World Spider Catalog

Lycosidae Sundevall, 1833

Family | ACCEPTED

N.B.: according to Brignoli, 1983c: 432, "Guy (1966, 1969) has proposed many synonymies and has downgraded many genera to subgenus rank; as this author based his conclusions more on nomenclature than on the study of material (or even of the original descriptions)", Platnick (WSC 15.0) did not accept Guy's conclusions; the changes by Guy, 1966 are noted (except where more recent work has superseded them); Guy, 1969 is merely a literature compilation that reflects those changes and is not cited separately; several genera of Lycosidae have been found not to be monophyletic (Murphy et al., 2006). Piacentini & Ramírez, 2019: 233-237 confirm ten valid subfamilies of Lycosidae: Venoniinae Lehtinen & Hippa, 1979, Zoicinae Lehtinen & Hippa, 1979 (= Piratinae Zyuzin, 1993), Evippinae Zyuzin, 1985, Sosippinae Dondale, 1986 (= Hygrolycosinae He & Song, 1996), Artoriinae Framenau, 2007, Tricassinae Alderweireldt & Jocqué, 1993 (= Arctosinae He & Song, 1996), Hippasinae Simon, 1898, Allocosinae Dondale, 1986, Pardosinae Simon, 1898 (= Wadicosinae Zyuzin, 1985) and Lycosinae Sundevall, 1833; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 518, f. 17-18.

Gen. Piratula Roewer, 1960 urn:lsid:nmbe.ch:spidergen:04202

type Pirata hygrophilus Thorell, 1872; gender feminine

N.B.: already used by Roewer, 1955c: 287 as nomen nudum; removed from the synonymy of Pirata Sundevall, 1833 by Omelko, Marusik & Koponen, 2011: 213, contra Dondale & Redner, 1981b: 107, Breitling, 2019b: 21 and Breitling, 2021: 26 (the two latter mainly based on COI, see FAQ 7).

In synonymy:

Piratula bilobata (Tullgren, 1901, T from Pardosa) = Piratula insularis (Emerton, 1885) (Wallace, 1950: 77, sub Pirata)

Piratula eugeni (Roewer, 1951) = Piratula knorri (Scopoli, 1763) (Buchar, 1966a: 217, sub Pirata)

Piratula exigua (Banks, 1892) = Piratula minuta (Emerton, 1885) (Chamberlin, 1908: 303, sub Pirata)

Piratula humicola (Montgomery, 1902, T from Allocosa) = Piratula minuta (Emerton, 1885) (Wallace & Exline, 1978: 96, sub Pirata)

Piratula japonica (Tanaka, 1974) = Piratula piratoides (Bösenberg & Strand, 1906) (Yaginuma, 1978a: 16, sub Pirata)

Piratula libera (Montgomery, 1902) = Piratula insularis (Emerton, 1885) (Chamberlin, 1908: 311, sub Pirata)

Piratula moodyi (Hull, 1950) = Piratula uliginosa (Thorell, 1856) (Buchar, 1966a: 218, sub Pirata)

Piratula nanshanensis (Yin & Wang, 1984) = Piratula procurva (Bösenberg & Strand, 1906) (Yu & Song, 1988c: 119, sub Pirata)

Piratula piccolo (Dahl, 1908) = Piratula insularis (Emerton, 1885) (Hackman, 1954: 32, sub Pirata)

Piratula praedatoria (Schenkel, 1953) = Piratula piratoides (Bösenberg & Strand, 1906) (Song, 1987: 240, sub Pirata)

Piratula procurva sinensis (Song, Shang, Wang, C. F. Cheng & S. X. Cheng, 1978) = Piratula procurva (Bösenberg & Strand, 1906) (Song, 1987: 241, sub Pirata)

Piratula qinlingensis (Zheng & Qiu, 1980) = Piratula yaginumai (Tanaka, 1974) (Song, 1987: 244, sub Pirata)

Piratula sagaphila (Strand, 1918, T from Hyaenosa) = Piratula procurva (Bösenberg & Strand, 1906) (Tanaka, 1988a: 57, sub Pirata)

Piratula wuchangensis (Schenkel, 1963) = Piratula piratoides (Bösenberg & Strand, 1906) (Song, 1987: 240, sub Pirata)

Homonym replaced:

Piratula exigua (Tanaka, 1974) -- see Piratula tanakai Brignoli, 1983 Brignoli, 1983c

Piratula acuminata Sudhin, Sankaran & Sen, 2025 | | India urn:lsid:nmbe.ch:spidersp:065508

Piratula acuminata Sudhin, Sankaran & Sen, 2025: 594, f. 1A-L, 2A-F (Dmf) Original description

Piratula borea (Tanaka, 1974) | | Russia (Far East), China, Japan urn:lsid:nmbe.ch:spidersp:019059

Pirata boreus Tanaka, 1974a: 33, f. 18-20 (Dmf) Original description

Pirata boreus Tanaka, 1988a: 55, f. 25-28 (mf)

Pirata boreus Song, Zhu & Chen, 1999: 335, f. 200D (f)

Pirata boreus Tanaka, 2009: 225, f. 26-27 (mf)

Piratula borea Omelko, Marusik & Koponen, 2011: 218, f. 48-52 (mf, T from Pirata)

Pirata boreus Suzuki, 2019c: 84, f. 11e-f (f)

Piratula borea Tanikawa, 2021: 68, f. 3 (f)

Piratula canadensis (Dondale & Redner, 1981) | | Russia (Far East, Kamchatka), Canada urn:lsid:nmbe.ch:spidersp:019064

Pirata canadensis Dondale & Redner, 1981a: 194, f. 1-8 (Dmf) Original description

