World Spider Catalog

Linyphiidae Blackwall, 1859

Family | ACCEPTED

N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; Tian et al., 2022 show five subfamilies: Linyphiinae (incl. Mynogleninae Lehtinen, 1967: 250 and Dubiaraneinae Millidge, 1993c: 145), Erigoninae Emerton, 1882: 7, Micronetinae Hull, 1920a: 8, Ipainae Saaristo, 2007b: 34 and Stemonyphantinae Wunderlich, 1986: 120; da Silva-Moreira, Kulkarni & Hormiga, 2025: 60-62 list resp. newly circumscribe six subfamilies: Erigoninae, Lepthyphantinae Simon, 1929: 562, Linyphiinae (syn. nov.: Dubiaraneinae), Mynogleninae, Pocobletinae Silva-Moreira & Hormiga, 2025: 61 (new subfamily) and Stemonyphantinae, state Ipainae and Micronetinae to be polyphyletic, leave several linyphiid clades unranked and also several genera as incertae sedis.

Gen. Pityohyphantes Simon, 1929 urn:lsid:nmbe.ch:spidergen:01201

type Linyphia phrygiana C. L. Koch, 1836; gender masculine

Transferred to other genera:

Pityohyphantes palilis (L. Koch, 1870) -- see Neriene

In synonymy:

Pityohyphantes costatus annulipes (Banks, 1892) = Pityohyphantes costatus (Hentz, 1850) (Sherwood, 2024b: 220)

Pityohyphantes phrygianus pirini (Drensky, 1921) = Pityohyphantes phrygianus (C. L. Koch, 1836) (Deltshev & Blagoev, 2001: 110)

Pityohyphantes vancouveranus Chamberlin & Ivie, 1942 = Pityohyphantes rubrofasciatus (Keyserling, 1886) (Crawford, 1988: 22)

Pityohyphantes alticeps Chamberlin & Ivie, 1943 | | Canada, USA urn:lsid:nmbe.ch:spidersp:012349

Pityohyphantes alticeps Chamberlin & Ivie, 1943: 27, f. 45-47 (Dm) Original description

Pityohyphantes brachygynus Chamberlin & Ivie, 1942 | | Canada, USA urn:lsid:nmbe.ch:spidersp:012350

Pityohyphantes brachygynus Chamberlin & Ivie, 1942a: 57, f. 140 (Df) Original description

Pityohyphantes costatus (Hentz, 1850) | | Canada, USA urn:lsid:nmbe.ch:spidersp:012351

Linyphia costata Hentz, 1850: 31, pl. 4, f. 11 (Df) Original description

Linyphia phrygiana Emerton, 1882: 63, pl. 19, f. 1a-h (mf; misidentified per Chamberlin & Ivie, 1943: 28)

Linyphia phrygiana Keyserling, 1886b: 60, pl. 12, f. 164 (mf; misidentified sensu Chamberlin & Ivie, 1943: 28)

Linyphia phrygianus annulipes Banks, 1892a: 42 (D) [urn:lsid:nmbe.ch:spidersp:012352]

Linyphia phrygiana Emerton, 1902: 141, f. 332-334 (mf; misidentified sensu Chamberlin & Ivie, 1943: 28)

Linyphia phrygiana Comstock, 1910: 171, f. 10-11 (m; misidentified sensu Chamberlin & Ivie, 1943: 28)

Linyphia phrygiana Comstock, 1912: 394, f. 407-409 (mf; misidentified sensu Chamberlin & Ivie, 1943: 28)

Pityohyphantes phrygianus Blauvelt, 1936: 137, pl. 12, f. 80-84, 86 (mf; misidentified sensu Chamberlin & Ivie, 1943: 28)

Linyphia phrygiana Comstock, 1940: 112, 408, f. 102-103, 379, 386 (m; misidentified sensu Chamberlin & Ivie, 1943: 28)

Pityohyphantes phrygianus Muma, 1943: 81, pl. XIII, f. 13-14 (mf; misidentified sensu Chamberlin & Ivie, 1943: 28)

Pityohyphantes costatus Chamberlin & Ivie, 1943: 28 (removed from S of Pityohyphantes phrygianus, rejecting Emerton, 1882: 63)

Pityohyphantes phrygianus Kaston, 1948: 119, f. 240-248 (mf; misidentified sensu Chamberlin & Ivie, 1943: 28)

Pityohyphantes costatus Kaston, 1972a: 125, f. 16, 282-283 (mf)

Pityohyphantes costatus Paquin & Dupérré, 2003: 146, f. 1615-1617 (mf)

Pityohyphantes costatus Sherwood, 2024b: 220 (S of Pityohyphantes costatus annulipes)

Pityohyphantes cristatus Chamberlin & Ivie, 1942 | | Canada, USA urn:lsid:nmbe.ch:spidersp:012353

Pityohyphantes cristatus Chamberlin & Ivie, 1942a: 58, f. 141-143 (Dmf) Original description

Pityohyphantes cristatus coloradensis Chamberlin & Ivie, 1942 | | USA urn:lsid:nmbe.ch:spidersp:012354

Pityohyphantes cristatus coloradensis Chamberlin & Ivie, 1942a: 58, f. 144-145 (Dmf) Original description

Pityohyphantes hesperus (Chamberlin, 1920) | | USA urn:lsid:nmbe.ch:spidersp:012355

Linyphia hespera Chamberlin, 1920a: 194, f. 19.4., 20.1-2 (Dm) Original description

Pityohyphantes hesperus Gertsch, 1935a: 22 (Df)

Pityohyphantes kamela Chamberlin & Ivie, 1943 | | Canada, USA urn:lsid:nmbe.ch:spidersp:012356

Pityohyphantes kamela Chamberlin & Ivie, 1943: 28, f. 51-53 (Dmf) Original description

