-8 (Dmf)

Aranea marmorata Fourcroy, 1785: 535 (D)

Aranea agraria Olivier, 1789: 215 (D)

Aranea obscura Fabricius, 1793: 419 (D)

Araneus mirabilus Martyn, 1793: 50, pl. 7, f. 1-2 (mf)

Dolomedes mirabilis Walckenaer, 1805: 16, pl. 2, f. 17-18 (j)

Ocyale mirabilis Audouin, 1826: 373

Aranea bivittata Risso, 1826: 164 (D)

Dolomedes mirabilis Hahn, 1829: 1, pl. 2, f. A-C (m)

Dolomedes mirabilis Hahn, 1834: 35, f. 120 (f)

Ocyale murina C. L. Koch, 1837b: 23

Ocyale mirabilis C. L. Koch, 1847: 107, f. 1346 (m)

Ocyale rufofasciata C. L. Koch, 1847: 110, f. 1347 (Df)

Ocyale murina C. L. Koch, 1847: 111, f. 1348 (Df)

Dolomedes scheuchzeri Bremi-Wolff in Menzel, 1849: 12, f. 17 (D)

Dolomedes mirabilis Blackwall, 1861a: 37, pl. 2, f. 18 (mf)

Ocyale mirabilis Thorell, 1872a: 349

Ocyale mirabilis L. Koch, 1878b: 56, pl. 2, f. 3b (m)

Ocyale mirabilis Menge, 1879: 506, pl. 83, f. 287 (mf)

Ocyale mirabilis Hansen, 1882: 75, pl. 5, f. 8 (mf)

Ocyale mirabilis Becker, 1882b: 81, pl. 5, f. 1-17 (mf)

Pisaura mirabilis Simon, 1886g: 354

Pisaura mirabilis Simon, 1898a: 294, f. 303 (m)

Pisaura mirabilis Kulczyński, 1899: 422, pl. 9, f. 115 (m)

Pisaura mirabilis Smith, 1902: 329, f. a-d (mf)

Pisaura mirabilis Bösenberg, 1903: 408, pl. 39, f. 604 (mf)

Pisaura rufofasciata Kulczyński, 1903a: 54

Pisaura mirabilis Järvi, 1905: 26, pl. 4, f. 1-4, pl. 5, f. 1-2 (f)

Pisaura mirabilis Järvi, 1908: 755, f. 3 (f)

Pisaura listeri Dahl, 1908: 258, f. 21-23 (Dmf)

Ocyale mirabilis albida Franganillo, 1913: 120 (Df)

Ocyale mirabilis fusca Franganillo, 1913: 120 (Df)

Pisaura listeri Dahl & Dahl, 1927: 7, f. 6-9 (mf)

Pisaura mirabilis Berland, 1927b: 22, f. 17-18

Pisaura listeri Reimoser, 1932: 63, f. 9 (mf)

Pisaura listeri Spassky, 1935: 193, f. 1, 3 (mf)

Pisaura mirabilis Simon, 1937: 1048, 1051, f. 1615-1616 (mf)

Pisaura listeri Holm, 1947: 40, f. 17a-b, 18, pl. 8, f. 90, pl. 10, f. 48 (mf)

Pisaura mirabilis Locket & Millidge, 1951: 293, f. 141, 142A-B (mf)

Aranea bivittata Roewer, 1955c: 1395 (nicht zu deuten = nomen dubium)

Pisaura mirabilis Wiebes, 1959b: 67, f. 91, 110 (mf)

Pisaura mirabilis Azheganova, 1968: 22, f. 25-26 (mf)

Pisaura mirabilis Loksa, 1969: 130, f. 87, 88A-B (mf)

Pisaura mirabilis Miller, 1971: 171, pl. XXVII, f. 13-14 (mf)

Pisaura mirabilis Blandin, 1976c: 919, f. 1, 7a, 10, 13, 15, 18 (mf)

Pisaura mirabilis Brignoli, 1977f: 63, f. 37-40 (f)

Pisaura mirabilis Barrientos, 1978b: 18 (S)

Pisaura mirabilis Brignoli, 1978i: 206, f. 1-2 (f)

Pisaura mirabilis Hu, 1984: 263, f. 277.1-2 (m)

Pisaura mirabilis Brignoli, 1984c: 35, f. 1-2, 13, 15 (mf)

Pisaura mirabilis Roberts, 1985: 154, f. 67c (mf)

Pisaura mirabilis Song, 1987: 211, f. 171 (mf)

Pisaura mirabilis Wunderlich, 1987a: 229, f. 607-608 (m)

Pisaura mirabilis Hu & Wu, 1989: 241, f. 198.1-2 (m)

Pisaura mirabilis Izmailova, 1989: 45, f. 33 (f)

Pisaura mirabilis Sierwald, 1990: 31, f. 7, 30, 45 (m)

Pisaura sp. Logunov, 1990: 37, f. 2и (mf)

Pisaura mirabilis Heimer & Nentwig, 1991: 350, f. 911 (mf)

Pisaura mirabilis Zhao, 1993: 307, f. 141 (m)

Pisaura mirabilis Baldacchino et al., 1993: 51, f. 7d (m)

Pisaura mirabilis Samm, 1994: 1, f. 2 (m)

Pisaura mirabilis Roberts, 1995: 237, f. (mf)

Pisaura mirabilis Zhao, 1995: 1024, f. 480 (m)

Pisaura mirabilis Mcheidze, 1997: 246, f. 533-535 (mf)

Pisaura mirabilis Roberts, 1998: 254, f. (mf)