Pirata canadensis Dondale & Redner, 1990: 251, f. 366-373 (mf, removed from S of Pirata procurvus, rejecting Liu, 1987b: 85)

Pirata canadensis Paquin & Dupérré, 2003: 167, f. 1870-1873 (mf)

Piratula canadensis Omelko, Marusik & Koponen, 2011: 221, f. 4b, 53-64, 146-147, 158 (mf, T from Pirata)

Piratula cantralli (Wallace & Exline, 1978) | | Canada, USA urn:lsid:nmbe.ch:spidersp:019065

Pirata cantralli Wallace & Exline, 1978: 46, f. 81-82, 88-89 (Dmf) Original description

Pirata cantralli Dondale & Redner, 1981a: 193, f. 9-10 (m)

Pirata cantralli Dondale & Redner, 1990: 253, f. 374-379 (mf)

Pirata cantralli Paquin & Dupérré, 2003: 167, f. 1874-1877 (mf)

Piratula cantralli Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Piratula clercki (Bösenberg & Strand, 1906) | | China, Korea, Taiwan, Japan urn:lsid:nmbe.ch:spidersp:019068

Tarentula clercki Bösenberg & Strand, 1906: 316, pl. 8, f. 107, pl. 13, f. 320 (Df) Original description

Pirata clercki Fox, 1935: 456

Piratula clercki Roewer, 1955c: 287

Pirata clercki Yaginuma, 1960: 87, f. 78.2 (f)

Pirata clercki Yaginuma, 1965b: 366, f. 9.8 (f)

Pirata clercki Yaginuma, 1971: 87, f. 78.2 (f)

Pirata clercki Tanaka, 1974a: 34, f. 21-24 (mf)

Pirata clercki Yoshida, 1978b: 10, f. 16-17 (mf)

Pirata clercki Hu, 1984: 242, f. 251.1-2 (f)

Pirata clercki Yaginuma, 1986a: 167, f. 92.3 (mf)

Pirata clercki Liu, 1987a: 45, pl. 1, f. 4 (m)

Pirata clercki Liu, 1987b: 86, f. 1-5 (mf)

Pirata clercki Tanaka, 1988a: 42, f. 9-12 (mf)

Pirata clercki Chikuni, 1989b: 113, f. 18 (mf)

Pirata clercki Chen & Zhang, 1991: 215, f. 220.1-4 (mf)

Pirata clercki Barrion & Litsinger, 1994: 317, f. 1662-1665 (mf)

Pirata clercki Yin et al., 1997c: 27, f. 9a-d (mf)

Pirata clercki Song, Zhu & Chen, 1999: 335, f. 200E, Q (mf)

Pirata clercki Namkung, 2002: 340, f. 20.34a-b (mf)

Pirata clercki Kim & Cho, 2002: 219, f. 487-492 (f)

Pirata clercki Namkung, 2003: 342, f. 20.34a-b (mf)

Pirata clercki Lee et al., 2004: 99, unnumbered f. (m)

Pirata clercki Tanaka, 2009: 225, f. 18-19 (mf)

Pirata clercki Zhu & Zhang, 2011: 282, f. 204A-E (mf)

Piratula clercki Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Pirata clercki Yin et al., 2012: 772, f. 384a-d (mf)

Piratula denticulata (Liu, 1987) | | Russia (Middle Siberia to Far North-East), China, Taiwan urn:lsid:nmbe.ch:spidersp:019071

Pirata denticulata Liu, 1987a: 46, pl. 2, f. 8 (m, nomen nudum)

Pirata denticulata Liu, 1987c: 367, f. 1-6 (Dmf) Original description

Pirata denticulata Chen & Zhang, 1991: 211, f. 215.1-6 (mf)

Pirata denticulata Yin et al., 1997c: 29, f. 10a-f (mf)

Pirata denticulatus Song, Zhu & Chen, 1999: 335, f. 200F, R (mf)

Pirata denticulatus Tso & Chen, 2004: 407, f. 39-40 (m)

Piratula denticulata Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Pirata denticulata Yin et al., 2012: 774, f. 385a-f (mf)

Piratula gigantea (Gertsch, 1934) | | USA urn:lsid:nmbe.ch:spidersp:019074

Pirata gigantea Gertsch, 1934a: 12 (Df) Original description

Arkalosula gigantea Roewer, 1955c: 232

Pirata giganteus Wallace & Exline, 1978: 89, f. 187-191 (Dm, f, T from Arkalosula)

Pirata giganteus Dondale & Redner, 1981b: 107, f. 5-6 (m)

Pirata giganteus Dondale & Redner, 1990: 259, f. 390-395 (mf)

Piratula gigantea Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Piratula hiroshii (Tanaka, 1986) | | Japan urn:lsid:nmbe.ch:spidersp:019076

Pirata hiroshii Tanaka, 1986b: 19, f. 1-4 (Dmf) Original description

Pirata hiroshii Tanaka, 1988a: 66, f. 41-44 (mf)

Pirata hiroshii Tanaka, 2009: 227, f. 38-39 (mf)

Piratula hiroshii Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Piratula hokkaidensis (Tanaka, 2003) | | Japan urn:lsid:nmbe.ch:spidersp:038690

Pirata hokkaidensis Tanaka, 2003: 83, f. 1-4 (Dmf) Original description

Pirata hokkaidensis Tanaka, 2009: 227, f. 36-37 (mf)

Piratula hokkaidensis Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Piratula hurkai (Buchar, 1966) | | Ukraine, Caucasus (Russia, Georgia), Turkey urn:lsid:nmbe.ch:spidersp:019078

Pirata hurkai Buchar, 1966a: 210, f. 1A-B, E-F, 2A, 4E (Dmf) Original description

Pirata hurkai Ovtsharenko, 1979: 40, f. 33 (m)

