World Spider Catalog

Liocranidae Simon, 1897

Family | ACCEPTED

N.B.: a diagnosis has not been presented (see Lehtinen, 1967: 290-291, 340 for comments); family name predated by Cybaeodinae Simon, 1893, but that name has remained unused; considered a subgroup of Zoridae by Wunderlich, 2008c: 486; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 521, f. 19.

Gen. Scotina Menge, 1873 urn:lsid:nmbe.ch:spidergen:02205

type Agelena gracilipes Blackwall, 1859; gender feminine

Transferred to other genera:

Scotina occulta Kritscher, 1996 -- see Anagraphis

In synonymy:

Scotina bertkaui (Bösenberg, 1899, T from Apostenus) = Scotina palliardii (L. Koch, 1881) (Simon, 1932: 973)

Scotina diversa (O. Pickard-Cambridge, 1913, T from Agroeca) = Scotina palliardii (L. Koch, 1881) (Simon, 1932: 973)

Scotina praelongipes (O. Pickard-Cambridge, 1861) = Scotina gracilipes (Blackwall, 1859) (O. Pickard-Cambridge, 1879d: 39, sub Liocranum)

Scotina celans (Blackwall, 1841) | | Europe, Morocco, Algeria, Turkey urn:lsid:nmbe.ch:spidersp:024394

Agroeca celans Blackwall, 1841a: 624 (Dmf) Original description

Argus celans Walckenaer, 1847a: 504

Agroeca celans Blackwall, 1861a: 161, pl. 10, f. 103 (mf)

Liocranum celans Canestrini & Pavesi, 1868: 774

Agroeca celans O. Pickard-Cambridge, 1895b: 107, pl. 2, f. 8 (f)

Agroeca celans Becker, 1896: 304, pl. 17, f. 14 (mf)

Agroeca celans Chyzer & Kulczyński, 1897: 245, pl. 10, f. 3 (mf)

Scotina celans Bösenberg, 1902: 259, pl. 24, f. 376 (mf)

Scotina celans Simon, 1932: 945, 972, f. 1457-1458 (mf)

Scotina celans Reimoser, 1937a: 84, f. 75-76 (mf)

Scotina celans Tullgren, 1946: 55, pl. 9, f. 112-114 (mf)

Scotina celans Locket & Millidge, 1951: 153, f. 78B, E (mf)

Scotina celans Millidge & Locket, 1952: 60, f. 2e-f (m)

Scotina celans Locket & Millidge, 1953: 412, f. 246E-F (m)

Scotina celans Miller, 1971: 108, pl. XII, f. 15-17 (mf)

Scotina celans Locket, Millidge & Merrett, 1974: 16, f. 8D (f)

Scotina celans Brignoli & Gaddini, 1979: 12, f. 6 (f)

Scotina celans Roberts, 1985: 90, f. 35e (mf)

Scotina celans Grimm, 1986: 63, f. 7, 13, 60, 65-66 (mf)

Scotina celans Heimer & Nentwig, 1991: 394, f. 1026 (mf)

Scotina celans Roberts, 1995: 140, f. (mf)

Scotina celans Roberts, 1998: 149, f. (mf)

Scotina celans Bosmans, 1999: 31, f. 13-16 (mf)

Scotina celans Trotta, 2005: 167, f. 316-317 (mf)

Scotina celans Almquist, 2007: 347, f. 301a-e (mf)

Scotina celans Russell-Smith, 2015: 14, unnumbered f. (mf)

Scotina celans Kunt, Danışman & Coşar, 2024: 2, f. 2A-G, 3A-F (mf)

Scotina celans Bosselaers, 2024b: 11, f. 1E, H, 3H-I, 6A (f)

Scotina celans Zamani et al., 2024b: 210, f. 15, 35-36 (m)

Scotina celans Lecigne, Moutaouakil & Lips, 2025: 138, f. 82A-B (m)

Scotina gracilipes (Blackwall, 1859) | | Europe urn:lsid:nmbe.ch:spidersp:024395 TYPE SPECIES

Agelena gracilipes Blackwall, 1859a: 97 (Dm) Original description

Agelena gracilipes Blackwall, 1861a: 162, pl. 10, f. 104 (m)

Drassus praelongipes O. Pickard-Cambridge, 1861a: 428 (Dmf) [urn:lsid:nmbe.ch:spidersp:062467]

Drassus palliardii O. Pickard-Cambridge, 1871a: 413 (j, nomen nudum; misidentified per Thorell, 1873: 436)