Pisaura mirabilis Song, Zhu & Chen, 1999: 348, f. 204F, N (mf)

Pisaura mirabilis Levy, 1999a: 55, f. 29A-C, 30A-F (mf)

Pisaura mirabilis Zhang, 2000a: 4, f. 4A-F (mf)

Pisaura mirabilis Hu, 2001: 220, f. 116.1-4 (mf)

Pisaura mirabilis Zhang, Zhu & Song, 2004d: 394, f. 128-133, 238-243 (mf)

Pisaura mirabilis Almquist, 2005: 259, f. 249a-h (mf)

Pisaura mirabilis Kovblyuk et al., 2011: 38, f. 35-36, 39-40, 42 (f)

Pisaura mirabilis Kunt et al., 2012: 642, f. 7e-h, 8a-f (mf)

Pisaura mirabilis Nadolny et al., 2012: 256, f. 1, 3, 5, 7, 9-11, 15-16, 19, 22-23, 26-27, 31-32, 35, 39 (mf)

Pisaura mirabilis Fedoriak & Moscaliuc, 2013: 152, f. 4B, 5B (mf)

Pisaura mirabilis Kovblyuk, 2014: 47, f. 32 (mf)

Pisaura mirabilis Fedoriak, 2015: 106, f. 2.6.7B, 2.6.8B (mf)

Pisaura mirabilis Esyunin & Sozontov, 2015: 326, f. 1-4, 9-12, 17-18 (mf)

Pisaura mirabilis Fomichev, Marusik & Koponen, 2018: 334, f. 4a-b (m)

Pisaura mirabilis Breitling, 2020: 353, f. 28 (f)

Pisaura mirabilis Lecigne, 2021a: 33, f. 46 (f)

Pisaura mirabilis Sarkar, Siliwal & Uniyal, 2023c: 647, f. 12, 15 (mf)

Pisaura novicia (L. Koch, 1878) | | Mediterranean to Central Asia, India urn:lsid:nmbe.ch:spidersp:019760

Ocyale novicia L. Koch, 1878b: 54, pl. 2, f. 3, 3a (Dmf) Original description

Pisaura novicia Simon, 1898a: 289

Pisaura novicia Blandin, 1976c: 922, f. 12 (Df)

Pisaura novicia Brignoli, 1984c: 36, f. 3-4 (f)

Pisaura mirabilis Logunov, 1990: 37, f. 2в-з,3II (mf) [misidentified after Nadolny et al., 2012: 260]

Pisaura novicia Mcheidze, 1997: 246, f. 536 (f)

Pisaura novicia Kovblyuk et al., 2011: 38, f. 37-38, 41, 43 (f)

Pisaura novicia Nadolny et al., 2012: 260, f. 2, 4, 6, 8, 12-14, 17-18, 20-21, 24-25, 28-30, 33-34, 36-38, 40-45 (mf)

Pisaura novicia Fedoriak & Moscaliuc, 2013: 152, f. 4A, 5A (mf)

Pisaura novicia Kovblyuk, 2014: 47, f. 31-32 (mf)

Pisaura novicia Fedoriak, 2015: 106, f. 2.6.7A, 2.6.8A (mf)

Pisaura novicia Esyunin & Sozontov, 2015: 327, f. 5-8, 13-16, 19 (mf)

Pisaura novicia Sarkar, Siliwal & Uniyal, 2023c: 644, f. 1-11, 13-14, 16 (mf)

Pisaura orientalis Kulczyński, 1913 | | Mediterranean urn:lsid:nmbe.ch:spidersp:019761

Pisaura mirabilis orientalis Kulczyński, 1913a: 28 (Df) Original description

Pisaura orientalis Brignoli, 1984c: 38, f. 5-8 (f, elevated from subspecies)

Pisaura podilensis Patel & Reddy, 1990 | | India urn:lsid:nmbe.ch:spidersp:019763

Pisaura podilensis Patel & Reddy, 1990a: 38, f. 2a-g (Dmf) Original description

Pisaura quadrilineata (Lucas, 1838) | | Canary Is., Madeira urn:lsid:nmbe.ch:spidersp:019765

Philodromus quadrilineatus Lucas, 1838: 34, pl. 6, f. 11 (Dj) Original description

Pisaura mirabilis maderiana Kulczyński in Schmitz, 1895: 197 (Dmf)

Pisaura mirabilis maderiana Kulczyński, 1899: 422, pl. 9, f. 116, 123 (f)

Tibellus quadrilineatus Denis, 1941a: 121

Thanatus quadrilineatus Roewer, 1955c: 792

Pisaura maderiana Brignoli, 1984c: 38, f. 9-11 (f, elevated from subspecies of P. mirabilis)

Pisaura maderiana Wunderlich, 1987a: 230, f. 609-610 (m)

Pisaura quadrilineata Wunderlich, 1992a: 438, f. 660-662 (Tm from Thanatus, Sf)

Pisaura sublama Zhang, 2000 | | China urn:lsid:nmbe.ch:spidersp:019767

Pisaura sublama Zhang, 2000a: 5, f. 6A-E (Dmf) Original description

Pisaura sublama Zhang, Zhu & Song, 2004d: 396, f. 134-140, 244-249 (mf)

Pisaura sublama Zhu & Zhang, 2011: 298, f. 217A-G (mf)

Pisaura swamii Patel, 1987 | | India urn:lsid:nmbe.ch:spidersp:019768

Pisaura swamii Patel, 1987a: 64, f. a-e (Dmf) Original description