Pirata hurkai Mcheidze, 1997: 244, f. 529-530 (mf)

Pirata hurkai Nadolny & Kovblyuk, 2011: 176, f. 1, 6, 10, 16-31, 35-46 (mf)

Piratula hurkai Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Piratula hurkai Türkeş, 2024b: 339, f. 2, 3A-B (m)

Piratula hygrophila (Thorell, 1872) | | Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Kazakhstan, Central Asia urn:lsid:nmbe.ch:spidersp:019079 TYPE SPECIES

Lycosa piscatoria Blackwall, 1861a: 36, pl. 2, f. 17 (mf; misidentified per Thorell, 1872a: 346 and Thorell, 1873: 419)

Pirata hygrophilus Thorell, 1872a: 346 (new name for Lycosa piscatoria sensu Blackwall) Original description

Lycosa hygrophila O. Pickard-Cambridge, 1873c: 523

Pirata hygrophilus Simon, 1876a: 297 (Dmf)

Pirata hygrophilus Menge, 1879: 514, pl. 84, f. 291 (mf)

Pirata knorri Becker, 1882b: 118, pl. 9, f. 6b, c (f; misidentified per Dahl, 1908: 401)

Pirata hygrophilus Becker, 1882b: 120, pl. 9, f. 7a (f; m f. 7b see Pardosa amentata)

Pirata hygrophilus Chyzer & Kulczyński, 1891: 72, pl. 2, f. 7 (mf)

Pirata hygrophilus Bösenberg, 1903: 406, pl. 38, f. 599 (mf)

Pirata hygrophilus Dahl, 1908: 294, f. 29 (mf)

Pirata hygrophilus Dahl & Dahl, 1927: 59, f. 155-156 (mf)

Lycosa hygrophila Simon, 1937: 1116, 1140, f. 1760-1761 (mf)

Pirata hygrophilus Palmgren, 1939: 68, f. 102, 107, 112 (mf)

Pirata hygrophilus Holm, 1947: 9, f. 7a, pl. 1, f. 3-4, pl. 9, f. 1 (mf)

Pirata hygrophilus Locket & Millidge, 1951: 289, f. 139D, H (mf)

Piratula hygrophila Roewer, 1955c: 287

Piratula hygrophila Muller, 1958: 233, f. 5-6 (mf)

Pirata hygrophilus Wiebes, 1959b: 63, f. 94, 96 (mf)

Pirata hygrophilus Azheganova, 1971: 11, f. 16 (mf)

Pirata hygrophilus Tystshenko, 1971: 182, f. 528, 532 (mf)

Pirata hygrophilus Fuhn & Niculescu-Burlacu, 1971: 206, f. 100a-e (mf)

Pirata hygrophilus Miller, 1971: 168, pl. XXVI, f. 14-15 (mf)

Pirata hygrophilus Loksa, 1972: 58, f. 52A-B, 55C-D (mf)

Pirata hygrophilus Kronestedt, 1980b: 65, f. 9E-F, 10B (m)

Pirata hygrophilus Dondale & Redner, 1981b: 107, f. 7 (m)

Pirata hygrophilus Kronestedt, 1984: 105, f. 11-12

Pirata hygrophilus Roberts, 1985: 150, f. 65c (mf)

Pirata hygrophilus Zyuzin, 1985a: 44, f. 8

Pirata hygrophilus Izmailova, 1989: 41, f. 30 (f)

Pirata hygrophilus Heimer & Nentwig, 1991: 346, 894 (mf)

Pirata hygrophilus Roberts, 1995: 233, f. (mf)

Pirata hygrophilus Roberts, 1998: 251, f. (mf)

Pirata hygrophilus Almquist, 2006: 241, f. 236a-h (mf)

Pirata hygrophilus Nadolny & Kovblyuk, 2011: 180, f. 54-78 (mf)

Piratula hygrophila Omelko, Marusik & Koponen, 2011: 223, f. 2a, 65-70, 163 (mf, T from Pirata)

Piratula insularis (Emerton, 1885) | | North America, Europe, Russia (Europe to Middle Siberia) urn:lsid:nmbe.ch:spidersp:019081

Pirata insularis Emerton, 1885: 492, pl. 48, f. 8 (Df) Original description

Pardosa bilobata Tullgren, 1901a: 22, pl. 1, f. 12 (Df) [urn:lsid:nmbe.ch:spidersp:063997]

Pirata liber Montgomery, 1902: 578, pl. 30, f. 42-43 (Dmf) [urn:lsid:nmbe.ch:spidersp:063998]

Pirata insularis Chamberlin, 1908: 309, pl. 22, f. 3 (m, S of Pirata liber; f f. 4 see Pirata aspirans)

Pirata piccolo Dahl, 1908: 301, f. 30 (Dmf) [urn:lsid:nmbe.ch:spidersp:063996]

Pirata aspirans Chamberlin, 1908: 303, pl. 22, f. 6 (f; misidentified per Kaston, 1948: 310)

Pirata insularis Emerton, 1909: 208, pl. 6, f. 7

Pirata sylvestris Emerton, 1909: 209, pl. 6, f. 8a (m; misidentified per Kaston, 1948: 310)

Pirata aspirans Comstock, 1912: 643, f. 724b (f; misidentified per Kaston, 1948: 310)

Pirata piccolo Dahl & Dahl, 1927: 60, f. 157-158 (mf)

Pirata insularis Kaston, 1938a: 16, f. 10 (f)

Pirata piccolo Palmgren, 1939: 68, f. 108, 110, 113 (f)

Pirata insularis Comstock, 1940: 655, f. 724b (f)

Pirata insularis Muma, 1943: 53, pl. XII, f. 13 (f)

Pirata piccolo Holm, 1947: 10, pl. 1, f. 9-10, pl. 9, f. 5 (mf)

