World Spider Catalog

Linyphiidae Blackwall, 1859

Family | ACCEPTED

N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; Tian et al., 2022 show five subfamilies: Linyphiinae (incl. Mynogleninae Lehtinen, 1967: 250 and Dubiaraneinae Millidge, 1993c: 145), Erigoninae Emerton, 1882: 7, Micronetinae Hull, 1920a: 8, Ipainae Saaristo, 2007b: 34 and Stemonyphantinae Wunderlich, 1986: 120; da Silva-Moreira, Kulkarni & Hormiga, 2025: 60-62 list resp. newly circumscribe six subfamilies: Erigoninae, Lepthyphantinae Simon, 1929: 562, Linyphiinae (syn. nov.: Dubiaraneinae), Mynogleninae, Pocobletinae Silva-Moreira & Hormiga, 2025: 61 (new subfamily) and Stemonyphantinae, state Ipainae and Micronetinae to be polyphyletic, leave several linyphiid clades unranked and also several genera as incertae sedis.

Gen. Piniphantes Saaristo & Tanasevitch, 1996 urn:lsid:nmbe.ch:spidergen:01200

type Lepthyphantes pinicola Simon, 1884; gender masculine

N.B.: the synonymy with Oryphantes Hull, 1932, proposed by Breitling, 2019b: 30 and Breitling, 2021: 21, is not accepted here as it is based mainly on COI (see FAQ 7).

In synonymy:

Piniphantes pinicola valesiacus (Schenkel, 1925) = Piniphantes pinicola (Simon, 1884) (Thaler, 1986d: 230, sub Lepthyphantes)

Piniphantes tauricola (Strand, 1910) = Piniphantes pinicola (Simon, 1884) (Tanasevitch & Eskov, 1987: 185, sub Lepthyphantes)

Piniphantes agnellus (Maurer & Thaler, 1988) | | Alps (France, Italy) urn:lsid:nmbe.ch:spidersp:011073

Lepthyphantes agnellus Maurer & Thaler, 1988: 338, f. 17-19 (Df) Original description

Piniphantes agnellus Milano et al., 2019: 39, f. 6A-B, 7A-D, 8A-C (Dm, f, T from Lepthyphantes)

Piniphantes cinereus (Tanasevitch, 1986) | | Kyrgyzstan urn:lsid:nmbe.ch:spidersp:012341

Lepthyphantes cinereus Tanasevitch, 1986: 150, f. 43-49 (Dmf) Original description

Piniphantes cinereus Saaristo & Tanasevitch, 1996b: 180 (T from Lepthyphantes)

Piniphantes cirratus (Thaler, 1986) | | France (Corsica) urn:lsid:nmbe.ch:spidersp:012342

Lepthyphantes cirratus Thaler, 1986d: 226, f. 1-7 (Dm) Original description

Piniphantes cirratus Saaristo & Tanasevitch, 1996b: 180 (T from Lepthyphantes)

Piniphantes deosaicola (Caporiacco, 1935) | | Pakistan urn:lsid:nmbe.ch:spidersp:011145

Lepthyphantes deosaicola Caporiacco, 1935b: 165, pl. 3, f. 6 (Df) Original description

Piniphantes deosaicola Tanasevitch, 2025b: 112, f. 9A (f, T from Lepthyphantes)

Piniphantes himalayensis (Tanasevitch, 1987) | | Pakistan, Nepal urn:lsid:nmbe.ch:spidersp:012343

Lepthyphantes sp. prope pinicola Thaler, 1987b: 37, f. 22-27, 43-44 (mf; here per Tanasevitch, 1987b: 60)

Lepthyphantes uzbekistanicus himalayensis Tanasevitch, 1987b: 60, f. 87-90 (Dmf) Original description

Lepthyphantes himalayensis Saaristo & Tanasevitch, 1993: 60 (elevated from subspecies of Lepthyphantes uzbekistanicus)

Piniphantes himalayensis Saaristo & Tanasevitch, 1996b: 180 (T from Lepthyphantes)

Piniphantes macer (Tanasevitch, 1986) | | Kyrgyzstan urn:lsid:nmbe.ch:spidersp:012344