Pityohyphantes limitaneus (Emerton, 1915) | | Canada, USA urn:lsid:nmbe.ch:spidersp:012357

Linyphia limitanea Emerton, 1915a: 137, pl. 1, f. 5 (Dm) Original description

Pityohyphantes limitanea Blauvelt, 1936: 142, pl. 12, f. 85, pl. 13, f. 87-89 (m, Df)

Pityohyphantes limitaneus Paquin & Dupérré, 2003: 146, f. 1618-1620 (mf)

Pityohyphantes lomondensis Chamberlin & Ivie, 1941 | | USA urn:lsid:nmbe.ch:spidersp:012358

Pityohyphantes lomondensis Chamberlin & Ivie, 1941b: 13, f. 49 (Df) Original description

Pityohyphantes minidoka Chamberlin & Ivie, 1943 | | Canada, USA urn:lsid:nmbe.ch:spidersp:012359

Pityohyphantes minidoka Chamberlin & Ivie, 1943: 29, f. 54-56 (Dmf) Original description

Pityohyphantes navajo Chamberlin & Ivie, 1942 | | USA urn:lsid:nmbe.ch:spidersp:012360

Pityohyphantes navajo Chamberlin & Ivie, 1942a: 60, f. 151-152 (Dm) Original description

Pityohyphantes pallidus Chamberlin & Ivie, 1942 | | USA urn:lsid:nmbe.ch:spidersp:012362

Pityohyphantes pallidus Chamberlin & Ivie, 1942a: 59, f. 147-148 (Dm) Original description

Pityohyphantes phrygianus (C. L. Koch, 1836) | | Europe, Russia (Europe to Far East), Kazakhstan, Japan urn:lsid:nmbe.ch:spidersp:012363 TYPE SPECIES

Linyphia phrygiana C. L. Koch, 1836a: 83, f. 229-230 (Dmf) Original description

Linyphia phrygiana Emerton, 1882: 63 (S of Linyphia costata, revalidated by Chamberlin & Ivie, 1943: 28, see Pityohyphantes costatus)

Linyphia phrygiana Chyzer & Kulczyński, 1894: 56, pl. 2, f. 21 (mf)

Linyphia phrygiana Bösenberg, 1901: 69, pl. 5, f. 70 (mf)

Linyphia phrygiana pirini Drensky, 1921: 10, 76 (Dmf) [urn:lsid:nmbe.ch:spidersp:012364]

Pityohyphantes phrygianus Simon, 1929: 623, 741, f. 935-937 (mf)

Pityohyphantes phrygianus Reimoser, 1930b: 53, f. 6 (mf)

Pityohyphantes phrygianus Wiehle, 1956: 275, f. 455-459 (mf)

Pityohyphantes phrygianus Wiehle, 1967b: 187, f. 17 (f)

Pityohyphantes phrygianus Fuhn & Niculescu-Burlacu, 1969: 76, pl. II, f. 8a-b (mf)

Pityohyphantes phrygianus Tystshenko, 1971: 228, f. 641, 645 (mf)

Pityohyphantes phrygianus Miller, 1971: 219, pl. XLII, 5-6 (mf)

Pityohyphantes phrygianus Palmgren, 1975: 38, f. 9.4-7 (mf)

Pityohyphantes phrygianus Ashmole et al., 1978: 283, f. 2-10 (mf)

Pityohyphantes phrygianus Roberts, 1987: 159, f. 82b (mf)

Pityohyphantes phrygianus Coddington, 1990: 15, f. 68 (m)

Pityohyphantes phrygianus Heimer & Nentwig, 1991: 234, f. 625 (mf)

Pityohyphantes phrygianus Roberts, 1995: 364, f. (mf)

Pityohyphantes phrygianus Roberts, 1998: 374, f. (mf)

Pityohyphantes phrygianus Deltshev & Blagoev, 2001: 110 (S of Pityohyphantes phrygianus pirini)

Pityohyphantes phrygianus Uhl & Gunnarsson, 2001: 368, f. 1a-d (f)

Pityohyphantes phrygianus Deltshev, 2003: 137, f. 1-4 (mf)

Pityohyphantes phrygianus Gunnarsson, Uhl & Wallin, 2004: 138, f. 5-6 (f)

Pityohyphantes phrygianus Ono, Matsuda & Saito, 2009: 337, f. 1166-1168 (mf)

Pityohyphantes phrygianus Lecigne, 2018b: 71, f. 8J (f)

Pityohyphantes phrygianus Geci et al., 2023: 541, f. 18 (f)

Pityohyphantes rubrofasciatus (Keyserling, 1886) | | Canada, USA urn:lsid:nmbe.ch:spidersp:012365

Linyphia rubrofasciata Keyserling, 1886b: 66, pl. 13, f. 168 (Dmf) Original description

Pityohyphantes vancouveranus Chamberlin & Ivie, 1942a: 59, f. 146 (Dm) [urn:lsid:nmbe.ch:spidersp:056142]

Pityohyphantes rubrofasciatus Crawford, 1988: 22 (S of Pityohyphantes vancouveranus)

Pityohyphantes subarcticus Chamberlin & Ivie, 1943 | | Alaska, Canada urn:lsid:nmbe.ch:spidersp:012366

Pityohyphantes subarcticus Chamberlin & Ivie, 1943: 28, f. 48-50 (Dmf) Original description

Pityohyphantes subarcticus Paquin & Dupérré, 2003: 147, f. 1621-1622 (mf)

Pityohyphantes tacoma Chamberlin & Ivie, 1942 | | Alaska, USA urn:lsid:nmbe.ch:spidersp:012367

Pityohyphantes tacoma Chamberlin & Ivie, 1942a: 60, f. 149-150 (Dm) Original description