Pirata insularis Kaston, 1948: 310, f. 987-988, 1005, 1011 (mf)

Pirata insularis Wallace, 1950: 77, pl. 2, f. 9 (f, S of Pardosa bilobata)

Pirata insularis Hackman, 1954: 8, 32, 95 (S of Pirata piccolo)

Allocosa insularis Roewer, 1955c: 210

Pirata piccolo Tystshenko, 1971: 182, f. 533 (f)

Pirata piccolo Fuhn & Niculescu-Burlacu, 1971: 212, f. 103a-c (mf)

Pirata insularis Wallace & Exline, 1978: 40, f. 2, 77-80, 83-86 (mf, T from Allocosa)

Pirata piccolo Kronestedt, 1980b: 65, f. 9I, K (m)

Pirata insularis Dondale & Redner, 1981a: 193, f. 11-12 (m)

Pirata insularis Dondale & Redner, 1990: 255, f. 380-386 (mf)

Pirata piccolo Heimer & Nentwig, 1991: 346, f. 896 (mf)

Pirata insularis Roberts, 1995: 234, f. (mf)

Pirata piccolo Mcheidze, 1997: 243, f. 527-528 (f)

Pirata insularis Roberts, 1998: 251, f. (mf)

Pirata insularis Paquin & Dupérré, 2003: 167, f. 1878-1881 (mf)

Pirata insularis Almquist, 2006: 241, f. 237a-e (mf)

Piratula insularis Omelko, Marusik & Koponen, 2011: 224, f. 71-79, 154-155, 166 (mf, T from Pirata)

Piratula iriomotensis (Tanaka, 1989) | | Japan (Ryukyu Is.) urn:lsid:nmbe.ch:spidersp:019082

Pirata iriomotensis Tanaka, 1989d: 109, f. 1-4 (Dmf) Original description

Pirata iriomotensis Tanaka, 2009: 227, f. 30-31 (mf)

Piratula iriomotensis Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Piratula knorri (Scopoli, 1763) | | Europe, Caucasus, Russia (Europe to West Siberia) urn:lsid:nmbe.ch:spidersp:019084

Aranea knorrii Scopoli, 1763: 403 (D) Original description

Lycosa piscatoria C. L. Koch, 1847: 6, f. 1417-1419 (mf; misidentified per Dahl, 1908: 296)

Pirata knorri Simon, 1876a: 296 (Dmf)

Pirata knorri Bösenberg, 1903: 407, pl. 39, f. 601 (mf)

Pirata knorri Dahl, 1908: 296, f. 32 (mf)

Lycosa knorri Lessert, 1910b: 501 (Dmf)

Pirata knorri Dahl & Dahl, 1927: 62, f. 161-163 (mf)

Pirata knorri Saito, 1934b: 352, pl. 15, f. 82 (f)

Lycosa knorri Simon, 1937: 1118, 1140, f. 1764 (mf)

Lycosa furcifera Simon, 1937: 1118, 1140, f. 1765-1766 (Dmf; preoccupied by Thorell, 1875, see Pardosa furcifera)

Pirata knorri Saito, 1939: 72, f. 6.4 (f)

Pirata eugeni Roewer, 1951: 440 (replacement name for Lycosa furcifera) [urn:lsid:nmbe.ch:spidersp:063985]

Piratula eugeni Roewer, 1955c: 287

Piratula knorri Roewer, 1955c: 288

Pirata knorri Saito, 1959: 58, text-f. 7.8, f. 43a-e (mf)

Pirata knorri Wiebes, 1959b: 64, f. 89, 93 (mf)

Pirata knorri Buchar, 1966a: 212, f. 1C-D, 2B, 4A-D (mf, S of Lycosa furcifera and [automatically] of its replacement name Pirata eugeni)

Pirata knorri Tystshenko, 1971: 182, f. 536 (f)

Pirata knorri Fuhn & Niculescu-Burlacu, 1971: 208, f. 101a-f (mf)

Pirata knorri Miller, 1971: 169, pl. XXVI, f. 20 (m)

Pirata knorri Loksa, 1972: 60, f. 53A-B, 54A-B (mf)

Pirata knorrii Paik, 1978e: 107, f. 25.3 (f; probably misidentified, per Komnenov, 2014: 125)

Pirata knorri Tanaka, 1988a: 69, f. 45-47 (mf)

Pirata knorri Heimer & Nentwig, 1991: 346, f. 897 (mf)

Pirata knorri Roberts, 1995: 235, f. (mf)

Pirata knorri Mcheidze, 1997: 244, f. 531-532 (f)

Pirata knorri Roberts, 1998: 252, f. (mf)

Pirata knorri Nadolny & Kovblyuk, 2011: 176, f. 2, 11, 32-34, 79-82 (mf)

Piratula knorri Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Piratula knorri Demircan & Topçu, 2015: 181, f. 7A-C (f)

Piratula knorri Kuralt, Pajek Arambašič & Kostanjšek, 2025: 115, f. 2i-j (mf)

Piratula latitans (Blackwall, 1841) | | Europe, Turkey, Caucasus, Iran urn:lsid:nmbe.ch:spidersp:019085

Lycosa latitans Blackwall, 1841a: 612 (Dmf) Original description

Lycosa palustris C. L. Koch, 1847: 4, f. 1415-1416 (mf; misidentified per Thorell, 1873: 419)

Potamia palustris C. L. Koch, 1850: 35 (misidentified sensu Thorell, 1873: 419)

Lycosa latitans Blackwall, 1861a: 33, pl. 2, f. 15 (mf)

Pirata latitans Thorell, 1873: 419

Pirata latitans Bösenberg, 1903: 407, pl. 39, f. 600 (mf)

