World Spider Catalog

Linyphiidae Blackwall, 1859

Family | ACCEPTED

N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; Tian et al., 2022 show five subfamilies: Linyphiinae (incl. Mynogleninae Lehtinen, 1967: 250 and Dubiaraneinae Millidge, 1993c: 145), Erigoninae Emerton, 1882: 7, Micronetinae Hull, 1920a: 8, Ipainae Saaristo, 2007b: 34 and Stemonyphantinae Wunderlich, 1986: 120; da Silva-Moreira, Kulkarni & Hormiga, 2025: 60-62 list resp. newly circumscribe six subfamilies: Erigoninae, Lepthyphantinae Simon, 1929: 562, Linyphiinae (syn. nov.: Dubiaraneinae), Mynogleninae, Pocobletinae Silva-Moreira & Hormiga, 2025: 61 (new subfamily) and Stemonyphantinae, state Ipainae and Micronetinae to be polyphyletic, leave several linyphiid clades unranked and also several genera as incertae sedis.

Gen. Tiso Simon, 1884 urn:lsid:nmbe.ch:spidergen:01310

type Neriene vagans Blackwall, 1834; gender masculine

Transferred to other genera:

Tiso clavatus Schenkel, 1927 -- see Scotinotylus

Tiso lancearius Tanasevitch, 1987 -- see Allotiso

Tiso nemoralis Holm, 1939 -- see Panamomops

In synonymy:

Tiso carpaticus (Kulczyński, 1882) = Tiso aestivus (L. Koch, 1872) (Kulczyński, 1887: 270)

Tiso hamipalpis (Menge, 1868, T from Tmeticus) = Tiso vagans (Blackwall, 1834) (Simon, 1884a: 750)

Tiso longimanus (C. L. Koch, 1841, T from Neriene) = Tiso vagans (Blackwall, 1834) (Simon, 1884a: 750)

Tiso longipalpis (Wider, 1834, T from Micryphantes) = Tiso vagans (Blackwall, 1834) (Simon, 1884a: 750)

Tiso morosus Simon, 1884 = Tiso aestivus (L. Koch, 1872) (Denis, 1949a: 148)

Nomina dubia:

Tiso megalops Caporiacco, 1935b: 178, pl. 1, f. 18 (mf, India) -- Tanasevitch, 2025b: 106

Tiso strandi Kolosváry, 1934b: 15, f. 8 (f, Hungary) -- Breitling et al., 2016b: 62

Tiso aestivus (L. Koch, 1872) | | Canada, Greenland, Europe, Russia (Europe to Far East), Nepal, Mongolia, Japan urn:lsid:nmbe.ch:spidersp:012928

Erigone aestiva L. Koch, 1872b: 271 (Dmf) Original description

Erigone carpatica Kulczyński, 1882b: 16, pl. 2, f. 10 (Dm) [urn:lsid:nmbe.ch:spidersp:053618]

Tiso morosus Simon, 1884a: 509, f. 296 (Dm) [urn:lsid:nmbe.ch:spidersp:064881]

Tiso aestivus Simon, 1884a: 510

Tiso carpathicus Simon, 1884a: 510

Tiso aestivus Kulczyński, 1887: 270 (S of Tiso carpaticus)

Tiso aestivus Chyzer & Kulczyński, 1894: 127, pl. 5, f. 7 (mf)

Tiso aestivus Jackson, 1914: 127, pl. 3, f. 9-11 (mf)

Tiso morosus Simon, 1926: 371, 493, f. 659-661 (m, Df)

Tiso aestivus Brændegaard, 1946: 51, f. 29 (f)

Tiso aestivus Denis, 1949a: 148 (S of Tiso morosus)

Tiso aestivus Locket & Millidge, 1953: 264, f. 159F-G, 160B (mf)

Tiso aestivus Tullgren, 1955: 382, f. 83a-c (mf)

Tiso aestivus Brændegaard, 1958: 48, f. 42-43 (mf)

Tiso aestivus Wiehle, 1960a: 399, f. 729-735 (mf)

Tiso aestivus Thaler, 1970: 269, f. 40-43 (m)

Tiso aestivus Tystshenko, 1971: 262, f. 900, 904 (mf)

Tiso aestivus Miller, 1971: 271, pl. LVI, f. 18-19 (mf)

Tiso aestivus Palmgren, 1976: 104, f. 19.4-5 (mf)