Scotina gracilipes Menge, 1873: 337, pl. 59, f. 192 (mf)

Liocranum gracilipes Thorell, 1873: 436

Liocranum praelongipes O. Pickard-Cambridge, 1873b: 439, pl. 33, f. 4 (f)

Agraeca praelongipes Simon, 1878a: 311

Liocranum gracilipes O. Pickard-Cambridge, 1879d: 39 (S of Liocranum praelongipes)

Agroeca gracilipes Chyzer & Kulczyński, 1897: 245, pl. 10, f. 2 (mf)

Scotina praelongipes Simon, 1897a: 137, f. 146 (m)

Agroeca gracilipes Bösenberg, 1902: 260, pl. 24, f. 377 (mf)

Scotina gracilipes Simon, 1932: 945, 972, f. 1459-1460 (mf, S of Scotina praelongipes confirmed)

Scotina gracilipes Tullgren, 1946: 53, f. 15, pl. 9, f. 104-107 (mf)

Scotina gracilipes Locket & Millidge, 1951: 154, f. 78C, F (mf)

Scotina gracilipes Millidge & Locket, 1952: 60, f. 2b (f)

Scotina gracilipes Locket & Millidge, 1953: 412, f. 246B (f)

Scotina gracilipes Brændegaard, 1966: 201, f. 169-171 (mf)

Scotina gracilipes Roberts, 1985: 90, f. 36a (mf)

Scotina gracilipes Grimm, 1986: 66, f. 61-63 (mf)

Scotina gracilipes Heimer & Nentwig, 1991: 394, f. 1027 (mf)

Scotina gracilipes Roberts, 1995: 140, f. (mf)

Scotina gracilipes Roberts, 1998: 149, f. (mf)

Scotina gracilipes Almquist, 2007: 349, f. 302a-h (mf)

Scotina gracilipes Russell-Smith, 2015: 14, unnumbered f. (mf)

Scotina palliardii (L. Koch, 1881) | | Europe, Turkey, Korea urn:lsid:nmbe.ch:spidersp:024397

Liocranum palliardii L. Koch, 1881b: 60, pl. 2, f. 10 (Dmf; patronym for Palliardi) Original description

Apostenus bertkaui Bösenberg, 1899: 118, pl. 1, f. 6 (Dm) [urn:lsid:nmbe.ch:spidersp:062468]

Apostenus bertkaui Bösenberg, 1902: 255, pl. 24, f. 369 (m)

Apostenus fuscus Bösenberg, 1902: 256, pl. 24, f. 370 (mf; misidentified per Simon, 1932: 973)

Agroeca diversa O. Pickard-Cambridge, 1913: 113, pl. A, f. 1-3 (Df) [urn:lsid:nmbe.ch:spidersp:062469]

Scotina paillardi Simon, 1932: 945, 973 (S of Agroeca diversa and Apostenus bertkaui)

Scotina palliardi Reimoser, 1937a: 85, f. 77-78 (mf)

Scotina palliardi Palmgren, 1943: 62, f. 66 (mf)

Scotina palliardii Tullgren, 1946: 54, pl. 9, f. 108-111 (mf)

Scotina palliardii Millidge & Locket, 1952: 60, f. 2a, c-d (mf)

Scotina palliardii Locket & Millidge, 1953: 410, f. 246A, C-D (mf)

Scotina paillardi Miller, 1971: 108, pl. XII, f. 18 (m)

Scotina palliardii Pakhorukov & Utochkin, 1977a: 91, f. 3 (f)

Scotina palliardii Thaler, 1982c: 406, f. 10-12, 16 (mf)

Scotina palliardii Roberts, 1985: 90, f. 36b (mf)

Scotina palliardii Grimm, 1986: 68, f. 64 (mf)

Scotina palliardii Roberts, 1995: 141, f. (mf)

Scotina palliardii Roberts, 1998: 150, f. (mf)

Scotina palliardii Almquist, 2007: 349, f. 303a-d (mf)

Scotina palliardii Seo, 2011a: 101, f. 9-12 (f)

Scotina palliardii Russell-Smith, 2015: 15, unnumbered f. (f)

Scotina zoraoides Wunderlich, 2024 | | Portugal urn:lsid:nmbe.ch:spidersp:062544

Scotina zoraoides Wunderlich, 2024b: 26, f. 53-57, photo 3 (Dm) Original description