Pirata latitans Dahl, 1908: 298, f. 31 (mf)

Pirata latitans Dahl & Dahl, 1927: 61, f. 159-160 (mf)

Lycosa latitans Simon, 1937: 1117, 1139, f. 1762-1763 (mf)

Pirata latitans Locket & Millidge, 1951: 289, f. 139C, E (mf)

Pirata latitans Muller, 1955: 159, f. 2 (m)

Piratula latitans Roewer, 1955c: 288

Pirata latitans Wiebes, 1959b: 65, f. 106-107 (mf)

Pirata latitans Tystshenko, 1971: 182, f. 529, 534 (mf)

Pirata latitans Fuhn & Niculescu-Burlacu, 1971: 210, f. 1021-e (mf)

Pirata latitans Miller, 1971: 169, pl. XXVI, f. 18-19 (mf)

Pirata latitans Loksa, 1972: 59, f. 52D-E, 55E-F (mf)

Pirata latitans Roberts, 1985: 150, f. 65e (mf)

Pirata latitans Alderweireldt, 1989: 14, f. 11 (f)

Pirata latitans Heimer & Nentwig, 1991: 346, f. 898 (mf)

Pirata latitans Roberts, 1995: 234, f. (mf)

Pirata latitans Roberts, 1998: 252, f. (mf)

Pirata latitans Guryanova, 2003: 6, f. 14 (f)

Pirata latitans Bayram et al., 2007b: 80, f. 1d (f)

Pirata latitans Nadolny & Kovblyuk, 2011: 182, f. 3, 7, 12, 83-103 (mf)

Piratula latitans Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Piratula logunovi Omelko, Marusik & Koponen, 2011 | | Russia (Far East) urn:lsid:nmbe.ch:spidersp:044370

Piratula logunovi Omelko, Marusik & Koponen, 2011: 224, f. 4a, 80-91, 150-151, 161, 173 (Dmf) Original description

Piratula logunovi Zamani & Marusik, 2021f: 454, f. 23-25 (m)

Piratula longjiangensis (Yin, Peng, Xie, Bao & Wang, 1997) | | China urn:lsid:nmbe.ch:spidersp:019086

Pirata longjiangensis Yan et al., 1997: 17, f. 1-5 (Dmf) Original description

Pirata longjiangensis Yin et al., 1997c: 32, f. 12a-e (mf)

Pirata longjiangensis Song, Zhu & Chen, 1999: 335, f. 200H, T (mf)

Piratula longjiangensis Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Piratula meridionalis (Tanaka, 1974) | | Russia (Far East), China, Korea, Japan urn:lsid:nmbe.ch:spidersp:019089

Pirata meridionalis Tanaka, 1974a: 31, f. 15-17 (Dmf) Original description

Pirata meridionalis Sohn & Paik, 1981: 20, f. 1-10 (mf)

Pirata meridionalis Liu, 1987a: 46, pl. 2, f. 1-3 (m)

Pirata meridionalis Liu, 1987c: 371, f. 13-17 (mf)

Pirata meridionalis Tanaka, 1988a: 52, f. 21-24 (mf)

Pirata meridionalis Chikuni, 1989b: 114, f. 23 (mf)

Pirata meridionalis Barrion & Litsinger, 1994: 315, f. 1648-1652 (mf)

Pirata meridionalis Yin et al., 1997c: 34, f. 13a-d (mf)

Pirata meridionalis Song, Zhu & Chen, 1999: 344, f. 200I, U (mf)

Pirata meridionalis Namkung, 2002: 341, f. 20.35a-b (mf)

Pirata meridionalis Namkung, 2003: 343, f. 20.35a-b (mf)

Pirata meridionalis Tanaka, 2009: 225, f. 24-25 (mf)

Piratula meridionalis Omelko, Marusik & Koponen, 2011: 226, f. 4c-d, 92-101, 152-153, 165, 170 (mf, T from Pirata)

Pirata meridionalis Yin et al., 2012: 777, f. 387a-d (mf)

Piratula meridionalis Tanikawa, 2021: 68, f. 2 (f)

Piratula meridionalis Wang et al., 2021a: 55, f. 52A-H, 53A-E, 54A-D (mf)

Piratula meridionalis Hidaka, 2022b: 47, f. 7-9 (mf)

Piratula minuta (Emerton, 1885) | | North America urn:lsid:nmbe.ch:spidersp:019090

Pirata minutus Emerton, 1885: 493, pl. 48, f. 10 (Dmf) Original description

Pirata exigua Banks, 1892a: 72, pl. 1, f. 48 (Df) [urn:lsid:nmbe.ch:spidersp:063991]

Lycosa minuta Simon, 1898a: 335

Pirata humicolus Montgomery, 1902: 575, pl. 30, f. 40-41 (Dmf) [urn:lsid:nmbe.ch:spidersp:063986]

Aulonia humicola Montgomery, 1904: 265, pl. 20, f. 33 (f)

Pirata minuta Chamberlin, 1908: 301, pl. 21, f. 9, pl. 22, f. 33 (f, S of Pirata exigua)

Pirata minuta Comstock, 1912: 643, f. 724a

Pirata minuta Kaston, 1938a: 16, f. 9 (f)

Pirata minutus Comstock, 1940: 654, f. 724a (f)

Pirata minutus Muma, 1943: 53, pl. XII, f. 14 (f)

Pirata minuta Kaston, 1948: 307, f. 1001, 1008 (mf)

Allocosa humicola Roewer, 1955c: 210

Piratessa minutus Roewer, 1955c: 286 (in generic nomen nudum)

Pirata minutus Wallace & Exline, 1978: 96, f. 2, 205-216 (mf, T from Piratessa = Piratosa, S of Allocosa humicola)