Lepthyphantes macer Tanasevitch, 1986: 144, f. 20-26 (Dmf) Original description

Piniphantes macer Saaristo & Tanasevitch, 1996b: 180, f. 13C (m, T from Lepthyphantes)

Piniphantes pinicola (Simon, 1884) | | Europe, Turkey, Caucasus (Russia, Georgia, Armenia), Iran, Turkmenistan urn:lsid:nmbe.ch:spidersp:012345 TYPE SPECIES

Lepthyphantes pinicola Simon, 1884a: 312, f. 78-79 (Dmf) Original description

Lepthyphantes pinicola F. O. Pickard-Cambridge, 1891: 78, pl. 2, f. 3 (mf)

Lepthyphantes tauricola Strand, 1910d: 116 (Df) [urn:lsid:nmbe.ch:spidersp:065185]

Lepthyphantes pinicola valesiaca Schenkel, 1925d: 304 (Df) [urn:lsid:nmbe.ch:spidersp:065184]

Lepthyphantes pinicola Simon, 1929: 607, 609, 735, f. 923 (mf)

Lepthyphantes pinicola valesiacus Roewer, 1942a: 553

Lepthyphantes pinicola Locket & Millidge, 1953: 392, f. 233D, 235C (mf)

Lepthyphantes pinicola Wanless, 1971: 23, f. 3E-F, 7C (f)

Lepthyphantes pinicola Wanless, 1973: 131, f. III.1, V.10, VIII.13, IX.21 (f)

Lepthyphantes pinicola Thaler, 1983a: 142, f. 49 (f)

Lepthyphantes pinicola Thaler, 1986d: 230, f. 9-11, 17, 23-24 (mf, S of Lepthyphantes pinicola valesiacus)

Lepthyphantes pinicola Roberts, 1987: 152, f. 80a (mf)

Lepthyphantes pinicola Tanasevitch & Eskov, 1987: 185 (S of Lepthyphantes tauricola)

Lepthyphantes pinicola Tanasevitch, 1990: 28, f. 6.2-5 (mf)

Lepthyphantes pinicola Heimer & Nentwig, 1991: 196, f. 533 (mf)

Piniphantes pinicola Saaristo & Tanasevitch, 1996b: 180, f. 13A (m, T from Lepthyphantes)

Piniphantes pinicola Merrett, 2004b: 21, f. 4 (m)

Piniphantes pinicola Danışman et al., 2013: 588, f. 1A-B, 2A-D (mf)

Piniphantes plumatus (Tanasevitch, 1986) | | Kyrgyzstan urn:lsid:nmbe.ch:spidersp:012346

Lepthyphantes plumatus Tanasevitch, 1986: 146, f. 27-34 (Dmf) Original description

Piniphantes plumatus Saaristo & Tanasevitch, 1996b: 180, f. 13B (m, T from Lepthyphantes)

Piniphantes uzbekistanicus (Tanasevitch, 1983) | | Uzbekistan, Kyrgyzstan urn:lsid:nmbe.ch:spidersp:012347

Lepthyphantes uzbekistanicus Tanasevitch, 1983b: 1788, pl. 2, f. 1-6 (Dm; f f. 7-8 see Piniphantes zonsteini) Original description

Lepthyphantes uzbekistanicus Thaler, 1986d: 233, f. 12-13, 15-16, 19-21 (mf)

Lepthyphantes uzbekistanicus Tanasevitch, 1987b: 61, f. 91-93 (mf)

Lepthyphantes uzbekistanicus Tanasevitch, 1989a: 115, f. 85-88 (mf)

Piniphantes uzbekistanicus Saaristo & Tanasevitch, 1996b: 180 (T from Lepthyphantes)

Piniphantes zonsteini (Tanasevitch, 1989) | | Uzbekistan, Kyrgyzstan urn:lsid:nmbe.ch:spidersp:012348

Lepthyphantes uzbekistanicus Tanasevitch, 1983b: 1788, pl. 2, f. 7-8 (f; misidentified per Tanasevitch, 1989a: 115)

Lepthyphantes zonsteini Tanasevitch, 1989a: 116, f. 91-96 (Dmf) Original description

Piniphantes zonsteini Saaristo & Tanasevitch, 1996b: 180 (T from Lepthyphantes)