Tiso aestivus Millidge, 1977: 6, f. 4 (m)

Tiso aestivus Saito, 1982c: 15, f. 9, 35 (m)

Tiso aestivus Roberts, 1987: 70, f. 29e (mf)

Tiso aestivus Heimer & Nentwig, 1991: 258, f. 684 (mf)

Tiso aestivus Agnarsson, 1996: 91, f. 85A-B (mf)

Tiso aestivus Ono, Matsuda & Saito, 2009: 263, f. 61-62 (mf)

Tiso aestivus Marusik, 2015: 691, f. 13Y-Z (mf)

Tiso biceps Gao, Zhu & Gao, 1993 | | China urn:lsid:nmbe.ch:spidersp:037635

Tiso biceps Gao, Zhu & Gao, 1993: 41, f. 3-5 (Dm) Original description

Tiso camillus Tanasevitch, 1990 | | Azerbaijan urn:lsid:nmbe.ch:spidersp:012929

Tiso camillus Tanasevitch, 1990: 60, f. 15.6-9 (Dmf) Original description

Tiso golovatchi Tanasevitch, 2006 | | Russia (Far East) urn:lsid:nmbe.ch:spidersp:039984

Tiso golovatchi Tanasevitch, 2006c: 30, f. 19-25 (Dm; N.B.: erroneously listed as T. solivagus in English figure legend) Original description

Tiso incisus Tanasevitch, 2011 | | Pakistan, India urn:lsid:nmbe.ch:spidersp:044358

Tiso incisus Tanasevitch, 2011d: 590, f. 89-96 (Dmf) Original description

Tiso indianus Tanasevitch, 2011 | | India, Nepal urn:lsid:nmbe.ch:spidersp:044359

Tiso (?) indianus Tanasevitch, 2011d: 591, f. 97-100, 118 (Dm) Original description

Tiso vagans (Blackwall, 1834) | | Madeira, Europe, Caucasus (Russia, Georgia). Introduced to Canada urn:lsid:nmbe.ch:spidersp:012932 TYPE SPECIES

Theridion longipalpe Wider, 1834: 222, pl. 15, f. 7 (Dm) [urn:lsid:nmbe.ch:spidersp:064882]

Neriene vagans Blackwall, 1834b: 374 (Dmf) Original description

Micryphantes longipalpus C. L. Koch, 1837b: 11

Argus longipalpus Walckenaer, 1841: 354

Erigone longimana C. L. Koch, 1841a: 93, f. 661-662 (Dmf) [urn:lsid:nmbe.ch:spidersp:064883]

Neriene vagans Blackwall, 1864a: 257, pl. 18, f. 173 (mf)

Tmeticus hamipalpis Menge, 1868: 192, pl. 38, f. 95 (Dmf) [urn:lsid:nmbe.ch:spidersp:064884]

Neriene longimana O. Pickard-Cambridge, 1879d: 110

Tiso vagans Simon, 1884a: 507, f. 293-295 (mf, S of Micryphantes longipalpis, Neriene longimana and Tmeticus hamipalpis)

Microctenonyx longimana Dahl, 1886: 81

Tiso vagans Simon, 1894a: 653, f. 637-639 (m)

Tiso vagans Becker, 1896: 91, pl. 10, f. 1 (mf)

Tiso longimanus Bösenberg, 1902: 164, pl. 14, f. 222 (mf)

Tiso vagans Simon, 1926: 371, 493, f. 657-658 (mf, S of Tiso longimanus confirmed)

Tiso vagans Locket & Millidge, 1953: 261, f. 159D-E, 160A (mf)

Tiso vagans Tullgren, 1955: 381, f. 82a-d (mf)

Tiso vagans Wiehle, 1960a: 396, f. 722-728 (mf)

Tiso vagans Merrett, 1963b: 413, f. 88A-C (m)

Tiso vagans Tystshenko, 1971: 262, f. 901 (m)

Tiso vagans Miller, 1971: 271, pl. LVI, f. 20-21 (mf)

Tiso vagans Palmgren, 1976: 105, f. 19.1-3 (mf)

Tiso vagans Millidge, 1977: 6, f. 3 (m)

Tiso vagans Roberts, 1987: 70, f. 29d (mf)

Tiso vagans Heimer & Nentwig, 1991: 258, f. 685 (mf)

Tiso vagans Seropian et al., 2023: 256, f. 52 (f)