Pirata minutus Dondale & Redner, 1990: 262, f. 400-403 (mf)

Pirata minutus Paquin & Dupérré, 2003: 168, f. 1882-1884 (mf)

Piratula minuta Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Piratula montigena (Liu, 1987) | | China urn:lsid:nmbe.ch:spidersp:019094

Pirata montigena Liu, 1987a: 46, pl. 2, f. 7 (m, nomen nudum)

Pirata montigena Liu, 1987c: 369, f. 7-12 (Dmf) Original description

Pirata montigena Yin et al., 1997c: 35, f. 14a-d (mf)

Pirata montigenus Song, Zhu & Chen, 1999: 344, f. 200J, 201A (mf)

Piratula montigena Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Piratula piratoides (Bösenberg & Strand, 1906) | | Russia (Far East), China, Korea, Japan urn:lsid:nmbe.ch:spidersp:019103

Tarentula piratoides Bösenberg & Strand, 1906: 318, pl. 13, f. 336 (Df) Original description

Pirata praedatoria Schenkel, 1953b: 80, f. 37 (Df) [urn:lsid:nmbe.ch:spidersp:063988]

Piratula piratoides Roewer, 1955c: 288

Pirata wuchangensis Schenkel, 1963: 356, f. 206a-c (Df) [urn:lsid:nmbe.ch:spidersp:063989]

Pirata japonicus Tanaka, 1974a: 29, f. 11-14 (Dmf) [urn:lsid:nmbe.ch:spidersp:063987]

Pirata japonicus Yin, 1978: 11, f. 6A-D (mf)

Pirata praedatoria Song et al., 1978: 4, f. 8A-E (f, Dm)

Pirata piratoides Yaginuma, 1978a: 16 (S of Pirata japonicus)

Pirata japonicus Song, 1980: 170, f. 92a-e (mf)

Pirata japonicus Wang, 1981: 121, f. 61A-B (f)

Pirata praedatorius Brignoli, 1983c: 456

Pirata japonicus Hu, 1984: 242, f. 252.1-5 (mf)

Pirata praedatoria Hu, 1984: 243, f. 254.1-5 (mf)

Pirata praedatoria Guo, 1985: 124, f. 2-62.1-3 (mf)

Pirata preadatoria Zhu & Shi, 1985: 141, f. 125a-d (mf)

Pirata piratoides Yaginuma, 1986a: 167, f. 92.4 (mf)

Pirata piratoides Song, 1987: 240, f. 195 (mf, S of Pirata praedatoria and P. wuchangensis)

Pirata piratoides Zhang, 1987: 158, f. 134.1-4 (mf)

Pirata piratoides Liu, 1987a: 45, pl. 1, f. 5-6 (m, S of Pirata praedatoria confirmed)

Pirata piratoides Liu, 1987b: 85, f. 12-17 (mf, S of Pirata praedatoria and P. wuchangensis confirmed)

Pirata piratoides Tanaka, 1988a: 49, f. 17-20 (mf)

Pirata piratoides Chikuni, 1989b: 113, f. 20 (mf)

Pirata piratoides Chen & Gao, 1990: 130, f. 163a-b (mf)

Pirata piratoides Chen & Zhang, 1991: 213, f. 218.1-4 (mf)

Pirata piratoides Song, Zhu & Li, 1993: 873, f. 39A-E (mf)

Pirata piratoides Zhao, 1993: 105, f. 45a-c (mf)

Pirata piratoides Barrion & Litsinger, 1994: 315, f. 1653-1658 (mf)

Pirata piratoides Zhao, 1995: 823, f. 318a-c (mf)

Pirata piratoides Yin et al., 1997c: 39, f. 16a-d (mf)

Pirata piratoides Song, Zhu & Chen, 1999: 344, f. 200L, 201C (mf)

Pirata piratoides Song, Zhu & Chen, 2001: 257, f. 160A-E (mf)

Pirata piratoides Namkung, 2002: 342, f. 20.36a-b (mf)

Pirata piratoides Kim & Cho, 2002: 219, f. 493-498 (mf)

Pirata piratoides Namkung, 2003: 344, f. 20.36a-b (mf)

Pirata piratoides Tanaka, 2009: 225, f. 22-23 (mf)

Pirata piratoides Zhu & Zhang, 2011: 285, f. 207A-D (mf)

Piratula piratoides Omelko, Marusik & Koponen, 2011: 226, f. 2b, 102-113, 157a-b, 160, 171 (mf, T from Pirata)

Pirata piratoides Yin et al., 2012: 780, f. 389a-d (mf)

Piratula piratoides Dong, 2018: 65, f. 9-8.A-G, pl. 15 (mf)

Piratula piratoides 57 in Wang et al., 2021a: 57, f. 55A-H, 56A-D, 57A-D (mf)

Piratula piratoides Zhang, Peng & Zhang, 2022: 155, f. 113A-H (mf)

Piratula procurva (Bösenberg & Strand, 1906) | | China, Korea, Japan urn:lsid:nmbe.ch:spidersp:019106

Tarentula procurva Bösenberg & Strand, 1906: 315, pl. 13, f. 321 (Dmf) Original description

Tarentula sagaphila Strand, 1918: 99 (Dmf) [urn:lsid:nmbe.ch:spidersp:063994]

Hyaenosa sagaphila Roewer, 1955c: 260

Piratula procurva Roewer, 1955c: 288

Pirata procurva Yaginuma, 1960: 87, f. 78.3 (f)

Pirata procurva Yaginuma, 1961a: 83, f. D-E (m)

Pirata procurvus Yaginuma, 1965b: 366, f. 9.7 (f)

Pirata procurva Paik, 1970b: 87, f. 8-9 (f)

Pirata procurvus Yaginuma, 1971: 87, f. 78.3 (f)

Pirata procurvus Tanaka, 1974a: 38, f. 28-31 (mf)

Pirata procurvus Shimojana, 1977b: 113, pl. 6E (f)

Pirata procurvus sinensis Song et al., 1978: 5, f. 9A-D (Dmf) [urn:lsid:nmbe.ch:spidersp:063993]

Pirata procurvus sinensis Hu, 1984: 244, f. 255.1-4 (mf)

Pirata nanshanensis Yin & Wang, 1984b: 146, f. 1A-J (Dmf) [urn:lsid:nmbe.ch:spidersp:063992]

Pirata procurvus Yaginuma, 1986a: 167, f. 92.5 (mf)

Pirata nanshanensis Song, 1987: 238, f. 194 (mf)

Pirata procurvus Song, 1987: 241, f. 196 (mf, S of Pirata procurvus sinensis)

Pirata procurvus Liu, 1987a: 46, pl. 2, f. 6 (m)

Pirata procurvus Liu, 1987b: 85, f. 18-23 (mf, S of Pirata canadensis [revalidated by Dondale & Redner, 1990: 251], S of P. procurvus sinensis confirmed)

Pirata procurvus Tanaka, 1988a: 57, f. 29-32 (mf, S of Hyaenosa sagaphila)

Pirata procurvus Yu & Song, 1988c: 119 (S of Pirata nanshanensis)

Pirata procurvus Chikuni, 1989b: 113, f. 21 (mf)

Pirata procurvus Feng, 1990: 155, f. 130.1-2 (mf)

Pirata procurvus Chen & Zhang, 1991: 213, f. 217.1-5 (mf)

Pirata procurvus Song, Zhu & Li, 1993: 873, f. 40A-D (mf)

Pirata procurvus Zhao, 1993: 106, f. 46a-c (mf)

Pirata procurvus Barrion & Litsinger, 1994: 317, f. 1659-1661 (f)

Pirata procurvus Zhao, 1995: 824, f. 382a-c (mf)

Pirata procurvus Yin et al., 1997c: 41, f. 17a-d (mf)

Pirata procurvus Song, Zhu & Chen, 1999: 344, f. 200M, 201D (mf)

Pirata procurvus Song, Zhu & Chen, 2001: 259, f. 161A-D (mf)

Pirata procurvus Namkung, 2002: 343, f. 20.37a-b (mf)

Pirata procurvus Kim & Cho, 2002: 220, f. 499-504 (mf)

Pirata procurvus Namkung, 2003: 345, f. 20.37a-b (mf)

Pirata procurvus Lee et al., 2004: 99, unnumbered f. (f)

Pirata procurvus Tanaka, 2009: 227, f. 28-29 (mf)

Pirata procurvus Zhu & Zhang, 2011: 286, f. 208A-D (mf)

Piratula procurva Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Pirata procurvus Yin et al., 2012: 781, f. 390a-d (mf)

Piratula procurvus Dong, 2018: 66, f. 9-9.A-C, pl. 15 (f)

Piratula raika Zamani & Marusik, 2021 | | Iran urn:lsid:nmbe.ch:spidersp:053660

Piratula raika Zamani & Marusik, 2021f: 454, f. 15-22 (Dm) Original description

Piratula serrulata (Song & Wang, 1984) | | Russia (Far East), China urn:lsid:nmbe.ch:spidersp:019112

Pirata serrulatus Song & Wang, 1984: 149, f. 1-4 (Dmf) Original description

Pirata serrulatus Song, 1987: 242, f. 197 (mf)

Piratula serrulata Logunov, 1992e: 59, f. 5 (mf)

Piratula serrulata Omelko, Marusik & Koponen, 2011: 228, f. 5a, 114-123, 148-149, 159, 172 (mf, T from Pirata)

Piratula tanakai (Brignoli, 1983) | | Russia (Far East), Korea, Japan urn:lsid:nmbe.ch:spidersp:019123

Pirata exiguus Tanaka, 1974a: 37, f. 25-27 (Dmf; preoccupied by Banks, 1892, see Piratula minuta) [urn:lsid:nmbe.ch:spidersp:063999]

Pirata tanakai Brignoli, 1983c: 456 (replacement name) Original description

Pirata tanakai Yaginuma, 1986a: 169, f. 92.7 (f)

Pirata tanakai Tanaka, 1988a: 61, f. 33-36 (mf)

Pirata tanakai Chikuni, 1989b: 113, f. 22 (mf)

Pirata tanakai Namkung, 2002: 344, f. 20.38 (f)

Pirata tanakai Namkung, 2003: 346, f. 20.38 (f)

Pirata tanakai Tanaka, 2009: 227, f. 32-33 (mf)

Piratula tanakai Omelko, Marusik & Koponen, 2011: 228, f. 124-135, 162 (mf, T from Pirata)

Piratula tenuisetacea (Chai, 1987) | | China urn:lsid:nmbe.ch:spidersp:019125

Pirata tenuisetaceus Chai, 1987: 363, f. 2.1-12 (Dmf) Original description

Pirata tenuisetaceus Liu, 1987a: 45, pl. 2, f. 10 (m, nomen nudum)

Pirata tenuisetaceus Chen & Zhang, 1991: 212, f. 216.1-6 (mf)

Pirata enuisetaceus Song, Zhu & Li, 1993: 873, f. 41A-B (f)

Pirata tenuisetaceus Zhao, 1993: 111, f. 49a-c (mf)

Pirata tenuisetaceus Zhao, 1995: 829, f. 385a-c (mf)

Pirata tenuisetaceus Yin et al., 1997c: 46, f. 19a-e (mf)

Pirata tenuisetaceus Song, Zhu & Chen, 1999: 344, f. 200O, 201F (mf)

Pirata tenuisetaceus Zhu & Zhang, 2011: 288, f. 210A-D (mf)

Piratula tenuisetacea Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)

Pirata tenuisetaceus Yin et al., 2012: 785, f. 392a-e (mf)

Piratula uliginosa (Thorell, 1856) | | Europe, Russia (Europe to West Siberia) urn:lsid:nmbe.ch:spidersp:019131

Potamia uliginosa Thorell, 1856: 111 (Df) Original description

Lycosa uliginosa Westring, 1861: 533

Pirata uliginosus Thorell, 1870b: 194

Pirata uliginosus Dahl, 1908: 300, f. 28 (f, Dm)

Pirata uliginosus Dahl & Dahl, 1927: 59, f. 154 (mf)

Pirata uliginosus Palmgren, 1939: 67, f. 103, 108, 111 (f)

Pirata uliginosus Holm, 1947: 9, f. 7b, pl. 1, f. 5-6, pl. 9, f. 2 (mf)

Pirata moodyi Hull, 1950: 421, pl. 3, f. 1-2 (Df) [urn:lsid:nmbe.ch:spidersp:063990]

Pirata uliginosus Locket & Millidge, 1953: 416, f. 250A-B (mf)

Pirata uliginosus Muller, 1958: 232, f. 4 (m)

Pirata uliginosus Wiebes, 1964: 98, f. 1-2 (mf)

Pirata uliginosus Buchar, 1966a: 218 (S of Pirata moodyi)

Pirata uliginosus Fuhn & Niculescu-Burlacu, 1971: 217, f. 107a-d (mf)

Pirata uliginosus Miller, 1971: 169, pl. XXVI, f. 23, pl. LXI, f. 24 (mf)

Pirata uliginosus Loksa, 1972: 58, f. 52C, 55A-B (mf)

Pirata uliginosus Kronestedt, 1980b: 65, f. 9L, 10C (m)

Pirata uliginosus Loksa, 1981a: 101, f. 3, 5 (mf)

Pirata uliginosus Roberts, 1985: 150, f. 65d (mf)

Pirata uliginosus Heimer & Nentwig, 1991: 346, f. 895 (mf)

Pirata uliginosus Roberts, 1995: 234, f. (mf)

Pirata uliginosus Roberts, 1998: 252, f. (mf)

Pirata uliginosus Almquist, 2006: 245, f. 241a-f (mf)

Piratula uliginosa Blick et al., 2016: 387 (T from Pirata)

Pirata uliginosus Lecigne, 2016d: 33, f. 15F (f)

Piratula yaginumai (Tanaka, 1974) | | Russia (Far East), China, Korea, Japan urn:lsid:nmbe.ch:spidersp:019136

Pirata yaginumai Tanaka, 1974a: 27, f. 7-10 (Dmf) Original description

Pirata yaginumai Paik, 1976b: 85, f. 5-6 (f)

Pirata qinlingensis Zheng & Qiu, 1980: 379, f. 5-9 (Dmf) [urn:lsid:nmbe.ch:spidersp:063995]

Pirata qiulingensis Qiu, 1981: 15, f. 6A-E (mf)

Pirata qinlingensis Hu, 1984: 244, f. 256.1-4 (mf)

Pirata qinlingensis Guo, 1985: 124, f. 2-63.1-4 (mf)

Pirata qinlingensis Zhu & Shi, 1985: 142, f. 126a-d (mf)

Pirata yaginumai Yaginuma, 1986a: 167, f. 92.2 (mf)

Pirata yaginumai Song, 1987: 244, f. 199 (mf, S of Pirata qinlingensis)

Pirata yaginumai Zhang, 1987: 160, f. 136.1-6 (mf)

Pirata yaginumai Liu, 1987a: 46, pl. 2, f. 5 (m)

Pirata yaginumai Liu, 1987b: 84, f. 6-11 (mf, S of Pirata qinlingensis confirmed)

Pirata yaginumai Tanaka, 1988a: 46, f. 13-16 (mf)

Pirata yaginumai Chikuni, 1989b: 114, f. 24 (mf)

Pirata yaginumai Chen & Zhang, 1991: 210, f. 214.1-5 (mf)

Pirata yaginumai Yin et al., 1997c: 48, f. 20a-d (mf)

Pirata yaginumai Song, Zhu & Chen, 1999: 344, f. 200P, 201G (mf)

Pirata yaginumai Song, Zhu & Chen, 2001: 261, f. 163A-E (mf)

Pirata yaginumai Namkung, 2002: 339, f. 20.33a-b (mf)

Pirata yaginumai Kim & Cho, 2002: 220, f. 505-510 (mf)

Pirata yaginumai Namkung, 2003: 341, f. 20.33a-b (mf)

Pirata yaginumai Tanaka, 2009: 225, f. 20-21 (mf)

Pirata yaginumai Zhu & Zhang, 2011: 289, f. 211A-D (mf)

Piratula yaginumai Omelko, Marusik & Koponen, 2011: 229, f. 2c, 5b-c, 136-145, 156a-b, 164, 174 (mf, T from Pirata)

Pirata yaginumai Yin et al., 2012: 786, f. 393a-d (mf)

Piratula yaginumai Zhang, Peng & Zhang, 2022: 156, f. 114A-H (mf)

Piratula yesoensis (Tanaka, 1985) | | Japan urn:lsid:nmbe.ch:spidersp:019137

Pirata yesoensis Tanaka, 1985: 52, f. 1-4 (Dmf) Original description

Pirata yesoensis Tanaka, 1988a: 64, f. 37-40 (mf)

Pirata yesoensis Tanaka, 2009: 227, f. 34-35 (mf)

Piratula yesoensis Omelko, Marusik & Koponen, 2011: 216 (T from Pirata)